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Araba, B.D.; Crowell-Davis, S.L. |
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Title |
Dominance relationships and aggression of foals (Equus caballus) |
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1994 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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41 |
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1-2 |
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1-25 |
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aggression; dominance; horse; Equus caballus |
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Studied a herd of 15 Belgian brood-mares and 10 foals. Specific aspects of social structure studied were dominance-subordinance relationships, preferred associates, social spacing, aggression rates, the frequency of aggressions administered down the dominance hierarchy, and interactive play bouts. The rank order of the foals, both before and after weaning, was positively correlated with the rank order of their dams. There was also a significant relationship between a foal's rank and its total aggression or aggression rate per subordinate post-weaning. Higher ranking foals had higher rates of aggression. Over 80% of threats were directed down the dominance hierachy. The play-rank order of the foals, scored by the number of times foal left a play bout, was not significantly correlated with the rank order as scored by agonistic interactions. -from Authors |
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Dept Anatomy and Radiology, Univ of Georgia, Athens, GA, USA |
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790 |
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Lansade, L.; Bertrand, M.; Bouissou, M.-F. |
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Title |
Effects of neonatal handling on subsequent manageability, reactivity and learning ability of foals |
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2005 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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Volume |
92 |
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1-2 |
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143-158 |
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Horse; Neonatal handling; Manageability; Emotional reactivity; Learning-ability; Human-animal relationship |
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Behaviour is an important factor to be taken into account in the various uses of horses. Today horses are mainly used for sport and leisure activities. They should therefore be easy to manage, calm and not fearful. Early handling is known to improve manageability and learning ability and to reduce fearfulness in various species. It has become fashionable in the horse industry to use an early training procedure, referred to as “imprint training”, which is said to produce durable if not permanent effects. However, no studies concerning the long-term effects of such neonatal handling have been carried out in horses. The present study examines the short- and long-term effects of neonatal handling on manageability, general reactivity and learning ability of foals. Twenty-six Welsh foals were studied: 13 were handled daily for 14 days from birth and 13 were non-handled controls. The handling procedure consisted of fitting a halter, gently patting all parts of each foal's body, picking up feet and leading over 40 m. Two days, 3 months, 6 months and 1 year after the end of the handling period, foals underwent behavioural tests to measure their manageability and various aspects of their reactivity. The results showed that neonatal handling has only short-term effects on manageability: 2 days after the handling period, handled animals were significantly easier to handle than controls for the four parameters measured during this test (time to fit a halter, time to pick up feet, walk ratio that is time during which foal walks under constraint/total time measured during leading and number of defensive reactions). Two parameters (time to fit a halter and walk ratio) were still lower in handled foals than in non-handled foals 3 months later and only one 6 months later (walk ratio). One year later there was no difference between groups. In addition, there was no effect of handling on reactivity at any time of testing or in any of the tests (reaction to isolation from conspecifics, presence of a human, presence of a novel object and to a surprise effect). Finally, neonatal handling did not improve the spatial or discriminative learning abilities measured at 14 months of age. To conclude, the effects of neonatal handling are only temporary. |
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841 |
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Lansade, L.; Bertrand, M.; Boivin, X.; Bouissou, M.-F. |
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Title |
Effects of handling at weaning on manageability and reactivity of foals |
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2004 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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87 |
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1-2 |
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131-149 |
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Horse; Handling; Weaning; Manageability; Emotional reactivity; Fearfulness; Human-animal relationship |
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The horse's temperament, including its manageability and reactivity and/or fearfulness, is of importance as it can result in problems and can render horses unsuitable for inexperienced riders. Early experience, including handling during infancy, may influence the horse's adult behaviour and reduce its fear of humans and other potentially frigthening situations. In the various species studied, handling has generally been undertaken during the neonatal period. The aim of the present study was to test the effects of handling young horses around the time of weaning, a period which has been demonstrated to be effective in increasing ease of handling in cattle and goats. Sixteen Anglo-Arab foals were handled for 12 days either immediately following weaning (early handled: EH) or 21 days later (late handled: LH); eight additional non-handled foals served as controls (C). Handling consisted of haltering, gently petting all parts of the body, picking up feet and leading the foal over 120 m. During handling sessions, EH were easier to handle than LH: time taken to fit them with a halter, to pick up feet, and “walk-ratio” (time walking under constraint/total time walking) were significantly lower for EH. During subsequent tests conducted over 2 days, 4, and 7 months, as well as 10 months and to some extent 18 months after the end of handling period, EH and LH were easier to handle and less reactive than controls, although differences diminished with time. The period following weaning can therefore be qualified as an “optimal period” for handling. Some of the effects persist for at least 18 months. |
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842 |
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Houpt, K.A.; Fraser, A.F. |
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Przewalski horses |
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1988 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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21 |
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1-2 |
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1-3 |
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Equine Behaviour @ team @ |
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2302 |
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Klimov, V.V. |
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Spatial-ethological organization of the herd of Przewalski horses (Equus przewalskii) in Askania-Nova |
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Journal Article |
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1988 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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21 |
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1-2 |
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99-115 |
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The ethological structure of the herd of Przewalski horses includes hierarchic ranks of horses which determine their social roles in the herd. Besides the age ranks, the wild horses are characterized by the formation of harem groups, a “leading” group of females, a group of bachelor stallions, family groups, etc. The ethological structure determines the spatial one, which is the form of distribution of horses over the territory, and its assimilation and transformation into a system of informative spatial units. Under the influence of “internal” and “external” stimuli, the intragoup regulatory mechanims (social adaptations) manifest themselves, which allow the herd to function in the complicated situation of the reserve and allow humans to control the herd by using these mechanisms. There are grounds to believe that, given the balanced ethological structure of these groups, wild horses could be successfully acclimatized into natural biotopes. |
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Equine Behaviour @ team @ |
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2330 |
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Kolter, L.; Zimmermann, W. |
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Social behaviour of Przewalski horses (Equus p. przewalskii) in the Cologne Zoo and its consequences for management and housing |
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1988 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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21 |
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1-2 |
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117-145 |
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Between 1977 and 1986, two actual rank changes and two unsuccessful attempts occurred among the mares of the Cologne herd. The stallion was at first a low-ranking individual, but attained a dominant position during the last 3 years. At this time he started to split his group and thus to affect attachment relationships among his mares. During his absence of half a year, new bonds resulted and disappeared again some months after his return. Foals were tolerated by the sire for a long time. His behaviour to young, sub-adult mares varied with the individual. Protection of sub-adult mares by adult mares against the stallion's attacks may occur. Young mares protect and guard strange foals from their very first day. Management steps to cope with social and feeding problems consisted of enlargement and adding complexity to the enclosure, the establishment of more feeding sites, building a stable and temporary removal of the stallion. |
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Equine Behaviour @ team @ |
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2332 |
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Ryder, O.A.; Massena, R. |
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A case of male infanticide in Equus przewalskii |
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1988 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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21 |
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1-2 |
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187-190 |
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Following the introduction of a new stallion to a band of E. przewalskii mares two births, both of male foals, resulted in foal death due to injuries sustained in the first day of life. Neither foal was sired by the new herd stallion. The second foal death was the results of an observed attack on the newborn male and is described here. Subsequently births in the same enclosure and, in one instance, to the same mare whose previous foal was killed, were of foals sired by the new stallion and were uneventful, with 3 male foals surviving to date. |
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from Professor Hans Klingels Equine Reference List |
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yes |
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1539 |
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Boyd, L.E.; Carbonaro, D.A.; Houpt, K.A. |
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The 24-hour time budget of Przewalski horses |
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1988 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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21 |
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1-2 |
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5-17 |
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A herd of 8 Przewalski horses were observed on pasture in summer. Fifteen-minute focal animal samples were used to determine the time budget of the horses during the periods 00.00-04.00, 04.00-08.00, 08.00-12.00, 12.00-16.00, 16.00-20.00 and 20.00-24.00 h EDT. The behavioral states recorded were feeding (grazing and eating grain), nursing, drinking, standing, stand-resting, self-grooming, mutual grooming, locomoting, playing, and lying laterally and sternally. The average number of behavioral states occurring per hour, and the defecation, urination, aggression and vocalization rates were also determined. Overall, the horses spent 46.4 +/- 5.9% of their time feeding, 1.3 +/- 0.1% nursing, 0.5 +/- 0.1% drinking, 20.6 +/- 5.4% standing, 15.7 +/- 3.2% stand-resting, 1.7 +/- 0.2% self-grooming, 2.2 +/- 0.7% mutual grooming, 7.4 +/- 1.0% locomoting, 1.2 +/- 0.3% playing, 1.2 +/- 0.5% lying laterally and 4.1 +/- 3.0% lying sternally. The horses averaged 45.2 +/- 5.8 behavioral states per hour, and 0.2 +/- 0.0 defecations, 0.3 +/- 0.0 urinations, 1.5 +/- 0.3 aggressions and 0.7 +/- 0.1 vocalizations per hour. The horses spent the greatest amount of time foraging between 20.00 and 04.00 h, when the temperatures were lower. They spent 68.2 +/- 2.2% of their time between 20.00 and 24.00 h feeding, but only 31.2 +/- 2.1% of their time feeding between 08.00 and 12.00 h. Recumbent rest was most common between 00.00 and 04.00 h. As temperatures rose during the daylight hours, the horses spent more time drinking and standing, rather than grazing. Stand-resting was the most common form of rest during the day. The horses exhibited the greatest number of activities per hour from 08.00 to 20.00 h. While standing in close proximity to one another during these hours, the horses exhibited the highest number of aggressions per hour (1.9-2.4). |
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McGreevy, P.D.; Thomson, P.C. |
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Differences in motor laterality between breeds of performance horse |
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2006 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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99 |
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183-190 |
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Horse; Lateralisation; Laterality; Breed; Training |
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This study examined the relationship between motor laterality in horses bred for different types of work and therefore different temperaments. Foreleg preference during grazing was measured in three populations of domestic horse, Thoroughbreds (TB, bred to race at the gallop), Standardbreds (SB, bred for pacing) and Quarter Horses (QH, in this case bred for so-called “cutting work” which involves manoeuvring individual cattle in and out of herds). With a one-sample t-test, TBs showed strong evidence of a left preference in motor laterality (P = 0.000), as did SBs (P = 0.002) but there was no convincing evidence for laterality in QH (P = 0.117). However, the increasing trend in left preference from QH to SBs then TBs was associated with increasing differences between individual horses within a breed. The overall preference (either left or right) increased with age (P = 0.008) and the rate of increase varied with breeds. The presence of a higher proportion of left-foreleg preferent individuals in TBs and SBs compared with QH may indicate that their training or selection (or both) has an effect on motor bias. |
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Caanitz, H.; O'Leary, L.; Houpt, K.; Petersson, K.; Hintz, H. |
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Effect of exercise on equine behavior |
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1991 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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31 |
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1-12 |
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The effect of short periods of strenuous exertion, in this case treadmill exercise, on the subsequent behavior of Standardbred horses was examined. Six horses were exercised on a high-speed treadmill 4 or 5 days per week, for 3-4 miles (approximately 1.8 m s-1 for 3 min, 5 m s-1 for 12 min, 9 m s-1 for 3 min, 3 m s-1 for 3 min, 1.8 m s-1 for 3 min). The behavior of the horses was observed in the horse's home stall immediately after exercise and 2-7 h after exercise. Focal animal sampling for a total of 150 h revealed that the horses spent significantly more time drinking and less time resting after exercise than they did on control (non-exercise or rest days). The greatest influence on behavior was seen immediately after exercise. The horses spent 13.2+/-2.7 s per 15 min drinking after exercise and 7.2+/-2.3 s per 15 min drinking on non-exercise days. They spent 7.3+/-1.5 min h-1 stand resting after exercise and 9.7+/-2.1 min h-1 on non-exercise days. These changes in behavior may be related to the physiological changes that accompany exercise. Eating, walking, elimination and self-grooming were not significantly influenced by exercise. In a second experiment the activities of two groups of six Standardbred mares were compared. One group was exercised on the treadmill and the other was not. The exercised horses spent more time drinking and lying, but urinated less than the non-exercised group. |
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1989 |
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