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	Author	Sigurjónsdóttir, H.; Haraldsson, H.
	Title	Significance of Group Composition for the Welfare of Pastured Horses			Type	Journal Article
	Year	2019	Publication	Animals	Abbreviated Journal	Animals
	Volume	9	Issue	14	Pages
	Keywords	horse welfare; aggression; allogrooming; pastured horses; Icelandic horse
	Abstract	We explore how herd composition and management factors correlate with frequencies of social interactions in horse groups. Since the welfare of horses correlates with low aggression levels and social contact opportunities, information of this kind is important. The data are a collection of records of social interactions of 426 Icelandic horses in 20 groups of at least eight horses. The complexities and limitations of the data prohibit useful statistical modelling so the results are presented descriptively. Interesting and informative patterns emerge which can be of use both in management and in future studies. Of special interest are the low levels of agonistic behaviours in breeding groups where one stallion was present. The horses were less agonistic when in groups with young foals and where group membership was stable. Unfamiliar yearlings in peer groups were especially aggressive. Allogrooming was most frequent in groups with relatively more young horses and in unstable and small groups. Interestingly, the horses allogroomed more if they had few preferred allogrooming partners. The findings show that composition (age/sex) and stability of groups are of great importance with respect to aggression levels and opportunities for establishing bonds.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title	Animals
	Series Volume	9	Series Issue	1	Edition
	ISSN	2076-2615	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6510
Permanent link to this record



	Author	Sigurjónsdóttir, H.; Haraldsson, H.
	Title	Significance of Group Composition for the Welfare of Pastured Horses			Type	Journal Article
	Year	2018	Publication	Animals	Abbreviated Journal	Animals
	Volume	9	Issue	1	Pages	14
	Keywords	horse welfare; aggression; allogrooming; pastured horses; Icelandic horse
	Abstract	Simple Summary Because of their social nature, horses need to have plenty of opportunities to interact with others to establish bonds and learn from their elders. Comparison of social behaviour of 426 horses in 20 groups of Icelandic horses in pastures, showed that aggression was lowest where the group composition was like the natural system, i.e., with a stallion, mares and their young foals. In groups without a stallion, the presence of foals is also associated with low aggression. Stability of the group with respect to group composition is of great importance; the horses are less aggressive in the more stable groups. The highest aggression was found in groups of unfamiliar yearlings. The horses allogroomed more in groups with relatively more young horses, which suggests they are forming bonds. Later, they groom less but prefer certain individuals. Horse owners should all be aware of the importance of planning the composition of horse groups and to keep the membership as stable as possible in order to ensure good welfare. Abstract We explore how herd composition and management factors correlate with frequencies of social interactions in horse groups. Since the welfare of horses correlates with low aggression levels and social contact opportunities, information of this kind is important. The data are a collection of records of social interactions of 426 Icelandic horses in 20 groups of at least eight horses. The complexities and limitations of the data prohibit useful statistical modelling so the results are presented descriptively. Interesting and informative patterns emerge which can be of use both in management and in future studies. Of special interest are the low levels of agonistic behaviours in breeding groups where one stallion was present. The horses were less agonistic when in groups with young foals and where group membership was stable. Unfamiliar yearlings in peer groups were especially aggressive. Allogrooming was most frequent in groups with relatively more young horses and in unstable and small groups. Interestingly, the horses allogroomed more if they had few preferred allogrooming partners. The findings show that composition (age/sex) and stability of groups are of great importance with respect to aggression levels and opportunities for establishing bonds.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6585
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	Author	Silk, J. B.
	Title	Patterns of intervention in agonistic contests among male bonnet macaques			Type	Book Chapter
	Year	1992	Publication	Coalitions and Alliances in Humans and Other Animals	Abbreviated Journal
	Volume		Issue		Pages	215-232
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Oxford University Press	Place of Publication	Oxford	Editor	Harcourt, A.H., and de Waal, F.B.M.
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5234
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	Author	Tomasello, M.
	Title	Do apes ape?			Type	Book Chapter
	Year	1996	Publication	Social learning in animals: the roots of culture	Abbreviated Journal
	Volume		Issue		Pages	319-346
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Academic Press	Place of Publication	London	Editor	Heyes, C. M.; Galef, B.G.
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5600
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	Author	Tomasello, M.; Call, J.
	Title	Do chimpanzees know what others see ? or only what they are looking at?			Type	Book Chapter
	Year	2006	Publication	Rational Animals?	Abbreviated Journal
	Volume		Issue		Pages	371-384
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Oxford University Press	Place of Publication	Oxford	Editor	Nudds, M.; Hurley, S.
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4094
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	Author	Villani, M.; Cairoli, F.; Kindahl, H.; Galeati, G.; Faustini, M.; Carluccio, A.; Veronesi, M.C.
	Title	Effects of mating on plasma concentrations of testosterone, cortisol, oestrone sulphate and 15-ketodihydro-PGF2alpha in stallions			Type	Journal Article
	Year	2006	Publication	Reproduction in Domestic Animals = Zuchthygiene	Abbreviated Journal	Reprod Domest Anim
	Volume	41	Issue	6	Pages	544-548
	Keywords	Animals; Dinoprost/analogs & derivatives/blood; Ejaculation/physiology; Estrone/analogs & derivatives/blood; Horses/blood/physiology; Hydrocortisone/blood; Male; Sexual Behavior, Animal/physiology; Testosterone/blood
	Abstract	Very little information is available regarding the physiological mechanisms involved in the normal sexual activity in the stallion and, in particular, the endocrine control of reproduction is still not clearly understood. This experiment was designed to determine the short-term effect of sexual stimulation on plasma concentrations of testosterone, cortisol, oestrone sulphate and 15-ketodihydro-PGF(2alpha) in stallions. Semen samples were collected from 10 lighthorse stallions of proven fertility using a Missouri model artificial vagina. At the same time, blood samples were collected from the jugular vein with heparinized tubes, 20 and 10 min before oestrous mare exposure, at exposure and 10, 20, 30 min after dismounting. Testosterone concentrations showed a sharp rise 10 min after mating (p < 0.001), reached a plateau, and then showed a further increase 30 min after mating (p < 0.001). Cortisol concentrations increased 10 min after mating (p < 0.001) and remained at high levels in the subsequent samples taken. A peak of oestrone sulphate was observed 10 min after mating (p < 0.001). 15-Ketodihydro-PGF(2alpha) concentrations decreased rapidly at the moment of the exposure of the stallions to an oestrous mare (p < 0.05), returned to pre-mating concentrations and then decreased again 30 min after mating (p < 0.05).
	Address	Dipartimento di Scienze Cliniche Veterinarie, Sez. Clinica Ostetrica e Ginecologica Veterinaria, Milan, Italy
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0936-6768	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17107515			Approved	no
	Call Number				Serial	1855
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	Author	Virányi, Zs.; Range, F.; Huber, L.
	Title	Attentiveness toward others and social learning in domestic dogs.			Type	Book Chapter
	Year	2008	Publication	Learning from Animals?: Examining the Nature of Human Uniqueness	Abbreviated Journal
	Volume		Issue		Pages	141-154
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Psychology Press	Place of Publication	New York, NY	Editor	Röska-hardy,L.S.. ;Neumann-held, E.
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN	978-1-84169-707-9	Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4974
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	Author	Waiblinger, S
	Title	Animal welfare and housing			Type	Book Chapter
	Year	2009	Publication	Welfare of Production Animals:: Assessment and Management of Risks (Food Safety Assurance and Veterinary Public Health)	Abbreviated Journal
	Volume		Issue		Pages	79-111
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Wageningen Acad. Publ.	Place of Publication	Wageningen	Editor	Smulders, F. J.
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN	978-90-8686-122-4.	Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5302
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	Author	Wingfield, J. C.,; Ramenofsky, M.
	Title	Hormones and the behavioral ecology of stress.			Type	Book Chapter
	Year	1999	Publication	Stress physiology in animals.	Abbreviated Journal
	Volume		Issue		Pages	1-51
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Sheffield Academic Press	Place of Publication	Sheffield, United Kingdom	Editor	Balm, P. H. M.
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4071
Permanent link to this record



	Author	Wolter, R.; Stefanski, V.; Krueger, K.
	Title	Parameters for the Analysis of Social Bonds in Horses			Type	Journal Article
	Year	2018	Publication	Animals	Abbreviated Journal	Animals
	Volume	8	Issue	11	Pages	191
	Keywords	feral horses; mutual grooming; social bonds; social bond analysis; spatial proximity
	Abstract	Social bond analysis is of major importance for the evaluation of social relationships in group housed horses. However, in equine behaviour literature, studies on social bond analysis are inconsistent. Mutual grooming (horses standing side by side and gently nipping, nuzzling, or rubbing each other), affiliative approaches (horses approaching each other and staying within one body length), and measurements of spatial proximity (horses standing with body contact or within two horse-lengths) are commonly used. In the present study, we assessed which of the three parameters is most suitable for social bond analysis in horses, and whether social bonds are affected by individual and group factors. We observed social behaviour and spatial proximity in 145 feral horses, five groups of Przewalskiâ��s horses (N = 36), and six groups of feral horses (N = 109) for 15 h per group, on three days within one week. We found grooming, friendly approaches, and spatial proximity to be robust parameters, as their correlation was affected only by the animalsâ�� sex (GLMM: N = 145, SE = 0.001, t = â��2.7, p = 0.008) and the group size (GLMM: N = 145, SE < 0.001, t = 4.255, p < 0.001), but not by the horse breed, the aggression ratio, the social rank, the group, the group composition, and the individuals themselves. Our results show a trend for a correspondence between all three parameters (GLMM: N = 145, SE = 0.004, t = 1.95, p = 0.053), a strong correspondence between mutual grooming and friendly approaches (GLMM: N = 145, SE = 0.021, t = 3.922, p < 0.001), and a weak correspondence between mutual grooming and spatial proximity (GLMM: N = 145, SE = 0.04, t = 1.15, p = 0.25). We therefore suggest either using a combination of the proactive behaviour counts mutual grooming and friendly approaches, or using measurements of close spatial proximity, for the analysis of social bonds in horses within a limited time frame.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	2076-2615	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6428
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