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Kabuga, J.D.; Gari-Kwaku, J.; Annor, S.Y. |
| Title |
Social status and its relationships to maintenance behaviour in a herd of N'dama and West African Shorthorn cattle |
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Journal Article |
| Year |
1991 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
31 |
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3-4 |
Pages |
169-181 |
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Social-related behaviour of N'dama and West African Shorthorn (WASH) cattle grazing together was studied over 10 consecutive days. Dominance was not related to age, liveweight or leadership when the animals were led into a weighing scale or into and out of the experimental paddock. Dominance had no influence on the use of shade, on drinking frequency or on grazing time, it was, however, positively associated with time spent ruminating and idling and with the frequency of allogrooming. Forced leadership into a weighing scale was negatively correlated (Spearmans rank correlation (rs=-0.69, P<0.05) with liveweight while voluntary leadership, out of the experimental paddock (rs=0.85, P<0.01) and into the experimental paddock (rs=0.76, P<0.05), was positively correlated with liveweight. Voluntary leadership also positively and significantly (P<0.01) influenced the frequency of visits to the water trough. All measures of leadership were significantly but negatively correlated with frequency of social association (close contact) between cows. N'dama were more aggressive than WASH and had higher dominance values. There was a slight tendency for WASH to associate more with their peers than N'dama with their peers. Social behaviours such as allogrooming were low and rubbing and sniffing absent in both breeds. |
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2034 |
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Kaseda, Y.; Khalil, A.M. |
| Title |
Harem size and reproductive success of stallions in Misaki feral horses |
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Journal Article |
| Year |
1996 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
47 |
Issue |
3-4 |
Pages |
163-173 |
| Keywords |
Harem size; Horse social organization; Reproduction; Blood types; Misaki feral horse |
| Abstract |
Over a 16-year period (1979-1994), long-term investigations were carried out on 14 Misaki feral stallions to analyze changes in harem size and the reproductive success. Harem size changed with the age of the stallions. Most stallions formed harem groups with four to five mares at the age of 4-6 and then the number of mares increased rapidly to the maximum at the age of 6-9 years. Thereafter, harem size decreased gradually to a minimum with advancing age. The harem size of 60 stable harem groups ranged from 1 to 9, and the average varied from a minimum mean of 1.8 in 1988 to a maximum mean of 5.3 in 1982. Mean harem size increased as adult sex ratio increased and a significant and positive correlation was found between them. One hundred and ninety-eight sire-foal pairs were determined by a paternity test with blood types and consort relations between stallions and mares during the study period. Out of 99 foals which were born in the stable harem groups, the true sires of 84 foals (85%) were the harem stallions in which the foals were born but the remaining 15 foals (15%) were sired by other harem stallions. Two out of three stallions which were studied throughout their lifetime produced 24 and 25 foals in 10 and 11 years of their reproductive lifespan, respectively. Another one produced only five foals in 6 years. The number of foals sired by the harem stallions was less than two over harem size 7 and some of the foals born in the harem were sired by other harem stallions. These results suggest that if a particular stallion monopolizes too many mares, he could not sire so many offspring because he could not always prevent his rival stallions from mating with his mares in wild or feral circumstances. |
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Equine Behaviour @ team @ |
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2308 |
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Kay, R.; Hall, C. |
| Title |
The use of a mirror reduces isolation stress in horses being transported by trailer |
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Journal Article |
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2009 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
116 |
Issue |
2-4 |
Pages |
237-243 |
| Keywords |
Horse; Isolation; Transport; Trailer; Mirror; Stress |
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Horse trailers are a common form of transportation for horses and ponies and often require the animal to travel alone or with a single companion. The current study investigated the effect of transporting horses alone, in company or with an acrylic safety mirror (measuring 81cm61.5cm) that provided surrogate companionship. The behavioural and physiological responses of 12 mature horses during a 30-min journey by trailer under the three treatments were compared. Behaviours (vocalisation, eating, head-tossing, pawing, and head-turning) were recorded. In order to assess circulatory changes that occur as part of the response to transport, heart rate (HR), rectal (Tr) and ear-pinna (Tp) temperatures were recorded. When travelling with a live companion significantly less time was spent vocalising (p<0.001), head-turning (p<0.001), head-tossing (p<0.01) and pawing (p<0.01); eating behaviour increased (p<0.05). Physiological responses (increases in HR and Tr and decreases in Tp) were also significantly reduced when travelling with a live companion (p<0.01). Travelling with the mirror did not significantly affect physiological responses compared with travelling alone, but the rise in Tr and fall in Tp was reduced (p=0.052 and p=0.051, respectively) and can be considered a trend. When travelling with a mirror significantly less time was spent turning the head (p<0.01), vocalising (p<0.05) and head-tossing (p<0.05); eating behaviour increased (p<0.05). The only significant difference between travelling with a live companion and a mirror was that the time spent turning the head round was less with a live companion (p<0.05). The provision of surrogate companionship in the form of a mirror was found to be preferable to travel alone, but where possible a live companion is recommended. Isolation during transportation was found to suppress feeding behaviour. Although peripheral blood flow (Tp) has been used to assess transport stress in other species it has not previously been used in the horse. Further evaluation of this non-invasive measure is now required. |
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0168-1591 |
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Equine Behaviour @ team @ |
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5097 |
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Keeling, L.J.; Bøe, K.E.; Christensen, J.W.; Hyyppä, S.; Jansson, H.; Jørgensen, G.H.M.; Ladewig, J.; Mejdell, C.M.; Särkijärvi, S.; Søndergaard, E.; Hartmann, E. |
| Title |
Injury incidence, reactivity and ease of handling of horses kept in groups: a matched case control study in four Nordic countries |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl Anim Behav Sci |
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horse; behaviour; reactivity; injury; welfare; management |
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Abstract There is increasing interest in keeping horses in groups, but progress is hampered by a lack of knowledge about which horses can and should be kept together. Therefore, our objective was to investigate the effect of group composition on the occurrence of injuries among horses, the ease of removing horses from groups and horses’ reactivity to a fearful stimulus. Using a matched case control design, 61 groups of horses were studied in Denmark, Norway, Finland and Sweden. They were allocated into groups of similar or different age and sex or where membership changed regularly or remained stable. Injuries were recorded before mixing the horses into treatment groups, the day after mixing and four weeks later. Reactivity of horses to a moving novel object and the behaviour of a horse being removed from its group and the reactions of other group members towards this horse and the handler were evaluated. It was hypothesized that a more socially variable group composition has beneficial effects on behaviour, ease of handling and reducing reactivity whereas frequent changes in group composition has negative consequences, resulting in more injuries. We found that differences in treatment effects were mainly related to breed, rather than group composition. Icelandic horses reacted less to the movement of the novel object (P = 0.007) and approached it more afterwards (P = 0.04). They also had fewer new injuries than warmbloods following mixing (P < 0.001) and fewer than all other groups 4 weeks later (P < 0.01). Most new injuries after mixing were minor and recorded on the horse’s head, chest, hind legs and rump. In conclusion, variations in sex and age composition of the group had little effect on injury level, reactivity and ease of handling compared to the general effect of breed. Concerns about the risk of severe injuries associated with keeping horses in groups are probably overestimated. Thus, we propose that horses can be successfully kept in groups of different sex and age composition. |
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Equine Behaviour @ team @ |
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6020 |
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Keiper, R.; Receveur, H. |
| Title |
Social interactions of free-ranging Przewalski horses in semi-reserves in the Netherlands |
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Journal Article |
| Year |
1992 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
33 |
Issue |
4 |
Pages |
303-318 |
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Social interactions were recorded in two bands of free-ranging Przewalski horses living on large (greater than 30 ha) pastures in the Netherlands. The average number of aggressive interactions per hour was 8.86 at Lelystad and 10.36 at Noorderheide. The most common aggressive interactions were lower intensity, lower cost displacements (17.2% of all aggressive acts at Lelystad, 13.2% at Noorderheide), threats to bite (42.3% and 40.7%, respectively) and threats to kick (15.4% and 23.9%, respectively). Analysis of aggression revealed that a clear, linear dominance hierarchy was present in each band. For each band there was a positive and highly significant correlation between the age of a horse and its rank in the hierarchy. In each band, the stallion was not the highest ranked horse. Non-agonistic behaviors exceeded the number of agonistic interactions (1253 vs. 558 for Lelystad; 1257 vs. 995 at Noorderheide). There was a negative correlation between the rank of a horse in the dominance hierarchy and the number of non-agonistic behaviors displayed. The group displaying the highest number of non-agonistic interactions were foals (48.9% of total non-agonistic behaviors at Lelystad; 51.1% at Noorderheide). The non-agonistic/agonistic ratio was greater than 1 for yearlings and the band stallion, as was also the case for foals, but was less than 1 for males. |
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refbase @ user @ |
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778 |
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Keiper, R.R. |
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Social interactions of the Przewalski horse (Equus przewalskii Poliakov, 1881) herd at the Munich Zoo |
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Journal Article |
| Year |
1988 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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21 |
Issue |
1-2 |
Pages |
89-97 |
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Data were collected on 972 aggressions and 233 acts of mutual grooming in a herd of 9 Przewalski horses in the Hellabrunn Tierpark in Munich, West Germany. The herd was composed of 1 adult stallion, 5 adult mares and 3 foals. A distinct linear dominance hierarchy was present in the herd, with the stallion being the top-ranking animal. Age was significantly correlated with rank. Almost 40% of all aggressions consisted of herding actions by the stallion. Threats to bite (20% of all aggressions) and threats to kick (11.4%) were next in frequency of occurrence. Most mutual grooms (71%) involved grooming the front part of the body. Although mutual grooming may be used to appease higher-ranked animals, most grooming bouts were between related horses. Foals initiate 47.6% of all allogrooming. Mutual grooming may reduce weaning conflict between a mare and her foal or may result in female coalitions that defend against predators or aggression by the herd stallion. |
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775 |
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Keiper, R.R.; Sambraus, H.H. |
| Title |
The stability of equine dominance hierarchies and the effects of kinship, proximity and foaling status on hierarchy rank |
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Journal Article |
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1986 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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16 |
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2 |
Pages |
121-130 |
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Dominance hierarchies were determined in four bands of feral horses living on Assateague Island. The bands varied in size from 10 to 16 horses, and consisted of one stallion, several mares and their offspring. The animals ranged in age from less than 1 to over 18 years. Field observation of all social interactions during the summer of 1981 was used to determine dominance. 1981 hierarchies for three of the bands were compared with hierarchies determined for the same bands in 1978, and showed that hierarchies change over time. Age was significantly correlated with rank. Mares with foals did not rank any higher in the hierarchies than mares without foals. Kinship did not appear to have an effect on dominance rank either, since neither juvenile nor adult offspring ranks correlated with the ranks of their mothers. The band stallion was not the highest-ranking animal of any band, but the location of the stallion peripheral to the main body of the band, the nature of his interactions with band members, and his length of residence in the band may have contributed to his low rank. |
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683 |
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Kendrick, K.M. |
| Title |
Intelligent perception |
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Journal Article |
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1998 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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57 |
Issue |
3-4 |
Pages |
213-231 |
| Keywords |
Intelligent perception; Environmental changes; Primates |
| Abstract |
For an animal from any species to exhibit intelligent perception it must be capable of being consciously aware of what it perceives and capable of learning from this experience. Although many organisms, and for that matter machines, are capable of rapid adaptive learning in response to perception of environmental changes, such adaptations can occur without them being consciously aware either of external stimuli or their response to them. While behavioural and neurophysiological evidence suggests that, apart from ourselves, other higher primates must also be capable of such awareness, an important central question is whether such awareness is a characteristic of primate evolution or if it also occurs in sub-primate mammals as well. In this review I will examine our behavioural and neurophysiological evidence from visual and olfactory recognition studies in the sheep to support the argument that they are likely to be aware of and learn about both social and non-social objects and that they are therefore capable of intelligent perception. However, the impact of motivational changes on these perceptual processes suggests that they may be limited in terms of both prospection and retrospection and dealing with symbolic associations. |
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796 |
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Khalil, A.M.; Kaseda, Y. |
| Title |
Early experience affects developmental behaviour and timing of harem formation in Misaki horses |
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Journal Article |
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1998 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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59 |
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4 |
Pages |
253-263 |
| Keywords |
Feral horse; Young male; Social behavior; Developmental stage |
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A study was made of the behavior of young male Misaki feral horses in the developmental stage, by observing nine of them once a week from January 1988 to December 1996. The relationship between behavior before separation and in the developmental stage was also investigated. This stage begins just after young males separate from their natal band or mothers, and it continues until they start to form harems. The duration of the developmental stage in the study ranged from 0.6 to 3.9 years, depending on the age of the young males at the time of separation. Young males associated with three types of social groups at the beginning of the developmental stage, according to their social groups before separation. These were bachelor groups, harem groups and wandering female groups. During this period, males joined the three groups, mixed sex groups and sometimes were solitary. It was considered that these associations provided a good opportunity for males to acquire different behavioral patterns and experiences before they entered the next stage. Depending on the groups with which they associated, young males that spent more time with bachelor groups had the longest average developmental stage. They associated with harem groups more often during the breeding season and more frequently with other groups or were solitary during the non-breeding season. This may be a transition period because by the end of this stage all males had spent time in solitude before forming their own harem bands. |
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2021 |
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Khalil, A.M.; Kaseda, Y. |
| Title |
Behavioral patterns and proximate reason of young male separation in Misaki feral horses |
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Journal Article |
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1997 |
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Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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54 |
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4 |
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281-289 |
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Misaki feral horse; Horse maternal-filial bond; Social behavior; Separation season; Separation reason |
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The present investigation was undertaken to study the proximate reasons why and the behavioral patterns of young male Misaki feral horses when they left their natal band or mothers. We observed a total of ten young males twice a month from January 1988 to December 1995. Almost all young males left their natal band or mothers at between 1 and 4 years of age. We found that, during the separation process, all the young males from first parity dams returned several times after the initial separation, indicating a strong attachment between primiparous mares and their male offspring. The other five separated only once without rejoining. Our observations showed five variable behavior patterns of young males at separation time, depending on the consort relation between their mothers and harem stallion and the reason for separation at that time. Eight young males separated in the non-breeding season at average 2.1 years and the other two separated in the breeding season at average 3 years and the average difference was not significant. These results revealed that 80% of the young males separated voluntarily when the natural resources become poor whereas 20% separated when their siblings were born. |
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Equine Behaviour @ team @ |
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