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Author Timney, B.
Title (up) Photopic Spectral Sensitivity and Wavelength Discrimination in the Horse (Equus caballus) Type Conference Article
Year 2008 Publication IESM 2008 Abbreviated Journal
Volume Issue Pages
Keywords
Abstract A number of studies have demonstrated that horses have chromatic vision and are able to distinguish colours from grey, independent of brightness. Anatomical and physiological data support the view that they are dichromats. In two experiments we provide behavioural evidence for their dichromacy. In the first we measured photopic spectral sensitivity. Using a spatial two alternative forced choice task, two horses were required to discriminate a coloured circular patch on an achromatic background from a blank field over a range of wavelengths . The obtained spectral sensitivity function was consistent with the presence of two cone classes with predicted spectral peaks at 429 nm and 545 nm, respectively. In the second experiment we obtained wavelength discrimination functions. The same animals were required to make discriminations between a series of standard wavelengths and a series of comparison wavelengths across the spectrum. The resultant wavelength discrimination function showed a single minimum at approximately 480 nm, which is also consistent with the view that horses are dichromats. Key Words: Horse, colour vision, spectral sensitivity, wavelength discrimination
Address
Corporate Author Timney, B. Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference IESM 2008
Notes Talk 15 min IESM 2008 Approved yes
Call Number Equine Behaviour @ team @ Serial 4463
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Author Wöhr, A.C.; Erhard, M.
Title (up) Polysonographic studies, about sleeping behaviour of horses Type Conference Article
Year 2008 Publication IESM 2008 Abbreviated Journal
Volume Issue Pages
Keywords
Abstract Objective: In the context of the ongoing discussion about keeping horses in individual stable boxes vs. in herds the lack of relaxation of the horse as a flight animal is an argument often put forward against individual housing. The long-term objective of our investigations is to determine the sleep phases in various housing systems in order to find a substantiated answer to that issue. For that purpose, the sleep waves measured by EEG have to be defined beforehand and allocated to the individual stages of sleep. The experiments described here are intended to support this effort. The objective is to establish a method which by means of a portable polysomnograph allows to capture the sleeping behaviour of horses for the purpose of defining the individual stages of sleep. It was investigated which stages of sleep horses undergo, and to what extent they may be comparable to those of humans.

Animals/materials/methods: Given the high technical effort involved, somnographic examination of large animals has so far been difficult and mostly required the animals to be sedated. Meanwhile, however, instruments such as the Somnoscreen by Somnomedics have become available. This is a completely portable polysomnograph with up to 28 channels and wireless online signal transmission and synchronous video transmission to a PC.

Using this instrument, the sleep profile of 5 Icelandic ponies and 10 horses (different race) aged 5-10 years was recorded and evaluated for 4 or 5 nights per horse. The following parameters were assessed: EEG (electroencephalogram), EOG (electrooculogram), EMG (electromyogram), ECG (electrocardiogram), thoracic and abdominal breathing motions as well as identification of the body posture. Synchronous video recordings were made. EEG recordings were obtained through gold-coated disk electrodes with long flexible cables, applied and secured to the scalp.

Results: As with humans, various stages of sleep can also be defined for horses using the above methods of recording. The waking condition is characterised by alpha waves, which just like in humans are within a range of 8-12 Hz. Typical REM phases as in humans were also detected, although not only when stretched completely on their side, as has hitherto always been described, but also when lying on their chest. Phases of deep sleep (stage 4) can also be measured, with the animals mostly in a standing position. The multi-stage human sleeping pattern, which is made up of 4-6 repeat phases of sleep (waking stage eyes open – waking stage eyes closed – REM phase – stage 1 – stage 2 – stage 3 – stage 4 – return to REM phase etc.) was found to be similar in horses in individual sequences. However, the sleep phases are shorter and more frequently interrupted by waking phases.

Conclusions: Horses are flight animals, which is why they have to be “on eye” in every situation so as to be able to flee in the face of danger. In a natural herd lying positions are only assumed if one or more members watch over the herd. In some publications the REM phase is treated as equivalent to the deep sleep phase. Although the REM phase is a phase of total muscle relaxation it is at the same time the dream phase and due to the high frequencies and the low amplitudes in the EOG resembles Stage I. This means that the sleeping horse can be awakened very quickly from this REM phase so as to be able to react to any dangerous situation. It therefore makes sense for the horses to assume a lying position during REM phases as the muscles are relaxed, yet a waking condition can be reached very quickly. A standing position seems to be preferred during deep sleep phases, where waking takes rather long, so that at least the position will not have to be changed. Whether the sleeping behaviour changes depending on age and race has yet to be investigated.
Address Institute of Animal Welfare, Ethology and Animal Hygiene, Ludwig-Maximilians University, Schwere-Reiter-Str. 9, 80637 Munich/Germany, woehr@lmu.de
Corporate Author Wöhr, A.C. Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference IESM 2008
Notes Talk 15 min IESM 2008 Approved yes
Call Number Equine Behaviour @ team @ Serial 4498
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Author Boross, B.; Maros, K.
Title (up) Possible indicators of the human-horse relationship among adult horses Type Conference Article
Year 2008 Publication IESM 2008 Abbreviated Journal
Volume Issue Pages
Keywords
Abstract According to scientists human-horse relationship is influenced by management conditions (Hausberger et al.; 2007) including handling methods (Henry et al.; 2005). Furthermore, Henry et al. (2005) show that mares foals“ behaviour toward humans may be influenced even by the mares. Although the familiarity of the testperson is said to be an important variable measuring relationship (Waiblinger et al.; 2006), other studies show that horses generalize from the experimenter to unfamiliar humans (Henry et al.; 2005) or from the caretaker to other human beings (Hausberger et al.; 2002).

Methods: 51 horses were observed in a 20x20 sandy arena (familiar to the subjects) to evaluate their relationship with a testperson.

First an arena test was carried out for 5 minutes. The horse was put into the arena alone. After that the person test came in which the testperson interacted with the horse.

The person test consisted of 3 or 4 different phases according to the horse”s behaviour. The phases were: (1.) voluntary animal approach test in which the testperson stood still and the subject was allowed to move wherever it wanted (Appr-vol); (2.) the testperson calling the horse to himself (Appr-call); (3.) the testperson making the horse stay beside him (Stand-still); (4.) the testperson making the horse follow him (without any tools and food reward) (Follow).

First 51 horses were tested with testpersons who were familiar to the animals. After that 39 horses were retested with unfamiliar testpersons within more than one week.

To analyze the potential effects of different human-related factors on the horse behaviour in the person test, such as time spent with training the horse (grooming, riding, working etc.), training methods, number of trainers etc. a questionnare has been used.

Results: Arena test: Horse behaviour was affected neither by their age (2-24 years) nor by their gender (30 geldings and 1 stallion vs 20 mares), however, mares spent more time in the entrance of the arena.

In the test with unfamiliar testpersons (retest) horses stood longer and walked less, which could reflect some habituation to the situation.

Person test: Gender and age of horses or time (years) elapsing since the horse started to be trained did not have any effect on the reactions of horses.

There was a positive correlation between the time (hours) spent with the horse weekly and the time of Follow.

The number of trainers had a significant effect on the Appr-vol and Follow responses: the less the number of the trainers, the sooner the horse approached the familiar testperson and the longer the animal followed him.

In the person test, significant difference was observed only in Appr-vol, thus horses followed the familiar testpersons much longer than the unfamiliar ones. Horses trained to follow their trainers without a lead rope followed significantly longer than animals without this kind of trainings.

Conclusion: Followership seemed to be the most sensitive behavioural response to human familiarity and it may also indicate some other aspects of horse-human relationship, however, training has a clear effect on this human related reaction.
Address Department of Organic Agriculture and Animal Welfare, Szent István University
Corporate Author Maros, K. Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference IESM 2008
Notes Talk 15 min IESM 2008 Fullpaper Approved yes
Call Number Equine Behaviour @ team @ Serial 4480
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Author Hothersall, B.
Title (up) Preliminary studies on visuo-spatial cue use in horses Type Conference Article
Year 2008 Publication IESM 2008 Abbreviated Journal
Volume Issue Pages
Keywords
Abstract A number of discrimination learning studies suggest that horses attend to spatial cues in preference to visual ones. However, global and relative spatial cues have generally been confounded so it is not clear what information horses actually used. There has also been no study of whether one cue is learnt at the expense of another, or whether multiple cues are encoded but prioritised differently. We therefore carried out a series of studies examining how horses use different cues in learning a food-finding discrimination task. All studies used a criterion for success of 7/10 on two consecutive days (binomial test, p=0.0004).

A pilot study showed that horses could discriminate between three buckets using visual cues (striped, chequered or plain patterns). All four subjects performed significantly above chance levels, although one horse’s performance did not meet the criterion. Three others completed the task in between two and 12 sessions. There were some indications that horses tried to solve the task using irrelevant spatial information.

Nine foals were then taught to find a food reward using the same three patterned buckets placed in a row in any corner of a test arena. In Stage 1 the rewarded bucket was signified both by visual cues (pattern) and by relative spatial cues (left, middle or right position). Foals took between 2 and 10 days to complete the task , whereupon the cues were separated. Foals then had to ignore the inappropriate cue in Stage 2. For the Spatial group (n=5), only relative position remained relevant; for the Visual Group (n=4) bucket pattern indicated the rewarded bucket. Spatial group foals completed Stage 2 faster than Visual group foals (Mann Whitney U, z=-2.71, p=0.008). Position cues seemed to overshadow learning about visual cues: Visual group foals persisted in responding to the previously correct position and none reached criterion within the maximum available time (15 days to complete both stages).

Finally, five adult horses and one 10 month old filly learnt to find a reward in one of three identical yellow buckets whose position in an array (left, middle or right) and location within the test arena were consistent. On reaching criterion, responses were recorded in two unrewarded probe trials where the buckets were either shifted along the same axis or rotated by 90º. When the array was shifted, five out of six horses chose the previously correct position (binomial test, p=0.018) that was now in a new location within the school. Choices were not significantly different from random when the array was rotated, suggesting that the horses had learnt something about global cues even if they did not use them in selecting a bucket.

At the small scale tested, relative spatial cues (position with regard to other goals) seemed to be more important to horses than either visual cues or global spatial cues, though the latter appeared to be encoded to some extent. Reliance on spatial cues may reflect their usefulness in the horse’s natural lifestyle in navigating over long distances and feeding on visually similar plants.
Address University of Bristol, Clinical Veterinary Science
Corporate Author Hothersall, B. Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference IESM 2008
Notes Talk 15 min IESM 2008 Approved yes
Call Number Equine Behaviour @ team @ Serial 4493
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Author Krueger, K. (ed)
Title (up) Proceedings of the International Equine Science Meeting 2008 Type Conference Volume
Year 2008 Publication IESM 2008 Abbreviated Journal
Volume Issue Pages
Keywords Equine Ecology; Equine Sociality; Equine Learning; Equine Cognition; Equine Welfare
Abstract Target group: Biologists, Psychologists, Veterinarians and Professionals

Meeting target: Because the last international meeting on Equine Science took place a couple years ago, there is an urgent need for equine scientists to exchange scientific knowledge, coordinate research provide knowledge for practical application, and discus research results among themselves and with professionals who work with horses. Additionally, dialog concerning the coordination of the study “Equitation Science” in Europe is urgently needed. Coordination and cooperation shall arise from the meeting, enrich the research, and advance the application of scientific knowledge for the horses` welfare.
Address
Corporate Author Thesis
Publisher Xenophon Verlag Place of Publication Wald Editor Krueger, K.
Language English Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN 978-3-9808134-0-2 Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 4508
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Author Walzer, C.; Kaczensky, P.; Ganbaatar, O.; Stauffer, C.
Title (up) Przewalski Horses, Satellites and Wild Asses Type Conference Article
Year 2008 Publication IESM 2008 Abbreviated Journal
Volume Issue Pages
Keywords
Abstract The Przewalski`s horse (Equus f. przewalskii) was extinct in the wild by the mid-sixties of the past century. The species only survived due to captive breeding from 13 founder individuals. In 1992 a reintroduction program was initiated in the Great Gobi B Strictly Protected Area in southwestern Mongolia. During it“s initial years, the project concentrated exclusively on P-horses. In the past decade activities have expanded significantly. The ecological project start point can be summed up as follows: i) species is extinct in the wild, ii) severe bottleneck, iii) practically no ecological data prior to extinction, iv) released into an extremely harsh, highly variable and poorly understood environment. The ecological knowledge constraints were compounded by i) highly emotional species, ii) simultaneous and competitive projects, iii) logistic nightmare, iv) non-scientific basis in planning phase. By the late 1990s project leadership and management was overhauled with research and scientific data firmly integrated into the decision-making process. Early scientific input concentrated on determining causes of death and low reproductive rates. The elucidation of the effects of endemic piroplasmosis on the population and subsequent management changes lead to remediation this deadly problem. P-horses have been fitted with ARGOS and GPS-ARGOS collars in order to determine home range and habitat preferences. Simultaneously the Mongolian wild ass and the wolf have been studied with these methods in the shared habitat. Satellite-based technologies provide the backbone for all habitat related project issues. At the onset (digitized Russian maps) data collection was restricted to the Eastern part of the Gobi B. Subsequently the spatial scale encompasses the entire Gobi Region in Mongolia and Northern Xingjian in China (e.g. Landsat, MODIS, NOAA, SRTM). Research has also focused on the role, needs and possible impacts of local semi-nomadic herders that use the protected area. Capacity building and training workshops (e.g. construction of fuel efficient stoves, felting) have been initiated. In 2007 a trans-boundary project in collaboration with the Xingjian Institute of Ecology and Geography of the Chinese Academy of Sciences, was initiated. This project aims to support rural communities of nomadic pastoralists living in the trans-boundary area of the Dzungarian Gobi, in China and Mongolia. Today, this project and the one in Hustain Nuruu (Mongolia) are the only ones that have resulted in free-ranging non-supplemented populations. In the Gobi B area some 120 (status 05.2008) P-horses roam in the protected area. In 2003 the IUCN downlisted the Przewalski\'s horse from to . Further downlisting to is predicted to occur in 2011. There is no consensus on when a reintroduction program is deemed successful. Clearly viewing the self-sustainable re-establishment of a population as a successful end-point is at best a short-term approach, constrained by time (today and now). Comprehensive interdisciplinary monitoring and research was and is the foundation for management strategies and decisions in this project. However, a self-sustaining financial base in conjunction with dedicated training and empowerment of local scientists and residents constitute essential prerequisites for the project”s future. Defining success and thereby inferring an end-point can easily lead to complacency compromising species persistence. As others have stated the ultimate project objective must be a constantly re-evaluated state of population persistence without intervention.
Address Research Institute of Wildlife Ecology University of Veterinary Medicine Vienna, Austria and International Takhi Group, Takhin Tal, Mongolia
Corporate Author Walzer, C. Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference IESM 2008
Notes Invited speaker IESM 2008 Approved yes
Call Number Equine Behaviour @ team @ Serial 4459
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Author Kaczensky, P.; Ganbaatar, O.; von Wehrden,H.; Walzer, C.
Title (up) Przewalski`s horses (Equus ferus przewalskii) and Asiatic wild asses (Equus hemionus): Similar Species, Same Habitat – Same Use? Type Conference Article
Year 2008 Publication IESM 2008 Abbreviated Journal
Volume Issue Pages
Keywords
Abstract Historic overlap zones of wild equids were small in Africa but extensive for Przewalski`s horses and Asiatic wild asses in Asia. Currently the Great Gobi B Strictly Protected Area in SW Mongolia is the only place where sympatric, free-ranging populations of these equids occur. This provides an unique opportunity to examine the co-existence of these little studied species and test the hypothesis that Przewalski`s horses are primarily adapted to mesic steppe habitats, whereas Asiatic wild asses are adapted to arid desert steppes and semi-deserts.

We monitored 9 Przewalski`s horses and 7 wild asses with satellite telemetry and superimposed the data on a habitat map derived from remote sensing (LANDSAT TM & ETM+-data) and ground sample plots. We tested for habitat preferences comparing use and availability with a logistic regression mixed model approach. Individuals were treated as random factors. Factor variables were tested for significant differences in subsequent Tukey post-hoc tests. Przewalski`s horses had non-exclusive home ranges of 152-826 km² and heavily selected for the most productive riparian plant communities. Asiatic wild asses also had non-exclusive home ranges, but with 4,449-6,835 km² they were 10 times larger than those of Przewalski`s horses. Asiatic wild asses seem to use plant communities more or less relative to their availability. Our results provide evidence for two parallel resource selection strategies. Our findings indicate that the Gobi areas provide an edge, rather than an optimal habitat for Przewalski`s horses. This leaves only small and isolated pockets of suitable habitat for future re-introductions. Asiatic wild asses, on the other hand, need access to large tracts of land to cope with the unpredictable resource distribution of the Gobi. Thus, Asiatic wild ass conservation requires a large scale approach.
Address
Corporate Author Kaczensky, P. Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference IESM 2008
Notes Talk 15 min IESM 2008 Approved yes
Call Number Equine Behaviour @ team @ Serial 4490
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Author Krueger, K.
Title (up) Social cognition and social learning in horses Type Conference Article
Year 2008 Publication IESM 2008 Abbreviated Journal
Volume Issue Pages
Keywords
Abstract In comparison to other social animals it appears to be far more difficult to show social learning in horses, although the social background of the animals prompts the prediction that social learning should be verifiable. This demands for a thorough analysis of conditions, context and effects of social learning which might not be as obvious in horses than in other social animals. Moreover, the horse“s social cognition in general, i.e. the processing, encoding, storage, retrieval, and application of social information, needs to be examined. In previous studies we addressed the horse”s perception of it“s social environment. Our results indicate that domestic horses are not only capable of distinguishing between familiar and unfamiliar horses but also of recognizing the social status of familiar horses relative to their own. Furthermore, they extract information from observing interactions between a familiar horse and a human experimenter, and adjust their behaviour according to the observed horse”s reaction and relative dominance status when they are themselves later confronted with the experimenter in a similar situation. Additionally, we showed in pilot experiments that horses paid more attention to the gaze of dominant horses from their own social group than to that of unknown or subordinate horses. In another study we suggested that horses of both sexes are capable of determining the social affiliation, the sex and, for their own group members, the identity of individuals, by sniffing their faeces. Hence, social affiliation and dominance relationships seemed to have a major influence on the horses` motivation to pay attention to their conspecifics. In the future we will take another step towards linking the horse“s social cognitive capacities with its social ecology, social structure, and individuality. In the context of comparative cognition research on the socially complex primates, or on a range of carnivores such as corvids, canids or dolphins, horses are particularly interesting, because they are highly social, but do not rely on social hunting or cooperative raising of offspring. Also foraging is simple for horses, predator avoidance is demanding and this may be the main reason for the horse”s complex social lives and their choice of learning tactics. Hence, we will investigate the horse"s investigative behaviour towards neutral, fear-eliciting and food related novel objects. We propose that, as in other species, sociality and distinct personalities are key features for the understanding of cognitive capacities and the choice of learning strategies.
Address University of Regensburg – Biology I
Corporate Author Krueger, K. Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference IESM 2008
Notes Talk 15 min IESM 2008 Approved yes
Call Number Equine Behaviour @ team @ Serial 4462
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Author Jørgensen, G. H.M.; Borsheim, L.; Mejdell, C. M.; Bøe, K. E.
Title (up) Social interactions and spacing in horses (Equus caballus) grouped according to gender Type Conference Article
Year 2008 Publication IESM 2008 Abbreviated Journal
Volume Issue Pages
Keywords
Abstract The aim of this study was to investigate the social behaviour and spacing between horses when grouped according to gender in homogeneous- or heterogeneous groups, and to see if the amount and severity of aggression differed in relation to gender composition.

A total of 66 horses divided in six batches were used. In each batch, horses were allotted into one mare group, one gelding group and one mixed gender group, with most groups consisting of three or four animals. Three batches were performed on Icelandic horses and three batches on riding horses and -ponies. After 4-6 weeks of acclimatisation, a trained observer recorded all social interactions using direct, continuous observation one hour in the morning and one hour in the afternoon for three consecutive days. In addition, the nearest neighbour of each horse was recorded using instantaneous sampling every 10 minutes. The horses were inspected for injuries before grouping, day one after grouping and after 4-6 weeks. Furthermore, a novel object test and a handling test (taking horse out of group) were conducted after the behavioural observations were completed.

No significant effect of gender composition was found on social interactions, spacing or injuries. Most of the aggressive interactions recorded were threats, and not aggressive interactions involving physical contact. Aggression seemed to increase with decreased space per animal and more horses per feeding place. Very few injuries were found and most were superficial. Horses showed a low degree of fearfulness in the novel object test and most horses were easy to handle in the “horse out of group test”.

In conclusion, gender composition does not seem to have any effect on aggression level, spacing or injuries in social groups of horses.
Address 2National Veterinary Institute, Department of Animal Health, P.O. Box 750 Sentrum, 0106 Oslo, Norway.
Corporate Author Jørgensen, G. H.M. Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference IESM 2008
Notes Talk 15 min IESM 2008 Approved yes
Call Number Equine Behaviour @ team @ Serial 4489
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Author Klingel, H .
Title (up) Social Organisation of the Equids Type Conference Article
Year 2008 Publication IESM 2008 Abbreviated Journal
Volume Issue Pages
Keywords
Abstract Two distinct types of social organisation have evolved in the Equidae. Type 1: In Horse Equus przewalskii, Plains Zebra E. quagga and Mountain Zebra E. zebra the adults live in non-territorial, stable, one-male families and as single bachelors and in bachelor groups. Family stallions have the exclusive mating rights with the mares in their harems. These consist of up to 6 unrelated mares plus their offspring, totalling up to 20 members. Mares stay in their harems until death. Stallions' tenure is from age 5-6 years, i.e. when they succeed in controlling a harem, for close to life time, but are replaced when dead or incapacitated. Harems are stable even in the absence of a stallion, indicating voluntary membership. Adolescent mares leave their parental families to become members of another harem. In Plains Zebra the adolescent mares are abducted, during an oestrus, by suitors who fight the defending family stallion/father. Successful stallions are bachelors who start a family, or family stallions enlarging their harem. Young stallions leave their parental families voluntarily at age 2-3 years and join bachelor stallion groups from where the family stallions are recruited. An individualised dominance hierarchy excists with the stallion in the alpha position. It is based on individual knowledge and recognistion of the members. Type 2: In Grevy's Zebra E. grevyi, African Wild Ass E. africanus and Asiatic Wild Ass E. hemionus adult stallions monopolise territories in which they have the exclusive mating rights. Stallions are tolerant of any conspecifics entering their territory. Bachelor stallions behave subordinately – or fight for the possession of the territory which is a prerequisite for reproduction. Mares join up to form anonymous and unstable groups or herds. The only stable unit is of a mare and her offspring. In Grevy's Zebra mares with foal join preferentially conspecifics of the same soial status, as do mares without foal. Matings take place inside the territory. There is no lasting relationship of the mare with a particular stallion, and the mare may be mated by any stallion whose territory she is visiting. Territories measure up to 10 or more square kilometres, and tenure is for several years. Grevy Zebra territorial owners leave their territories for a few hours to visit a water hole, or for months when grazing and water conditions are below requirements, and re-occupy it upon return, unchallenged. Except for a few small populations, the extant equids live in semi-arid to arid regions where environmental conditions force them to migrate seasonally or opportunistically. The ranges of the various species overlap: Mountain Zebra and Plains Zebra in South Africa and Namibia, Plains Zebra and Grevy's Zebra in Kenya and Ethiopia, Grevy's Zebra and African Wild Ass in Ethiopia, Asiatic Wild Ass and Przewalski Horse in Mongolia andChina. Although, in the overlap zones, individuals of different species are using the same resources like water and grazing simultaneously and next to each other, they rarely make closer contacts. Infanticide has been recorded in captive situations.
Address Technische Universität Braunschweig
Corporate Author Klingel, H . Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference IESM 2008
Notes Invited speaker IESM 2008 Approved yes
Call Number Equine Behaviour @ team @ Serial 4457
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