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Author Trösch, M.; Pellon, S.; Cuzol, F.; Parias, C.; Nowak, R.; Calandreau, L.; Lansade, L. url  doi
openurl 
  Title Horses feel emotions when they watch positive and negative horse-human interactions in a video and transpose what they saw to real life Type (up) Journal Article
  Year 2020 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume 23 Issue 4 Pages 643-653  
  Keywords  
  Abstract Animals can indirectly gather meaningful information about other individuals by eavesdropping on their third-party interactions. In particular, eavesdropping can be used to indirectly attribute a negative or positive valence to an individual and to adjust one's future behavior towards that individual. Few studies have focused on this ability in nonhuman animals, especially in nonprimate species. Here, we investigated this ability for the first time in domestic horses (Equus caballus) by projecting videos of positive and negative interactions between an unknown human experimenter (a “positive” experimenter or a “negative” experimenter) and an actor horse. The horses reacted emotionally while watching the videos, expressing behavioral (facial expressions and contact-seeking behavior) and physiological (heart rate) cues of positive emotions while watching the positive video and of negative emotions while watching the negative video. This result shows that the horses perceived the content of the videos and suggests an emotional contagion between the actor horse and the subjects. After the videos were projected, the horses took a choice test, facing the positive and negative experimenters in real life. The horses successfully used the interactions seen in the videos to discriminate between the experimenters. They touched the negative experimenter significantly more, which seems counterintuitive but can be interpreted as an appeasement attempt, based on the existing literature. This result suggests that horses can indirectly attribute a valence to a human experimenter by eavesdropping on a previous third-party interaction with a conspecific.  
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  ISSN 1435-9456 ISBN Medium  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Trösch2020 Serial 6649  
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Author Ahrendt, L.P.; Labouriau, R.; Malmkvist, J.; Nicol, C.J.; Christensen, J.W. url  doi
openurl 
  Title Development of a standard test to assess negative reinforcement learning in horses Type (up) Journal Article
  Year 2015 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 169 Issue Pages 38-42  
  Keywords Algometry; Horse behaviour; Learning performance; Operant conditioning; Pressure-release; Horse training  
  Abstract Most horses are trained by negative reinforcement. Currently, however, no standardised test for evaluating horses' negative reinforcement learning ability is available. The aim of this study was to develop an objective test to investigate negative reinforcement learning in horses. Twenty-four Icelandic horses (3 years old) were included in this study. The horses were tested in a pressure-release task on three separate days with 10, 7 and 5 trials on each side, respectively. Each trial consisted of pressure being applied on the hindquarter with an algometer. The force of the pressure was increased until the horse moved laterally away from the point of pressure. There was a significant decrease in required force over trials on the first test day (P<0.001), but not the second and third day. The intercepts on days 2 and 3 differed significantly from day 1 (P<0.001), but not each other. Significantly stronger force was required on the right side compared to the left (P<0.001), but there was no difference between first and second side tested (P=0.56). Individual performance was evaluated by median-force and the change in force over trials on the first test day. These two measures may explain different characteristics of negative reinforcement learning. In conclusion, this study presents a novel, standardised test for evaluating negative reinforcement learning ability in horses.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0168-1591 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6650  
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Author Mejdell, C.M.; Buvik, T.; Jørgensen, G.H.M.; Bøe, K.E. url  doi
openurl 
  Title Horses can learn to use symbols to communicate their preferences Type (up) Journal Article
  Year 2016 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 184 Issue Pages 66-73  
  Keywords Operant conditioning; Blanket; Rug; Thermoregulation; Cognition; Clicker training  
  Abstract This paper describes a method in which horses learn to communicate by touching different neutral visual symbols, in order to tell the handler whether they want to have a blanket on or not. Horses were trained for 10-15min per day, following a training program comprising ten steps in a strategic order. Reward based operant conditioning was used to teach horses to approach and touch a board, and to understand the meaning of three different symbols. Heat and cold challenges were performed to help learning and to check level of understanding. At certain stages, a learning criterion of correct responses for 8-14 successive trials had to be achieved before proceeding. After introducing the free choice situation, on average at training day 11, the horse could choose between a “no change” symbol and the symbol for either “blanket on” or “blanket off” depending on whether the horse already wore a blanket or not. A cut off point for performance or non-performance was set to day 14, and 23/23 horses successfully learned the task within this limit. Horses of warm-blood type needed fewer training days to reach criterion than cold-bloods (P<0.05). Horses were then tested under differing weather conditions. Results show that choices made, i.e. the symbol touched, was not random but dependent on weather. Horses chose to stay without a blanket in nice weather, and they chose to have a blanket on when the weather was wet, windy and cold (χ2=36.67, P<0.005). This indicates that horses both had an understanding of the consequence of their choice on own thermal comfort, and that they successfully had learned to communicate their preference by using the symbols. The method represents a novel tool for studying preferences in horses.  
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  ISSN 0168-1591 ISBN Medium  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6651  
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Author Krueger, K.; Esch, L.; Byrne, R. url  doi
openurl 
  Title Need or opportunity? A study of innovations in equids Type (up) Journal Article
  Year 2021 Publication Plos One Abbreviated Journal Plos One  
  Volume 16 Issue 9 Pages e0257730  
  Keywords  
  Abstract Debate persists over whether animals develop innovative solutions primarily in response to needs or conversely whether they innovate more when basic needs are covered and opportunity to develop novel behaviour is offered. We sourced 746 cases of “unusual” behaviour in equids by contacting equid owners and caretakers directly and via a website (https://innovative-behaviour.org), and by searching the internet platforms YouTube and Facebook for videos. The study investigated whether differences in need or opportunity for innovation were reflected in the numbers of different types of innovations and in the frequencies of repeating a once-innovative behaviour (i) with respect to the equids' sex, age, and breed type, (ii) across behavioural categories, and whether (iii) they were affected by the equids' management (single vs group housing, access to roughage feed, access to pasture, and social contact). We found that the numbers of different types of innovation and the frequency of displaying specific innovations were not affected by individual characteristics (sex, age, breed or equid species). Few types of innovation in escape and foraging contexts were observed, whilst the comfort, play, and social contexts elicited the greatest variety of innovations. We also found higher numbers of different types of innovations in horses kept in groups rather than in individual housing, and with unlimited rather than with restricted access to pasture and roughage. Equids in permanent social contact performed high rates of once-innovative behaviour. We suggest that equids produce goal-directed innovations and repeat the behaviour at high frequency in response to urgent needs for food and free movement or when kept in conditions with social conflict. However, equids devise the greatest variety of innovations when opportunity to play and to develop comfort behaviour arises and when kept in good conditions.  
  Address  
  Corporate Author Thesis  
  Publisher Public Library of Science Place of Publication Editor  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6653  
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Author Piro, M.; Benjouad, A.; Karom, A.; Nabich, A.; Benbihi, N.; El Allali, K.; Machmoum, M.; Ouragh, L. url  doi
openurl 
  Title Genetic Structure of Severe Combined Immunodeficiency Carrier Horses in Morocco Inferred by Microsatellite Data Type (up) Journal Article
  Year 2011 Publication Journal of Equine Veterinary Science Abbreviated Journal J. Equine Vet. Sci.  
  Volume 31 Issue 11 Pages 618-624  
  Keywords Scid; Arab horses; Arab-Barb horses; Microsatellite; Dna; Genetic structure  
  Abstract A total of 17 microsatellite deoxyribonucleic acid loci used routinely for horse parentage control were used to evaluate genetic diversity among normal Arabian horses and severe combined immunodeficiency (SCID) carrier Arabian horses (ArS) and normal Arab-Barb horses and SCID carrier Arab-Barb horses (ArbeS). On the basis of the genotype of 186 horses, mean allelic diversity was estimated as 6.82, 5.53, and 6.7059 in normal Arabian horses, ArS, and for both groups of Arab-Barb horses, respectively. Five specific alleles were observed in ArS and ArbeS, with one common with ArS at HMS6, whereas five alleles common between ArS and ArbeS had a high frequency. Expected and observed heterozygosity showed great heterogeneity in the population studied and were similar or higher when compared with other studies on Arabian horses. Coefficient of gene differentiation Gst of Nei associated with Nei's genetic distance and multivariate correspondence analysis indicated a possible differentiation between the studied populations when analyzed separately according to breed. Probability of assignment of a horse to a specific group was assessed using a full and partial Bayesian approach. In all, 80.6% of Arab horses and 78.2% of Arab-Barb horses were assigned properly with a partial Bayesian test, which provided better results than the full one. These findings will be useful for identification of SCID carrier horses by using the microsatellite deoxyribonucleic acid loci used routinely for horse parentage control in our laboratory.  
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  Series Volume Series Issue Edition  
  ISSN 0737-0806 ISBN Medium  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6657  
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Author Bandini, E.; Tennie C. doi  openurl
  Title Exploring the role of individual learning in animal tool-use Type (up) Journal Article
  Year 2020 Publication PeerJ Abbreviated Journal PeerJ  
  Volume 25 Issue Pages 8:e9877  
  Keywords  
  Abstract The notion that tool-use is unique to humans has long been refuted by the growing number of observations of animals using tools across various contexts. Yet, the mechanisms behind the emergence and sustenance of these tool-use repertoires are still heavily debated. We argue that the current animal behaviour literature is biased towards a social learning approach, in which animal, and in particular primate, tool-use repertoires are thought to require social learning mechanisms (copying variants of social learning are most often invoked). However, concrete evidence for a widespread dependency on social learning is still lacking. On the other hand, a growing body of observational and experimental data demonstrates that various animal species are capable of acquiring the forms of their tool-use behaviours via individual learning, with (non-copying) social learning regulating the frequencies of the behavioural forms within (and, indirectly, between) groups. As a first outline of the extent of the role of individual learning in animal tool-use, a literature review of reports of the spontaneous acquisition of animal tool-use behaviours was carried out across observational and experimental studies. The results of this review suggest that perhaps due to the pervasive focus on social learning in the literature, accounts of the individual learning of tool-use forms by naïve animals may have been largely overlooked, and their importance under-examined.  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6659  
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Author Bandini , E.; Motes-Rodrigo, A.; Steele, M.P.; Rutz, C.; Tennie, C. doi  openurl
  Title Examining the mechanisms underlying the acquisition of animal tool behaviour Type (up) Journal Article
  Year 2020 Publication Biology Letters Abbreviated Journal Biol. Lett.  
  Volume 16 Issue 2020122 Pages  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6660  
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Author Solmsen, E.-H.; Bathen, M.; Grüntjens, T.; Hempel, E.; Klose, M.; Krüger, K.; Martin, H.; Meyer, A.; Schütte, P.; Vogel, L.; Wiezorek, S.; Wittor, B. openurl 
  Title Protecting horses against wolves in Germany Type (up) Journal Article
  Year 2021 Publication Carnivore Damage Prevention News Abbreviated Journal CPDnews  
  Volume 23 Issue Pages 12-19  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6661  
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Author Maloney, S.J. url  openurl
  Title The Relationship Between Asymmetry and Athletic Performance: A Critical Review Type (up) Journal Article
  Year 2019 Publication The Journal of Strength & Conditioning Research Abbreviated Journal  
  Volume 33 Issue 9 Pages  
  Keywords symmetry; imbalance; power; strength  
  Abstract Maloney, SJ. The relationship between asymmetry and athletic performance: A critical review. J Strength Cond Res 33(9): 2579-2593, 2019--Symmetry may be defined as the quality to demonstrate an exact correspondence of size, shape, and form when split along a given axis. Although it has been widely asserted that the bilateral asymmetries are detrimental to athletic performance, research does not wholly support such an association. Moreover, the research rarely seeks to distinguish between different types of bilateral asymmetry. Fluctuating asymmetries describe bilateral differences in anthropometric attributes, such as nostril width and ear size, and are thought to represent the developmental stability of an organism. There is evidence to suggest that fluctuating asymmetries may be related to impaired athletic performance, although contradictory findings have been reported. Sporting asymmetries is a term that may better describe bilateral differences in parameters, such as force output or jump height. These asymmetries are likely to be a function of limb dominance and magnified by long-standing participation within sport. Sporting asymmetries do not seem to carry a clear influence on athletic performance measures. Given the vast discrepancy in the methodologies used by different investigations, further research is warranted. Recent investigations have demonstrated that training interventions can reduce sporting asymmetries and improve performance. However, studies have not sought to determine whether the influence of sporting asymmetry is independent of improvements in neuromuscular parameters. It may be hypothesized that the deficient (weaker) limb has a greater potential for adaptation in comparison to the strong limb and may demonstrate greater responsiveness to training.  
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  Series Volume Series Issue Edition  
  ISSN 1064-8011 ISBN Medium  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ 00124278-201909000-00032 Serial 6662  
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Author Byström, A.; Clayton, H.M.; Hernlund, E.; Rhodin, M.; Egenvall, A. url  doi
openurl 
  Title Equestrian and biomechanical perspectives on laterality in the horse Type (up) Journal Article
  Year 2020 Publication Comparative Exercise Physiology Abbreviated Journal Comp. Exerc. Physiol.  
  Volume 16 Issue 1 Pages 35-45  
  Keywords  
  Abstract It has been suggested that one of the underlying causes of asymmetrical performance and left/right bias in sound riding horses is laterality originating in the cerebral cortices described in many species. The aim of this paper is to review the published evidence for inherent biomechanical laterality in horses deemed to be clinically sound and relate these findings to descriptions of sidedness in equestrian texts. There are no established criteria to determine if a horse is left or right dominant but the preferred limb has been defined as the forelimb that is more frequently protracted during stance and when grazing. Findings on left-right differences in forelimb hoof shape and front hoof angles have been linked to asymmetric forelimb ground reaction forces. Asymmetries interpreted as motor laterality have been found among foals and unhandled youngsters, and the consistency or extent of asymmetries seems to increase with age. Expressions of laterality also vary with breed, sex, training and handling, stress, and body shape but there are no studies of the possible link between laterality and lameness. In a recent study of a group of seven dressage horses, a movement pattern in many ways similar to descriptions of sidedness in the equestrian literature, e.g. one hind limb being more protracted and placed more laterally than the other, has been documented. The role of innate laterality versus painful conditions, training, human handedness and simply habit remains to be determined. Understanding the biomechanical manifestations of laterality in healthy horses, including individual variation, would yield a potential basis for how laterality should be taken into account in relation to training/riding and rehabilitation of lameness.  
  Address  
  Corporate Author Thesis  
  Publisher Wageningen Academic Publishers Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1755-2540 ISBN Medium  
  Area Expedition Conference  
  Notes doi: 10.3920/CEP190022 Approved no  
  Call Number Equine Behaviour @ team @ Serial 6663  
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