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| Author |
Crowell-Davis, S.L. |
| Title |
Self-grooming by mares and foals of the Welsh pony (Equus caballus) |
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Journal Article |
| Year |
1987 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
17 |
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3-4 |
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197-208 |
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Self-grooming behaviour of 15 mare-foal pairs was studied weekly for the first 24 weeks of the foal's life. Mares self-groomed at mean rates of 1.2-2.2 times h-1 depending on the 4-week period. Foals self-groomed more often, with a peak rate of 12.3 times h-1 occurring during Week 5-8 and a subsequent decline to 6.0 times h-1 by Weeks 21-24. Self-grooming bouts of mares and foals also differed in the proportion devoted to each type of self-grooming. During a greater proportion of their total self-grooming bouts, foals scratched their head and neck with a hind limb or bit and scratched the trunk or hind limbs with the teeth. The mares spent a greater proportion of their self-grooming bouts rubbing their head or neck on an inanimate object, another pony, or their fore limb, or rolling on the ground. Particular types of individual self-grooming bouts were engaged in for different lengths of time. Foals had longer bouts of scratching the head and neck with a hind limb, scratching or biting the fore limb or hind limb with the teeth, and rubbing the head and neck on an inanimate object, while mares had longer bouts of rolling. Some of the differences in preference for particular types of self-grooming may be the result of differences in relative size of various body parts and, consequently, relative ease of a given type of self-grooming. |
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Equine Behaviour @ team @ |
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2275 |
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Crowell-Davis, S.L.; Houpt, K.A.; Carini, C.M. |
| Title |
Mutual grooming and nearest-neighbor relationships among foals of Equus caballus |
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Journal Article |
| Year |
1986 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
15 |
Issue |
2 |
Pages |
113-123 |
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A 3-year study was carried out on the developmental behavior of foals from birth to 24 weeks of age and the behavior of mares living with foals. Mutual-grooming partners of foals were primarily other foals. The peak frequency of mutual grooming occurred during Weeks 9-12, when fillies mutual-groomed 1.6 times h-1 and colts mutual-groomed 0.9 times h-1. Fillies mutual-groomed more frequently than colts (P < 0.025). Fillies mutual-groomed randomly with colts and other fillies (P < 0.05), whereas colts mutual-groomed almost exclusively with fillies (P = 0.03). At all ages studied, if a foal's nearest neighbor was not its mother, it was more likely to be another foal than would be expected if the foal was associating randomly with non-mother ponies. Fillies were more likely than expected to have a filly rather than a colt as their nearest neighbor (P = 0.01). Thus, during their first few months of life, the foals studied exhibited patterns of behavior which were consistent with the development of the usual social milieu of unmanaged adults, in which several mares form a cohesive herd with one or more stallions associating with them. |
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Equine Behaviour @ team @ |
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2276 |
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Kaseda, Y.; Khalil, A.M. |
| Title |
Harem size and reproductive success of stallions in Misaki feral horses |
Type |
Journal Article |
| Year |
1996 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
47 |
Issue |
3-4 |
Pages |
163-173 |
| Keywords |
Harem size; Horse social organization; Reproduction; Blood types; Misaki feral horse |
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Over a 16-year period (1979-1994), long-term investigations were carried out on 14 Misaki feral stallions to analyze changes in harem size and the reproductive success. Harem size changed with the age of the stallions. Most stallions formed harem groups with four to five mares at the age of 4-6 and then the number of mares increased rapidly to the maximum at the age of 6-9 years. Thereafter, harem size decreased gradually to a minimum with advancing age. The harem size of 60 stable harem groups ranged from 1 to 9, and the average varied from a minimum mean of 1.8 in 1988 to a maximum mean of 5.3 in 1982. Mean harem size increased as adult sex ratio increased and a significant and positive correlation was found between them. One hundred and ninety-eight sire-foal pairs were determined by a paternity test with blood types and consort relations between stallions and mares during the study period. Out of 99 foals which were born in the stable harem groups, the true sires of 84 foals (85%) were the harem stallions in which the foals were born but the remaining 15 foals (15%) were sired by other harem stallions. Two out of three stallions which were studied throughout their lifetime produced 24 and 25 foals in 10 and 11 years of their reproductive lifespan, respectively. Another one produced only five foals in 6 years. The number of foals sired by the harem stallions was less than two over harem size 7 and some of the foals born in the harem were sired by other harem stallions. These results suggest that if a particular stallion monopolizes too many mares, he could not sire so many offspring because he could not always prevent his rival stallions from mating with his mares in wild or feral circumstances. |
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Equine Behaviour @ team @ |
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2308 |
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Rapaport, L.; King, N.E. |
| Title |
The behavioral research program at the Washington Park Zoo |
Type |
Journal Article |
| Year |
1987 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
18 |
Issue |
1 |
Pages |
57-66 |
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For the past decade, the Washington Park Zoo, Portland, Oregon, has had an active behavioral research program. The research department is both a zoo-supported research facility for visiting researchers and staff, and an educational facility that teaches practical behavioral research methods to undergraduates. The research education program utilizes students from any of a dozen local colleges and universities. Students receive academic credit for their participation. Active keeper-participation plays a major role in many research projects. Not only does keeper-cooperation facilitate research, but their knowledge of the individual animals often proves invaluable. In addition to involvement in student projects, keepers have also conducted their own research projects. |
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Equine Behaviour @ team @ |
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2324 |
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Lamoot, I.; Callebaut, J.; Demeulenaere, E.; Vandenberghe, C.; Hoffmann, M. |
| Title |
Foraging behaviour of donkeys grazing in a coastal dune area in temperate climate conditions |
Type |
Journal Article |
| Year |
2005 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
92 |
Issue |
1-2 |
Pages |
93-112 |
| Keywords |
Grazing behaviour; Habitat use; Donkey; Equid; Diet composition |
| Abstract |
A small herd of donkeys was introduced in a coastal dune reserve `Houtsaegerduinen' (ca. 80 ha) in Belgium, in order to slow down expansion of dominant grass and shrub species. The Houtsaegerduinen is a nutrient poor scrub-dominated dune system with a spatially heterogeneous vegetation pattern. Different aspects of the grazing behaviour (grazing time, bite rate, habitat use, diet composition) of the free-ranging donkeys are described and analysed. Behavioural data (of maximum six adult mares) were collected through continuous focal animal observation in three consecutive years (1998-2001). Temporal variation in grazing time, habitat use and diet composition was determined. During daylight, donkeys spent most of their time on grazing (56%). In all 3 years, grazing time was significantly shorter in summer (45% of their time), longest grazing times were achieved in spring (64%). In spring, the donkeys also achieved the highest bite rate (21.5 bites/min). The grassy habitat was preferred for foraging in all seasons, while the use of scrub and woodland was variable over time. Averaged over the four seasons, the general diet consisted for 80% of graminoids, 10% of forbs and 10% of woody plants. However, diet composition varied not only among seasons and years, but depended also on the foraged habitat type. We discuss the possible role of the donkeys in nature management. |
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Equine Behaviour @ team @ |
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2338 |
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Lamoot, I.; Callebaut, J.; Degezelle, T.; Demeulenaere, E.; Laquiere, J.; Vandenberghe, C.; Hoffmann, M. |
| Title |
Eliminative behaviour of free-ranging horses: do they show latrine behaviour or do they defecate where they graze? |
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Journal Article |
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2004 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
86 |
Issue |
1-2 |
Pages |
105-121 |
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Equids; Faeces avoidance; Grazing behaviour; Spatial differentiation; Urine; Horse marking |
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In contrast to horses in pastures, it is thought that free-ranging horses do not perform latrine behaviour, i.e. a behavioural pattern whereby the animals graze and defecate in separate areas. However, few studies deal with this particular subject, reporting contrasting conclusions. We hypothesize that horses free-ranging in large heterogeneous areas do not perform latrine behaviour. Thus, we believe that grazing and elimination behaviour are spatially related: where horses graze, they will also defecate. Behavioural data were collected from Konik horses, Haflinger horses, Shetland ponies and donkeys, grazing in different nature reserves (54-80 ha). Data for the different equids were analyzed separately, as well as data for mares and stallions (Konik and donkey stallions only). We investigated the proportion of the number of defecations/urinations while grazing on the total number of defecations/urinations; furthermore, we searched for the sequence of behaviours representing latrine behaviour in the strict sense. Additionally, we analyzed the correlation between grazing behaviour and eliminative behaviour on both vegetation type level and patch level. All the female equids often continued grazing while defecating. During urination, grazing ceases in the majority of instances. Cases where a mare terminated grazing in a certain vegetation type and sward height to eliminate in another vegetation type or in another sward height within the same vegetation type were rarely observed. On the vegetation type level as well as on the patch level, there was a highly significant (P<0.001) positive correlation between grazing time and number of eliminations (or eliminating time). The high values of the correlation coefficients (in case of the defecation variables r ranges between 0.553 and 0.955; in case of the urination variables r ranges between 0.370 and 0.839) illustrate that the spatial distribution of the eliminative behaviour can be explained to a high degree by the spatial distribution of the grazing behaviour. Results in the case of the stallions are preliminary, but indicate the same pattern. Horses, free-ranging in large heterogeneous areas, do not perform latrine behaviour, but defecate where they graze. Possibly, animal density is of major importance to explain this behavioural difference with horses in pastures. We suggest that also spatial vegetation heterogeneity and plant productivity of the grazed area, as well as parasite status of the grazing animals could play a role. |
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Equine Behaviour @ team @ |
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2339 |
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Moehlman, P.D.; Kebede, F.; Yohannes, H. |
| Title |
The African wild ass (Equus africanus): conservation status in the horn of Africa |
Type |
Journal Article |
| Year |
1998 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
60 |
Issue |
2-3 |
Pages |
115-124 |
| Keywords |
Equus africanus; Critically endangered; Extinction |
| Abstract |
From 1989 to 1996, surveys were made in most of the historic range of African wild asses in Somalia, Ethiopia, and Eritrea. From the 1970s to the mid 1990s populations of African wild asses (Equus africanus, Fitzinger, 1857) in Somalia and Ethiopia have declined from approximately 6 to 30 per 100 km2 to 1 or 2 per 100 km2. Given the current IUCN criteria, they are Critically Endangered (CR) and face extremely high risk of extinction in the wild in the immediate future, as their populations have been reduced by at least 80% over the last 10+ years (IUCN, 1994). Basic research is needed on this species as scientific information on its reproductive biology, behavior, ecology, and genetics is very limited. Improved support needs to be provided to existing parks and reserves and new multiple use reserves need to be established. |
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Equine Behaviour @ team @ |
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2380 |
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Moehlman, P.D. |
| Title |
Behavioral patterns and communication in feral asses (Equus africanus) |
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Journal Article |
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1998 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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60 |
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2-3 |
Pages |
125-169 |
| Keywords |
Equids; Feral asses; Behavior patterns; Facial expressions; Postures; Locomotion |
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The behavior of feral populations of the African wild ass (Equus africanus) were studied in the Northern Panamint Range of Death Valley National Monument for 20 months from 1970 to 1973 [Moehlman, P.D., 1974. Behavior and ecology of feral asses (Equus asinus). PhD dissertation, University of Wisconsin, Madison, 251 pp.; Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Natl. Geogr. Soc. Res. Reports, 1970: 405-411]. Maintenance behavior is described and behavior sequences that were used in social interactions are quantified by sex and age class. Agonistic, sexual, and greeting behavior patterns are described and analyzed in conjunction with the responses they elicited. Mutual grooming mainly occurred between adult males, and between females and their offspring. Five types of vocalizations were distinguished: brays, grunts, growls, snorts, and whuffles. A second population was studied for 1 month on Ossabaw Island, GA (Moehlman, 1979). This population had more permanent social groups and had a higher rate of mutual grooming and foal social play. |
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Equine Behaviour @ team @ |
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2381 |
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Moehlman, P.D. |
| Title |
Feral asses (Equus africanus): intraspecific variation in social organization in arid and mesic habitats |
Type |
Journal Article |
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1998 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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60 |
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2-3 |
Pages |
171-195 |
| Keywords |
Equids; Feral asses; Social organization; Mating systems; Intraspecific variation |
| Abstract |
Feral asses have been studied in the arid habitats of the southwestern United States [Moehlman, P.D., 1974. Behavior and ecology of feral asses (Equus asinus). PhD dissertation, University of Wisconsin, Madison, 251 pp.; Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Nat. Geogr. Soc. Res. Reports 1970, 405-411.; Woodward, S.L., 1979. The social system of feral asses (Equus asinus). Z. Tierpsychol. 49, 304-316] and in the mesic habitat of Ossabaw Island, Georgia [Moehlman, P.D., 1979, ibid; McCort, W.D., 1980. The feral asses (Equus asinus) of Ossabaw Island, Georgia. PhD dissertation, Pennsylvania State University, University Park, 219 pp]. The feral ass populations in these two locales exhibited intraspecific variation in polygynous mating systems and social organization which were consistent with the ecological classification of mating systems of Emlen and Oring (1977) [Emlen, S.T., Oring, S.W., 1977. Ecology, sexual selection, and the evolution of mating systems. Science 197 (4300), 215-223]. Feral asses in the arid environment have a `resource defense' polygynous mating system, and those in the mesic habitat exhibit `female (harem) defense' polygyny. The intraspecific variation observed in feral asses encompasses the interspecific variation observed in the family Equidae. |
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Equine Behaviour @ team @ |
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2382 |
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Moehlman, P.D.; Fowler, L.E.; Roe, J.H. |
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Feral asses (Equus africanus) of Volcano Alcedo, Galapagos: behavioral ecology, spatial distribution, and social organization |
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Journal Article |
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1998 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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60 |
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2-3 |
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197-210 |
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Equids; Feral asses; Social organization; Mating systems; Intraspecific variation; Galapagos |
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Feral asses were studied on Volcano Alcedo, Galapagos Islands, Ecuador, during the wet season of 1980. On the volcano rim during March/April, two stable groups were observed to have a `female (harem) defense' polygynous mating system [Emlen, S.T., Oring, S.W., 1977. Ecology, sexual selection, and the evolution of mating systems. Science 197 (4300), pp. 215-223] and social behavior patterns and feeding ecology similar to feral asses living in a habitat where forage and climate are similar, e.g., Ossabaw Island, Georgia [Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Nat. Geogr. Soc. Res. Rep., 1970, pp. 405-411; Moehlman, P.D., 1997. Feral asses (Equus africanus): intraspecific variation in social organization in arid and mesic habitats. J. Appl. Anim. Behav. Sci., this issue; McCort, W.D., 1980. The feral asses (Equus asinus) of Ossabaw Island, Georgia., PhD Dissertation, Pennsylvania State University, University Park, 219 pp.]. |
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Equine Behaviour @ team @ |
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2383 |
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