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Author Christensen, J.W.; Keeling, L.J.; Nielsen, B.L. doi  openurl
  Title Responses of horses to novel visual, olfactory and auditory stimuli Type Journal Article
  Year 2005 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 93 Issue 1-2 Pages 53-65  
  Keywords Horses; Novelty responses; Fear; Behaviour; Heart rate  
  Abstract Responses of horses towards fear-eliciting stimuli can have important consequences for both human and horse safety. This experiment was designed to investigate behavioural and heart rate (HR) responses of horses to novel visual, auditory and olfactory stimuli. Twenty-four 2-year-old, previously unhandled, stallions were habituated to receive a food reward from a container in a test arena. Each horse was exposed to three 2 min tests in a balanced design where in addition to the feed container, either a traffic cone (visual test), white noise (auditory test) or eucalyptus oil applied to the inside of the container (olfactory test) were used as the novel stimuli. Compared to the control, less time was spent eating during all tests. There was no difference in locomotion activity in the different test situations, but presentation of the novel visual and auditory stimuli elicited significantly increased HR responses in the horses, compared to their response to the arena without novel stimuli (control), whereas there was no increase in HR response to the olfactory stimulus. However, during the olfactory test, the horses had an increased number of eating bouts and became more vigilant towards their surroundings, whereas during the visual and auditory tests, more time was spent alert towards the stimulus. The horses also took significantly more steps backwards in response to the auditory test. The heart rate responses correlated between tests and reflect a non-differentiated activation of the sympathetic nervous system, while the behavioural responses were linked to the type of stimulus.  
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  Call Number (up) Equine Behaviour @ team @ Serial 2256  
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Author Christensen, J.W.; Ladewig, J.; Sondergaard, E.; Malmkvist, J. url  doi
openurl 
  Title Effects of individual versus group stabling on social behaviour in domestic stallions Type Journal Article
  Year 2002 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 75 Issue 3 Pages 233-248  
  Keywords Horse; Individual stabling; Group stabling; Social behaviour  
  Abstract Domestic horses (Equus caballus) are typically kept in individual housing systems, in which they are deprived of physical contact. In order to study the effects of social restrictions on behaviour in young horses, nineteen 2-year-old stallions were housed either singly (n=7), or in groups of three (n=12) for 9 months. Subsequently, the stallions were released into two separate 2 ha enclosures according to treatment, and recordings were made on social interactions and nearest neighbours during a 6-week-period, 28 h per week. Previously group stabled stallions frequently had a former group mate as their nearest neighbour (P=0.001), whereas previously singly stabled stallions did not associate more with their former box neighbours, to whom physical contact was limited by bars during the previous treatment. The nearest neighbour was more frequently recorded to be within one horselength of singly stabled than of group stabled stallions (P=0.005). More aggressive behaviour was recorded in the group of previously singly stabled stallions, i.e. bite threats (P=0.032), whereas group stabled stallions tended to make more use of subtle agonistic interactions (displacements, submissive behaviour). Singly stabled stallions also responded to the 9 months of social deprivation by significantly increasing the level of social grooming (P<0.001) and play behaviour (P<0.001), when subsequently interacting freely with other horses. The increased occurrence may relate to a build-up of motivation (a rebound effect), as well as to external factors, such as playful pasture companions and the increased space allowance of the pasture. It is concluded that 2-year-old domestic stallions are sensitive to social deprivation and that stabling has long-term effects, lasting 6 weeks at least, on the social behaviour in stallions.  
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  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2257  
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Author Barber, J.A.; Crowell-Davis, S.L. url  doi
openurl 
  Title Maternal behavior of Belgian (Equus caballus) mares Type Journal Article
  Year 1994 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 41 Issue 3-4 Pages 161-189  
  Keywords Belgian; Draft horse; Ontogeny; Maternal aggression; Maternal behavior; Nursing behavior; Recumbency response  
  Abstract The relationship between ten Belgian mares and their offspring was studied from the first day of foal life to 17 weeks of age. Mares and foals spent more time at greater distances from each other as foals matured. Mares exhibited the recumbency response, being in closer proximity to their foals when foals were recumbent than when they were upright. Foals initiated the majority of nursing bouts. Frequency and duration of nursing bouts and percentage of time resting recumbently declined as foals matured. Foals also terminated the their foals, and they were most likely to do so in the first month of foal life. Maternal initiation of nursing. There was usually no discernible foal response to maternal aggression. Little difference between maternal behavior directed towards colts and fillies was found for all aspects of the study. Maternal behavior in the Belgian draft horse was similar to that reported for other equid breeds.  
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  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2268  
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Author Crowell-Davis, S.L. url  doi
openurl 
  Title Daytime rest behavior of the Welsh pony (Equus caballus) mare and foal Type Journal Article
  Year 1994 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 40 Issue 3-4 Pages 197-210  
  Keywords Horse; Behavior; Rest; Sleep; Pony; Foal  
  Abstract Upright and recumbent rest of 15 Welsh pony foals and their mothers was studied over a 2 year period. During their first week of life, the foals spent 32% of the time in recumbent rest. Subsequently, the percentage of time spent in recumbent rest decreased, but was still greater than for the foal's mother by Week 21, when the foals spent 6.5% of their time in recumbent rest. Adults spent little time in recumbent rest. Foals rested upright only 3.5% of the time during their first week of life. Mares rested upright more than foals did to Week 13, at which time peak values for time spent in upright rest occurred for both mares (32.5%) and foals (23%). Subsequently, mares and foals spent equal, but decreasing, amounts of time resting upright. The total time spent resting by the foals decreased gradually, and was characterized by a transition from recumbent rest to upright rest. Foals were more likely to be resting, either recumbent or upright, if their mother was resting upright. During the late spring, summer, and early autumn, mares and foals were most likely to be resting upright between 09:00 and 17:00 h.  
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  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2269  
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Author Pickerel, T.M.; Crowell-Davis, S.L.; Caudle, A.B.; Estep, D.Q. url  openurl
  Title Sexual preference of mares (Equus caballus) for individual stallions Type Journal Article
  Year 1993 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 38 Issue 1 Pages 1-13  
  Keywords Horse; Sexual behavior; Sexual preference; Vocalization  
  Abstract Eight mares were tested to determine if they remained near one of two stallions longer than would be expected if association was random. Six stallions were paired in 30 combinations and each mare was tested 30 times. The mares (Equus caballus) demonstrated a definite preference for individual stallions throughout the breeding season. This preference was influenced by the estrous state of the mare. During estrus, mares' preferences for stallions were positively correlated with the rate at which a given stallion vocalized. During diestrus, mares spent significantly less time in the proximity of stallions and did not exhibit any preference for individual stallions.  
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  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2270  
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Author Hall, C.; Crowell-Davis, S.L.; Warren, R.J. url  doi
openurl 
  Title Maternal and developmental behavior of the feral horses of Cumberland Island, Georgia Type Journal Article
  Year 1993 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 37 Issue 1 Pages 85  
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  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2271  
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Author Smith-Funk, E.D.; Crowell-Davis, S.L. url  openurl
  Title Maternal behavior of draft mares (Equus caballus) with mule foals (Equus asinus x Equus caballus) Type Journal Article
  Year 1992 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 33 Issue 2-3 Pages 93-119  
  Keywords  
  Abstract Draft mares and their mule foals were observed from the day of birth to Week 17 of each foal's life. The rate of nursing was recorded and the duration of nursing activity to the nearest second. The rate at which foals engaged in nursing activity varied at each age. The duration of nursing bouts varied slightly as the foals matured. Aggression was recorded during both nursing and non-nursing activity for both the mares and foals. Maternal aggression was highest during nursing activity, especially during the pre-nurse nuzzling period. Maternal aggression increased as the foals matured. Mother-directed foal aggression was primarily in response to maternal aggression. Spatial relationships between each focal dyad were recorded when the foals were upright, not nursing and when they were recumbent. Spatial relationships differed based on the foal's state. The activity in which the mare engaged while her foal was recumbent was recorded. The movements of the mares were also recorded during foal recumbency. Mares approached or maintained their distance from their recumbent foal more than they left their recumbent foal in all weeks of the study, except Week 2.  
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  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2272  
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Author Crowell-Davis, S.L.; Caudle, A.B. url  doi
openurl 
  Title Coprophagy by foals: recognition of maternal feces Type Journal Article
  Year 1989 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 24 Issue 3 Pages 267-272  
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  Abstract Six foals were each observed for 4 h per week during the first 6 weeks of life in an experimental situation in which they had access to feces taken from their mother and from another mare which was not pregnant or lactating. The foals sniffed at the feces equally. Two foals engaged in a total of seven bouts of coprophagy. All bouts of coprophagy involved maternal feces (χ2; P<0.01).  
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  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2273  
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Author Crowell-Davis, S.L.; Houpt, K.A.; Kane, L. url  doi
openurl 
  Title Play development in Welsh pony (Equus caballus) foals Type Journal Article
  Year 1987 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 18 Issue 2 Pages 119-131  
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  Abstract The structure of the play of colts and fillies living on pasture was studied from birth (n = 15) for up to 24 weeks. Foal play was categorized as running and bucking alone, running and bucking in a group, interactive (contact or combat) play, play with an object, and play at an adult. The rate of play decreased with increasing age and ambient temperature. Fillies and colts played with equal frequency, but engaged in some different types of play at different rates. There was no difference between colts and fillies in the proportion of play bouts of running and bucking in a group or playing with an object. Fillies engaged in running and bucking alone more than colts. Colts engaged in interactive play and play at an adult more than fillies. While there was no significant difference between colts and fillies in the duration of either type of running and bucking play, the interactive play bouts of colts were significantly longer than those of fillies. Both mares and stallions were tolerant of foal play which involved use of their body as a play object, including mounting play. Both fillies and colts engaged in mounting play. Foals used various natural objects found in the pasture for repeated bouts of play with inanimate objects, a behaviour which may explain, from a developmental perspective, the occasional use of “tools” in adult equids. The sex differences in type of play were consistent with the social structure of unmanaged adults in which males must compete with each other in order to associate with females.  
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  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2274  
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Author Crowell-Davis, S.L. url  doi
openurl 
  Title Self-grooming by mares and foals of the Welsh pony (Equus caballus) Type Journal Article
  Year 1987 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 17 Issue 3-4 Pages 197-208  
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  Abstract Self-grooming behaviour of 15 mare-foal pairs was studied weekly for the first 24 weeks of the foal's life. Mares self-groomed at mean rates of 1.2-2.2 times h-1 depending on the 4-week period. Foals self-groomed more often, with a peak rate of 12.3 times h-1 occurring during Week 5-8 and a subsequent decline to 6.0 times h-1 by Weeks 21-24. Self-grooming bouts of mares and foals also differed in the proportion devoted to each type of self-grooming. During a greater proportion of their total self-grooming bouts, foals scratched their head and neck with a hind limb or bit and scratched the trunk or hind limbs with the teeth. The mares spent a greater proportion of their self-grooming bouts rubbing their head or neck on an inanimate object, another pony, or their fore limb, or rolling on the ground. Particular types of individual self-grooming bouts were engaged in for different lengths of time. Foals had longer bouts of scratching the head and neck with a hind limb, scratching or biting the fore limb or hind limb with the teeth, and rubbing the head and neck on an inanimate object, while mares had longer bouts of rolling. Some of the differences in preference for particular types of self-grooming may be the result of differences in relative size of various body parts and, consequently, relative ease of a given type of self-grooming.  
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  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2275  
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