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Author Hedman, J.
Title Heart rate response towards fear-eliciting stimuli in horses Type Manuscript
Year 2003 Publication Sveriges lantbruksuniversitetSveriges lantbruksuniversitet Veterinärprogrammet Abbreviated Journal
Volume 2004 Issue 40 Pages
Keywords (up) horse, fear, heart-rate, novel stimuli
Abstract Finding the right horse for each rider is a difficult task as it is just as important that the temperament of the horse fits the rider as it is that the horse is of the right size. Even though it is a commonly experienced problem, no objective method of easily measuring the horse“s temperament has yet been developed. If it is possible to test horses and get an objective measure of how reactive (emotional) they are, it could be a big help in finding the right horse for each rider. It would be desirable to have a way of testing the horse”s reaction in an unfamiliar (and potentially frightening) situation. In practice this test should be just as easy as it is getting a judgement of its conformation and gaits.

The aim of the present study was to measure individual variation in HR response to different novel objects in horses of the same age, breed and reared in the same environment. We wanted to see whether certain horses (i.e. more emotional horses) react more to novel stimuli, in general, than other horses (i.e. less emotional), irrespective of the type of stimulus. We also wanted to see if different novel stimuli elicited different responses within individuals. The hypothesis was that individuals will react in a similar way to various stimuli.

Twenty four Danish warmblood horses were included in this study. All horses were 2 year-old stallions, reared under similar environmental conditions. They had received a minimum of handling prior to the experimental period. Three different stimuli were used. They were chosen because they were novel to the horses and would elicit measurable fear-reactions in all horses, but not so much that the horses did not approach the feed within the duration of the test. The visual stimulus consisted of a 1meter high orange traffic cone with reflex stripes, placed 1 m in front of the tub, the olfactory stimulus was eucalyptus oil and the auditory stimulus was a radio tuned to white noise. The control was an empty arena.

The result was that only the HR response to the auditory and visual stimuli differed significantly from the control days. The olfactory stimulus did not seem to alarm the horses the way the other stimuli did. We found a tendency towards a correlation in reaction between the olfactory and auditory stimuli and between the auditory and visual stimuli within individuals. These results indicate that horses do not generalize completely in their reaction between different stimuli.
Address
Corporate Author Thesis Bachelor's thesis
Publisher Place of Publication Dept. of Animal Environment and Health, SLU. Examensarbete / Sveriges lantbruksuniversitet, Veterinä Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 1650-7045 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 4652
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Author Hanggi, E.B.
Title Discrimination learning based on relative size concepts in horses (Equus caballus) Type Journal Article
Year 2003 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 83 Issue 3 Pages 201-213
Keywords (up) Horse; Concept; Size transposition; Generalization; Learning; Training
Abstract This study explored whether or not horses (Equus caballus) could respond to stimuli using a concept based on relative size. In Experiment 1, after learning to respond to the larger of the two stimuli for six sets of two-dimensional (2D) training exemplars, one horse was tested for size transposition that used novel larger and smaller stimuli as well as three-dimensional (3D) objects (5 two-dimensional sets and 5 three-dimensional sets with large, medium, small, and tiny sizes). The horse correctly chose (significantly above chance) the larger of two stimuli regardless of novelty or dimension or combination. In Experiment 2, two additional horses were tested using a subset of the stimuli from Experiment 1. One horse was required to select the larger stimulus--as in Experiment 1--and the other the smaller stimulus. After learning the task, both horses responded correctly to new stimuli and showed size transposition. These results suggest that at least some horses are capable of solving problems based on relative size concepts. Moreover, they are able to generalize across situations that vary from flat, black shapes to objects of different materials and colors including balls, flower pots, and PVC connectors. These findings support earlier research that showed that horses could categorize certain stimuli, and provide new evidence that they are capable of using some form of concept for problem solving. Understanding that horses have more advanced learning abilities than was previously believed should help improve training methods and management.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
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ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number refbase @ user @ Serial 398
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Author McCall, C.A.; Salters, M.A.; Johnson, K.B.; Silverman, S.J.; McElhenney, W.H.; Lishak, R.S.
Title Equine utilization of a previously learned visual stimulus to solve a novel task Type Journal Article
Year 2003 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 82 Issue 3 Pages 163-172
Keywords (up) Horse; Learned stimulus; Novel task; Discrimination
Abstract Forty-four horses were used to determine if a learned stimulus could be used to increase acquisition of a response unrelated to the original learned task. Horses were paired by age, breed and sex. One randomly chosen horse from each pair served as the control while the remaining horse was trained to respond to a visual stimulus by pushing a lever to obtain a food reward. In Experiment I (n=28), the food reward was delivered in a feed box located equidistant between two levers located 2.7 m apart on one wall of the test stall. Trained horses were given 30 training trials daily until they achieved at least 85% correct responses in three nonconsecutive days. After each trained horse reached criterion, both horses in the pair were tested in 30 trials daily for five consecutive days in a modified Y maze. The stimulus that indicated the correct lever in the initial training task was used to signify the correct arm of the maze for both the control and trained horses. The correct arm of the maze was changed randomly during each daily session, and correct choices resulted in a food reward. In Experiment II (n=16), levers were located on opposite walls (11 m apart) of the test stall, and the stimulus and modified Y maze were simplified. Trained horses received 15 training trials daily until they reached at least 85% correct responses on three consecutive days. As each trained horse achieved this criterion, both horses in the pair were tested in a maze for 15 trials daily for 7 days. Again, the stimulus from initial lever-pressing task was used to signify the rewarded arm of the maze. Correct choices of trained and control horses in both experiments were compared using a paired t-test. In Experiment I, no differences (P>0.34) in mean correct responses were detected between trained and control horses on the first day in the maze (15.3 and 16.8, respectively) or after 5 days in the maze (84.0 and 82.1, respectively). Similarly, in Experiment II, trained and control horses did not differ (P>0.50) in mean correct responses on the first day (6.9 and 7.4, respectively) or after 7 days in the maze (63.6 and 61.6, respectively). These results indicate that the stimuli used to solve the lever-pressing task were not utilized by horses in the maze task, and that horses may have difficulty transferring learned visual stimuli to new tasks.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 3587
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Author Wallin, L.; Strandberg, E.; Philipsson, J.
Title Genetic correlations between field test results of Swedish Warmblood Riding Horses as 4-year-olds and lifetime performance results in dressage and show jumping Type Journal Article
Year 2003 Publication Livestock Production Science Abbreviated Journal
Volume 82 Issue 1 Pages 61-71
Keywords (up) Horse; Performance test; Competition results; Animal model; Heritability; Genetic correlation
Abstract The main objective of this study was to estimate genetic correlations between traits of young sport horses (4 years old) evaluated in the Swedish Riding Horse Quality Test (RHQT) and later competition results in dressage and show jumping. The data comprised 3708 Warmblood horses born between 1968 and 1982 that had participated in the RHQT as 4-year-olds and 25[punctuation space]605 horses born between 1953 and 1995 with competition records. According to the criteria between 1206 and 1879 horses were common to this two files and were available for the estimations of the genetic correlations. Competition performance traits were cumulative points and cumulative placings received during a horse's lifetime, and a log10 transformation was used to achieve a more normal distribution of the data. Genetic correlations between gait traits scored in the RHQT and competition results in dressage were favourable, in the range 0.63-0.75, and between jumping traits scored in the RHQT and results in show jumping 0.83-0.93. Estimated heritabilities for gait and jumping traits scored in the RHQT were in the range 0.09-0.27 and 0.10-0.18, respectively. Estimated heritabilities for the cumulative points and cumulative placings in dressage and show jumping were 0.17/0.16 and 0.23/0.27, respectively. Thus, the results from the RHQT have proved to be useful for early genetic evaluation and selection of both mares and stallions for sport performance traits.
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Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
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ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number refbase @ user @ Serial 3956
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Author Visser, E.K.; van Reenen, C.G.; Schilder, M.B.H.; Barneveld, A.; Blokhuis, H.J.
Title Learning performances in young horses using two different learning tests Type Journal Article
Year 2003 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 80 Issue Pages 311-326
Keywords (up) Horse; Personality; Learning performance; Consistency; Emotionality
Abstract To achieve optimal performance in equine sports as well as in leisure not only the physical abilities of the horse should be considered, but also the horse's personality. Besides temperamental aspects, like emotionality, or the horse's reactivity towards humans in handling situations, the learning ability of the horse is another relevant personality trait. To study whether differences in learning performance are consistent over time and whether individual learning performance differs between learning tests or is affected by emotionality, 39 young horses (Dutch Warmblood) were tested repeatedly in two learning tests. An aversive stimulus (AS) was used in one learning test (the avoidance learning test) and a reward was used in the other learning test (the reward learning test). During both learning tests behaviour as well as heart rate were measured. Each test was executed four times, twice when horses were 1 year of age, and twice when they were 2 years of age. Half of the horses received additional physical training from 6 months onwards. In both tests horses could be classified as either performers, i.e. completing the daily session, or as non-performers, i.e. returning to the home environment without having completed the daily session. There were some indications that emotionality might have caused non-performing behaviour, but these indications are not convincing enough to exclude other causes. Furthermore, there seem to be no simple relationships between measures of heart rate, behavioural responses putatively related to emotionality and learning performance. Horses revealed consistent individual learning performances within years in both tests, and in the avoidance learning test also between years. There was no significant correlation between learning performances in the avoidance learning test and the learning performances in the reward learning test. It is concluded that individual learning abilities are consistent over a short time interval for an avoidance learning test and a reward learning test and over a longer time for the avoidance learning test. Furthermore, results indicate that some horses perform better when they have to learn to avoid an aversive stimulus while others perform better when they are rewarded after a correct response. It is suggested that these differences may be relevant to design optimal individual training programmes and methods.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
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ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number refbase @ user @ Serial 2009
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Author Sondergaard, E.; Halekoh, U.
Title Young horses' reactions to humans in relation to handling and social environment Type Journal Article
Year 2003 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 84 Issue 4 Pages 265-280
Keywords (up) Horses; Human-animal relationship; Social environment; Handling
Abstract Forty Danish warmblood colts in two replicates were used to investigate the effect of housing and handling in the rearing period on the reactions to humans. The horses entered the experiment after weaning and were housed either individually (n=16) or in groups of three (n=24). Half of the horses from each housing group were handled three times per week for a period of 10 min. Approach tests were performed in the home environment when the horses were 6, 9, 12, 18, 21, and 24 months old, and an Arena and Human Encounter test was performed in a novel environment when the horses were 12 and 24 months old, respectively. In the home environment, single-housed horses approached sooner and were more easily approached by a human than group-housed horses where no effect of handling was observed. Horses approached sooner and were more easily approached with increasing age. In the Arena and Human Encounter test, single-housed horses expressed less restless behaviour, more explorative behaviour, and less vocalisation than group-housed horses. Handled horses showed lower increase in heart rate during the test than non-handled horses. There was no difference between the number of times single or group-housed horses touched an unfamiliar person in the Arena and Human Encounter test but handled horses approached sooner than non-handled horses. It is concluded that the social environment affected the way horses reacted to humans when tested in the home environment but not in a novel environment. In contrast, handling affected the reactions to humans when tested in the novel environment but not in the home environment. However, handled horses also reacted less to the novel environment in general, thus indicating that handling is a mean of avoiding potential dangerous situations.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number refbase @ user @ Serial 308
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Author Newman, M.E.J.
Title The Structure and Function of Complex Networks Type Journal Article
Year 2003 Publication SIAM Review Abbreviated Journal SIAM Rev.
Volume 45 Issue 2 Pages 167-256
Keywords (up) networks; graph theory; complex systems; computer networks; social networks; random graphs; percolation theory
Abstract Inspired by empirical studies of networked systems such as the Internet, social networks, and biological networks, researchers have in recent years developed a variety of techniques and models to help us understand or predict the behavior of these systems. Here we review developments in this field, including such concepts as the small-world effect, degree distributions, clustering, network correlations, random graph models, models of network growth and preferential attachment, and dynamical processes taking place on networks.
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Corporate Author Thesis
Publisher Siam Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5214
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Author Wilson, D.A.; Stevenson, R.J.
Title The fundamental role of memory in olfactory perception Type Journal Article
Year 2003 Publication Trends in Neurosciences Abbreviated Journal Trends. Neurosci.
Volume 26 Issue 5 Pages 243-247
Keywords (up) olfactory perception mammals
Abstract Current emphasis on odorant physiochemical features as the basis for perception largely ignores the synthetic and experience-dependent nature of olfaction. Olfaction is synthetic, as mammals have only limited ability to identify elements within even simple odor mixtures. Furthermore, olfaction is experience-bound, as exposure alone can significantly affect the extent to which stimuli can be discriminated. We propose that early analytical processing of odors is inaccessible at the behavioral level and that all odors are initially encoded as `objects' in the piriform cortex. Moreover, we suggest that odor perception is wholly dependent on the integrity of this memory system and that its loss severely impairs normal perception.
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Publisher Place of Publication Editor
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Series Editor Series Title Abbreviated Series Title
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Notes Approved no
Call Number refbase @ user @ Serial 795
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Author Visser, E.K.; Van Reenen, C.G.; Engel, B.; Schilder, M.B.H.; Barneveld, A.; Blokhuis, H.J.
Title The association between performance in show-jumping and personality traits earlier in life Type Journal Article
Year 2003 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 82 Issue 4 Pages 279-295
Keywords (up) Personality; Performance; Horses; Prediction; Individual differences; Behavioural tests
Abstract For a horse to succeed in a show-jumping career, the individual has to possess both excellent physical abilities as well as a suitable personality to perform under challenging conditions. Forty-one Dutch Warmblood horses were used to develop personality tests and correlations between test variables and early training performances in jumping were studied. In behavioural tests, during the first 2 years of the horses' lives, personality aspects like emotionality, reactivity to human and learning abilities were quantified. At the age of 3, horses were broken and received early training in show-jumping. The inter-relationship between several performance variables measured during this early training phase were studied using principal component analysis (PCA). Variables measured in the different personality tests (novel-object test, handling test, avoidance-learning test and a reward-learning test) showed no correlations, suggesting that these tests all triggered different aspects of a horse's personality. This study indicates that it is possible to predict a substantial part of the show-jumping performance of an individual horse later in life by personality traits earlier in life.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number refbase @ user @ Serial 326
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Author Vieuille, C.; Berger, F.; Le Pape, G.; Bellanger, D.
Title Sow behaviour involved in the crushing of piglets in outdoor farrowing huts--a brief report Type Journal Article
Year 2003 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 80 Issue 2 Pages 109-115
Keywords (up) Pig-maternal behaviour; Crushing; Free-ranging; Welfare
Abstract This study focuses on maternal activities involved in the trapping of piglets by the sow's body in outdoor farrowing systems and examine the mother-piglet context leading either to the death of piglets or to their survival. The behaviour of six Large-WhitexLandrace sows and their litters was continuously video recorded at their first and second parity, during the 40 h following parturition. Crushing mainly occurred at evening and night, during the first 12 h of farrowing and involved changes between lying, sitting and standing positions, as well as between udder and side lying. No piglet died from savaging. Nevertheless, aggressive behaviours of sows were observed, particularly in their first maternal experience. The immediate context of trapping was related to the labour of the sow and to the feeding and resting of piglets. The immediate crushing context was related to active avoidance of restless piglets while lying down, as well as sitting and standing behaviours. These results are discussed in terms of differential reactions of the sow to suckling attempts of piglets.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 2235
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