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Author Krueger, K. isbn  openurl
  Title (up) “Pferdehaltung und Ethologie der Pferde” im Bachelorstudiengang Pferdewirtschaft Type Book Chapter
  Year 2014 Publication Forschendes Lernen initiieren, umsetzen und reflektieren Abbreviated Journal  
  Volume Issue Pages 54-81  
  Keywords  
  Abstract  
  Address  
  Corporate Author Thesis  
  Publisher UniversitätsVerlag Webler Place of Publication Bielefeld Editor : S. Lepp und C. Niederdrenk-Felgner  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN 10: 3-937026-91-6 Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5944  
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Author TRUMLER E openurl 
  Title (up) “Unternehmen Attrappe” Type Journal Article
  Year 1962 Publication Abbreviated Journal Jb. d. G. v. Opel-Freigeheges  
  Volume 3 Issue Pages 30-43  
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  Area Expedition Conference  
  Notes from Professor Hans Klingels Equine Reference List Approved no  
  Call Number Serial 1662  
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Author Schönfelder, W.D. isbn  openurl
  Title (up) “Wertschätzung” – Voraussetzung für erzieherisches Verhalten im Voltigier- und Reitunterricht Type Book Chapter
  Year 1982 Publication Sportlehre für Reiten.Fahren. Voltigieren Abbreviated Journal  
  Volume Issue Pages 51-60  
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  Corporate Author Thesis  
  Publisher FN-Verlag Place of Publication Warendorf Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN 3885420155 Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4450  
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Author Clement, T.S.; Feltus, J.R.; Kaiser, D.H.; Zentall, T.R. openurl 
  Title (up) “Work ethic” in pigeons: reward value is directly related to the effort or time required to obtain the reward Type Journal Article
  Year 2000 Publication Psychonomic bulletin & review Abbreviated Journal Psychon Bull Rev  
  Volume 7 Issue 1 Pages 100-106  
  Keywords Animals; Behavior, Animal/*physiology; Cognition/physiology; Columbidae; Discrimination Learning/physiology; Female; Male; Reinforcement (Psychology); *Reward; Time Factors  
  Abstract Stimuli associated with less effort or with shorter delays to reinforcement are generally preferred over those associated with greater effort or longer delays to reinforcement. However, the opposite appears to be true of stimuli that follow greater effort or longer delays. In training, a simple simultaneous discrimination followed a single peck to an initial stimulus (S+FR1 S-FR1) and a different simple simultaneous discrimination followed 20 pecks to the initial stimulus (S+FR20 S-FR20). On test trials, pigeons preferred S+FR20 over S+FR1 and S-FR20 over S-FR1. These data support the view that the state of the animal immediately prior to presentation of the discrimination affects the value of the reinforcement that follows it. This contrast effect is analogous to effects that when they occur in humans have been attributed to more complex cognitive and social factors.  
  Address University of Kentucky, Lexington, USA  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language English Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1069-9384 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:10780022 Approved no  
  Call Number refbase @ user @ Serial 248  
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Author Wasilewski A. url  openurl
  Title (up) 'Friendship' in ungulates? – Sociopositive relationships between non-related herd members of the same species Type Manuscript
  Year 2003 Publication Abbreviated Journal  
  Volume Issue Pages  
  Keywords  
  Abstract http://www.staff.uni-marburg.de/~z-phylog/wisstaff/wasi-sum.htm

Summary

* The quantitative study presented here demonstrates and verifies for ungulates the existence of social bonds, which in humans are termed 'friendships'. Whereas 'friendships' in non-human primates have become increasingly accepted over the past 30 years, their existence in non-primates remained as yet to be proved.

* The investigation was conducted on ungulates, because like primates the majority of their species are highly social; at the same time they are phylogenetically distant to the primates. To date, research into the social behaviour of ungulates in general, and of domestic species in particular, focuses predominantly on agonistic (socionegative) components, such as conflicts and resulting dominance hierarchies; sociopositive behaviours have been largely neglected.

As a consequence, previous evidence in favour of 'friendships' among ungulates remained largely anecdotal. Studies explicitly investigating sociopositive relationships among herd members are rare and typically confound bonds based on 'friendships' with reproductive relationships (kinship and sexually motivated bonds).

* Of each of the two large ungulate orders two species were investigated: horses and donkeys (Perissodactyla, Equidae) and sheep and cattle (Artiodactyla, Bovidae). The comparative, individual-based (and hence highly discriminative) longitudinal approach addressed all three levels of social organisation: associations, interactions and bond patterns within social units. Due to its systematic, quantitative, individual-based and comparatively broad design, this work may be considered to some degree a 'pioneering study'. As a consequence, a substantial amount of time and effort was required, particularly with respect to developing methods for the collection and analysis of data.

* Two reasons led to conducting this study on domestic ungulates: For farm animals information about who is related to whom is usually readily available, facilitating discrimination between friendship and kinship. Furthermore, successful proof of 'friendships' among ungulates would imply practical consequences for animal husbandry, since a social environment accommodating the animals' psychosocial needs is an essential prerequisite for their well-being (ethical reasons for animal protection) and ultimately results in higher productivity and hence economic efficiency (anthropocentric reasons for animal protection).

* Field research was conducted in England. Cattle and sheep were studied under commercial husbandry conditions, whereas appropriately large herds of horses and donkeys could only be found at a sanctuary.

* Two to three herds of each of the four species, namely cattle, sheep, donkeys and horses, comprising 11 to 60 members (average herd size approx. 25) were studied in two consecutive years (1996: 36 weeks, 1997: 28 weeks). The herds of cattle mainly consisted of subadult females, whereas those of the other species were predominantly composed of adults. Equids were kept in mixed-sex herds of mares and geldings (castrated males); the bachelor flocks of sheep consisted of 'intact' males (rams). The data presented here are based on 234 individuals and approximately 1500 hours of observation.

* Since even human social psychology provides neither a consistent definition nor a universally applicable terminology, it was necessary to establish a generally valid, precise definition of 'friendship'. Care was taken to exclude characteristics containing a bias as to the function of friendships (criteria were limited to those regarding contents and form). The definition contains all relevant criteria including those necessary for discrimination against alternative sociopositive relationships.



Friendship is defined as a voluntary, reciprocal, non-reproductive bond between individuals. It is primarily dyadic and carries a subjective value to both participants. Friendship is characterised by positive affect ('affection') and is expressed in a consistent inter-individual preference.

This definition is applicable to social relationships among both human and non-human animals, as well as interspecific relationships.

* Two types of parameters were used as indicators of 'friendship': frequency of spatial proximity (nearest neighbours; consistent neighbour preferences as a reflection of associations) and frequencies of different spontaneous as well as experimentally induced sociopositive interactions. For different reasons, of the four kinds of interactions (social grooming s.l., resting in physical contact, sharing limited feed, documenting behaviours s.l.) only grooming and feed sharing proved reliable.

* Aspects of the animals' spontaneous behaviour were recorded for all herds and in both years, whereas husbandry conditions demanded that complementary experiments be restricted to a few selected herds only. Data were collected using different Sampling and Recording Techniques, which were usually combined for maximum efficiency (e.g. Scan Sampling with nearest neighbour data, Ad libitum Sampling and Continuous Recording with sociopositive interactions).

* Data were subjected to an extensive statistical analysis. In addition to commonly used uni- and bivariate methods (Binomial and Chi-Square Goodness-of-Fit Tests, bivariate Correlations and Mantel Tests) a number of multivariate statistics were applied (Multidimensional Scaling, hierarchical Cluster Analysis, partial Correlations and multiple Mantel-Tests). Dyadic data are best represented as matrices; matrix data, however, pose the difficulty of partial interdependence, requiring Randomisation Tests (such as the one developed by Mantel) for their analysis. The complexity and amount of neighbour data required the development and optimisation of a 'customised' evaluation software (NENESYS = NEarest NEighbour SYstematic Standardisation programme) in close interdisciplinary cooperation with computer scientists and mathematicians.

* Seven different aspects of 'ungulate friendships' were addressed. The presented results differ in extent and depth.

1. Demonstration and verification of friendships in ungulates

2. Quantification of inter-individual preferences

3. Situation-specific aspects of these preferences

4. Dynamics and duration of friendships

5. Asymmetry within the relationships

6. Conditions enhancing friendships

7. Functions of animal friendships

* All four species expressed distinct inter-individual ('personal') preferences both for certain neighbours and for specific partners for sociopositive interactions. Endeavouring to present conservative and hence reliable assessments, only those neighbour preferences exceeding a threshold criterion were considered as associations. As the set-up of the study ensured that these associations met the criteria outlined in the definition of 'friendship' by excluding alternative bases for bonds (such as kinship and sexual motivation = reproductive relationships), the first explicit attempt at verifying the existence of friendship can be considered successful for all four species of ungulates examined. As a consequence, the concept of 'friendship' needs to be extended beyond the order of primates.

* Due to methodological reasons, the frequencies of interactions are only semi-quantitative and hence can be compared within herds only.

Frequencies of spatial proximity, on the other hand, provide a fully quantitative indicator, enabling direct comparisons across species, between different herds of the same species and between individuals. Due to the 'spatial limitations' of this thesis, it mainly focuses on inter-species comparisons.

With the exception of donkeys, the species could be arranged in sequence of increasing values of three parameters characterising their associative structure: maximum strength of association, maximum group size and proportion of herd members associated in distinct groups. Cattle showed the lowest values in all three parameters, sheep were intermediate, and horses reached the highest values. In these three species the strength of their associations was positively related to group size and the proportion of individuals associated in distinct groups. The donkeys did not fit this pattern: On the one hand, their association pattern comprised the lowest proportion of associated members and the smallest groups (pairs); on the other hand, these pairs exhibited very strong associations.

* Comparing inter-individual preferences across up to four different situations clear interspecific differences became evident. Some of these were greater between species of the same order than between orders.

The comparison of structures of association during grazing to those during resting, revealed two distinct patterns: Herds of horses and sheep flocks divided themselves into separate, usually mutually exclusive and independent groups and their associations corresponded greatly in both situations. Donkeys and cattle, on the other hand, formed diffuse, incomplete associative networks. The proportion of associations corresponding in both situations was far lower than in horses and sheep.

In all four species frequencies of social grooming s.l. (sheep are generally believed not to engage in social grooming, but see below) were compared to proximity frequencies during grazing and resting; and additionally to frequencies of feed sharing in the equid species.

Whereas horses restricted their sociopositive interactions entirely (grooming) or mostly (feed sharing) to members of their own group, cattle showed very little if any correspondence between partner preferences when grazing, resting and grooming. Donkeys spread their preferences widely across herd members, 'typically' approaching individuals for grooming with whom they were not associated during either grazing or resting. Considering their patterns of feed sharing, two different types of behaviour became evident: Pairs of donkeys that were closely associated during both grazing and resting, preferentially shared feed between them, whereas the remaining herd members did not discriminate between associates and non-associates when sharing feed. For sheep the findings of this study substantially increased the likelihood that unidirectional head rubbing and reciprocal cheek-to-cheek contact serve to strengthen 'personal' bonds and reduce tension – two functions generally attributed to social grooming. Both behaviours occurred mostly, but not exclusively, within groups of associated animals; these associations corresponded largely during grazing and resting.

* Results indicate that different types of social interactions and associations during different activities, respectively, may be particularly suited to initiate, consolidate and intensify friendships and occur particularly frequently during different phases of friendship formation and development. In horses, for instance, social grooming appears to some extent to be associated with the initiation of bonds, whereas resting in close proximity seems to strengthen newly formed relationships.

The longitudinal character of the study permitted to trace existing bonds over a maximum of 18 months. The duration of bonds in horses and sheep was comparable to findings of other authors for herds whose social networks resulted from a mixture of bonds based on both friendship and kinship; cattle relationships lasted shorter than described in the literature, while those in donkeys turned out to be approximately twice as long as had been documented to date.

* Aspects of asymmetry within dyadic relationships become particularly evident in grooming behaviours in bovids, which are typically performed unilaterally (social licking in cattle and head rubbing in sheep). Both behaviours can occur either spontaneously or as a consequence of solicitation.

* Addressing the question of factors potentially encouraging friendships, relationships between the degree of similarity with respect to some physical characteristics and nearest neighbour frequencies were found to be statistically significant. Cattle of similar age associated more closely, and horned rams preferred other individuals with horns as neighbours. The widely accepted preference of horses for associates of similar coat colour (and height) was not verified.

It remains to be decided whether (in addition to this preference for similar individuals) a preference based on complementary characteristics does exist.

* Following a critical review of the methodology employed, the benefits ungulates derive from friendships and possible applications for animal husbandry are discussed. Very little convincing evidence of a (direct) concrete, practical benefit was found. A psychological benefit in the form of social / emotional support, on the opposite, became highly probable.

Anecdotal evidence indicates, that situations causing a state of anxiety, may favour the initiation and intensification of friendships. Moreover, the study was able to statistically prove connection between (roughly) simultaneous introduction to a new herd (novel social environment) and persisting associations (as an indicator of friendships) in horses and sheep. These findings suggest that ungulate friendships provide a feeling of security. Social / emotional support among friends is predominantly given in a passive manner, e.g. in the form of – both literally and metaphorically – standing by a friend during conflicts with a third party, followed by social grooming with the friend. Social grooming has been shown by other authors to reduce physiological symptoms of stress. In addition, external signs of relaxation indicate a decrease in psychological tension. Hence, psychological support enhances the animal's physical health (and ultimately increases its biological fitness), providing an indirect practical benefit.

* Since all larger species of farm animals are highly social ungulates, the demonstration and verification of friendships implies consequences for husbandry practice. Intensive husbandry conditions are characterised by high animal densities, resulting in increased social stress, and grouping of animals of similar ages, which interferes with or prevents the formation and maintenance of bonds between kin. As friendships develop preferentially between individuals of similar age, they can be accommodated with relatively little effort when deciding about regrouping. The study presented here provides explicit recommendations to interested stockpersons on how to expeditiously and efficiently identify existing friendships and on which aspects to take into account when regrouping established herds or introducing new members.

* The different aspects of the study and their complex and often interwoven interrelations are condensed into a 'Regelkreismodell'. This model facilitates the generation of precise hypotheses for future research. Further detailed investigation into a variety of questions raised by this study seems in many ways promising and rewarding.

* The successful demonstration and verification of the existence of friendship in ungulates requires an extension of the 'concept of friendship' beyond the order of primates and encourages more detailed research into the bonding behaviour of other social vertebrates (e.g. elephants, dolphins, carnivores and also parrots).

* This extension demands a reinterpretation of some behaviours which so far have been unsatisfactorily or controversially classified, such as social head or horn rubbing in sheep, which has been mentioned occasionally, but received very little attention in the literature. Due to its occurrence during aggressive interactions, some authors consider horn or head rubbing to be agonistic. Others suppose the preorbital gland to play a significant role in this type of interaction and interpret head or horn rubbing as a 'respectful gesture' (adopting the dominant sheep's individual scent).

The study presents the first description of cheek-to-cheek contact in sheep. These cheek contacts are unequivocally sociopositive in nature. Since they are frequently associated with head rubbing, a third interpretation of head rubbing behaviour is offered by classifying both behaviours within the context of social grooming s.l.. More detailed examinations of these behaviours will be conducted as part of a research project into the evolution of the bovidae.
 
  Address  
  Corporate Author Thesis Ph.D. thesis  
  Publisher University of Marburg Place of Publication Marburg Editor  
  Language German Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
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  Notes Approved no  
  Call Number refbase @ user @ Serial 824  
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Author Tschudin, A.J.-P.C. url  openurl
  Title (up) 'Mindreading' Mammals? Attribution of Belief Tasks with Dolphins Type Journal Article
  Year 2001 Publication Animal Welfare Abbreviated Journal Anim Welfare  
  Volume 10 Issue Pages 119-127  
  Keywords  
  Abstract 'Mindreading' or theory of mind (ToM) refers to the capacity to attribute mental states to others. This ability is regarded as a critical component of what has, to date, exclusively characterized the advanced social cognition displayed by humans. The false belief task is a key test for ToM in different animal species. On a standard non-verbal false belief task, humans pass from age 4, whereas non-human primates consistently fail. Neuroanatomical and behavioural evidence for dolphins, however, indicates that they are capable of passing ToM tasks. The current paper represents a synthesis of the relevant dolphin research on neocortical evolution and non-invasive behavioural tests of precursors for ToM and the attribution of beliefs. The success of dolphins on attribution of belief tasks, in the absence of learning or cueing, indicates that they are capable of 'mindreading'. What are the implications of animal 'mindreading'? ToM tasks probe for reflexive consciousness and, by this criterion, dolphins may display reflexive consciousness. The implication of this conclusion is that future behavioural studies of social cognition will have considerable ethical and legal implications for animal welfare.  
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  Notes Approved no  
  Call Number refbase @ user @ Serial 3491  
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Author Nakagawa, S.; Waas, J.R. url  doi
openurl 
  Title (up) 'O sibling, where art thou?' – A review of avian sibling recognition with respect to the mammalian literature Type Journal Article
  Year 2004 Publication Biological Reviews of the Cambridge Philosophical Society Abbreviated Journal  
  Volume 79 Issue 1 Pages 101-119  
  Keywords Birds; Direct familiarisation; Indirect familiarisation; Individual recognition; Kin discrimination; Kin recognition; Mammals; Sibling recognition  
  Abstract Avian literature on sibling recognition is rare compared to that developed by mammalian researchers. We compare avian and mammalian research on sibling recognition to identify why avian work is rare, how approaches differ and what avian and mammalian researchers can learn from each other. Three factors: (1) biological differences between birds and mammals, (2) conceptual biases and (3) practical constraints, appear to influence our current understanding. Avian research focuses on colonial species because sibling recognition is considered adaptive where 'mixing potential' of dependent young is high; research on a wider range of species, breeding systems and ecological conditions is now needed. Studies of acoustic recognition cues dominate avian literature; other types of cues (e.g. visual, olfactory) deserve further attention. The effect of gender on avian sibling recognition has yet to be investigated; mammalian work shows that gender can have important influences. Most importantly, many researchers assume that birds recognise siblings through 'direct familiarisation' (commonly known as associative learning or familiarity); future experiments should also incorporate tests for 'indirect familiarisation' (commonly known as phenotype matching). If direct familiarisation proves crucial, avian research should investigate how periods of separation influence sibling discrimination. Mammalian researchers typically interpret sibling recognition in broad functional terms (nepotism, optimal outbreeding); some avian researchers more successfully identify specific and testable adaptive explanations, with greater relevance to natural contexts. We end by reporting exciting discoveries from recent studies of avian sibling recognition that inspire further interest in this topic.  
  Address Department of Biological Sciences, University Waikato, Private Bag 3105, Hamilton, New Zealand  
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  Notes Cited By (since 1996): 9; Export Date: 23 October 2008; Source: Scopus Approved no  
  Call Number Equine Behaviour @ team @ Serial 4567  
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Author Byrne R.W. doi  openurl
  Title (up) - Animal Cognition in Nature, edited by Russell P. Balda, Irene M. Pepperberg and Alan C. Kamil Type Journal Article
  Year 2000 Publication Trends in Cognitive Sciences Abbreviated Journal Trends. Cognit. Sci.  
  Volume 4 Issue Pages 73-73  
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  Call Number refbase @ user @ Serial 3480  
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Author Cords, M. openurl 
  Title (up) 13th Cong. Int. Primat Soc Type Miscellaneous
  Year 1990 Publication Abbreviated Journal  
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  Call Number Equine Behaviour @ team @ Serial 4925  
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Author Heptner Vg, openurl 
  Title (up) 2. Wildpferd, Tarpan Equus (Equus przewalski Poljakov 1881) In: Die Säugetiere der Sowjetunion Type Book Chapter
  Year 1966 Publication In: Heptner V G & Naumov N P (ed.) Jena Abbreviated Journal  
  Volume Issue Pages 858-874  
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  Notes from Professor Hans Klingels Equine Reference List Approved no  
  Call Number Serial 1181  
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