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	Author	Byström, A.; Clayton, H.M.; Hernlund, E.; Rhodin, M.; Egenvall, A.
	Title	Equestrian and biomechanical perspectives on laterality in the horse			Type	Journal Article
	Year	2020	Publication	Comparative Exercise Physiology	Abbreviated Journal	Comp. Exerc. Physiol.
	Volume	16	Issue	1	Pages	35-45
	Keywords
	Abstract	It has been suggested that one of the underlying causes of asymmetrical performance and left/right bias in sound riding horses is laterality originating in the cerebral cortices described in many species. The aim of this paper is to review the published evidence for inherent biomechanical laterality in horses deemed to be clinically sound and relate these findings to descriptions of sidedness in equestrian texts. There are no established criteria to determine if a horse is left or right dominant but the preferred limb has been defined as the forelimb that is more frequently protracted during stance and when grazing. Findings on left-right differences in forelimb hoof shape and front hoof angles have been linked to asymmetric forelimb ground reaction forces. Asymmetries interpreted as motor laterality have been found among foals and unhandled youngsters, and the consistency or extent of asymmetries seems to increase with age. Expressions of laterality also vary with breed, sex, training and handling, stress, and body shape but there are no studies of the possible link between laterality and lameness. In a recent study of a group of seven dressage horses, a movement pattern in many ways similar to descriptions of sidedness in the equestrian literature, e.g. one hind limb being more protracted and placed more laterally than the other, has been documented. The role of innate laterality versus painful conditions, training, human handedness and simply habit remains to be determined. Understanding the biomechanical manifestations of laterality in healthy horses, including individual variation, would yield a potential basis for how laterality should be taken into account in relation to training/riding and rehabilitation of lameness.
	Address
	Corporate Author				Thesis
	Publisher	Wageningen Academic Publishers	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1755-2540	ISBN		Medium
	Area		Expedition		Conference
	Notes	doi: 10.3920/CEP190022			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6663
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	Author	Krueger, K.
	Title	“Pferdehaltung und Ethologie der Pferde” im Bachelorstudiengang Pferdewirtschaft			Type	Book Chapter
	Year	2014	Publication	Forschendes Lernen initiieren, umsetzen und reflektieren	Abbreviated Journal
	Volume		Issue		Pages	54-81
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	UniversitätsVerlag Webler	Place of Publication	Bielefeld	Editor	: S. Lepp und C. Niederdrenk-Felgner
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN	10: 3-937026-91-6	Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5944
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	Author	Grönemann, C.
	Title	Konfliktfeld Pferd und Wolf – Eine Untersuchung zu Einstellungen, Erwartungen und Befürchtungen von Pferdehaltern und Reitsportlern in Niedersachsen			Type	Manuscript
	Year	2015	Publication	master thesis	Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract
	Address
	Corporate Author				Thesis	Master's thesis
	Publisher	Universität Hildesheim	Place of Publication	Hildesheim	Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6683
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	Author	Wolter, R.
	Title	The behaviour and managementof Przewalski’shorsesin semi-reserves			Type	Manuscript
	Year	2018	Publication	Phd thesis	Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract	In recent years, Przewalski’s horses have been increasingly kept in semi-reserves. However,there areonly few studies ontheir behaviour and their ability to adaptto management interventions.In the main part of my dissertation, I focus on investigatingthe animals’ behaviour in different semi-reserves with varyinghabitats and living spaces. In addition, I investigate the horses’ behaviour during various management interventionsand analysetheensuing changes instress levels. Another aspect of my dissertation is the studyof social behaviour inPrzewalski’s horses. I investigate theparameters that should be used to demonstrate social bonds between individualsandassess whichdata provide the most meaningful results.In the commentary tochapter 1,several studies investigatingsocial bonds in horsesare discussed. Comparing the various studies, it is strikingthat no homogeneous analyses orevaluation criteria exist. While some authors only considersocial grooming, others include data onthe spatial proximity of the individuals in their evaluations, and various definitionsof proximity can also be found in the literature. Additionally, someauthors use friendly approaches between individuals asa furtherparameter wheninvestigating the social bonds.Continuing with this theme, in chapter 2I investigate the social behaviour of the horses and compare various analysis methods. I show that proactive behaviour, such as friendly approaches, is a good alternative to spatial proximity when investigating social bonds between group members, andis also useful for expanding the often very small data sets of mutual grooming in horses. Comparing Przewalski’s horses with wild living horses, I found no significant differences in the social behavior and the frequency of social interactions, regardlessof group size, group composition, habitat, and individual parameters such as age and gender.Inchapter 3,I investigate the behaviour of a Przewalski’s horse group when exploring a new area of their enclosure. Their behaviour changed, showing less resting and more feeding. Furthermore, the animals maintained greater distances from each other, and the alpha male, instead of herding the group from behind, led the group around the new area and walked in front of the other group members. Moreover, he showed a substantial increase in stress level during the first day.A general comparison of the behaviour of the Przewalski’s horses in different semi-reserves is provided in chapter 4. In it, the habitat choice of the animals and their reactions to various management interventions are investigated. It is shown that Przewalski’s horses prefer open grassland to dense woods, although keeping Przewalski’s horses in a pine forest does not influence the animals’ stress level. In contrast to habitat, food range, and changes in the group composition, which do not appear to change stress levels, individual factors, such as the hierarchy, influence the glucocorticoid level of the animals significantly. The largest increases in stress hormones were demonstrated when the horses were temporarily confined in smaller areas.The importance of the available space is also discussed in chapter 5, where it is shown that horses show less aggressive behaviour when more space is provided. In contrast, the husbandry system does not influence the animals’ aggression, but the way of feeding can additionally reduce agonistic behaviour, for example if food is offered ad libitum.In summary, the results of this study provide indications for the optimization of keeping Przewalski’s horses in semi-reserves. The animals can adapt themselves to the environment and thrive in habitats which do not correspond to their original steppe-like home. Nevertheless, the semi-reserves should provide sufficient grassland, as the horses prefer this type of habitat. General speaking, any types of habitat can only offer a suitable living space if the food range is sufficient for the number of horses. Otherwise, and especially during could winter months, supplementary feeding is necessary according to the body condition of the animals. This is particularly important for older, weakened, or very young animals, which are still adapting to life in the semi-reserve. Without sufficient food, stress hormones can increase and negatively influence the well-being of the horses. The same is true for management interventions: restricting the animals to small enclosures, for example, can adversely affect the horses’ well-being and should be only done if absolutely necessary. Targetedbehaviour observations allow the animals that have a special meaning for the group to be identified, and these should not be taken out of the group unless it is unavoidable, as young and unexperienced horses orientate themselves on those animals. This is especially true for the alpha male in a bachelor group, as these groups are often composed of young horses and the alpha-male provides the necessary stability and experience. Social bonds between individuals can be investigated by observing friendly and proactive behaviour, and social grooming and friendly approaches yield suitable data for such analysis.
	Address
	Corporate Author				Thesis	Ph.D. thesis
	Publisher	University Regensburg	Place of Publication	Regensburg	Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6639
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	Author	Golynski, M.; Szczepanik, M.P.; Wilkolek, P.M.; Adamek, L.R.; Sitkowski, W.; Taszkun, I.
	Title	Influence of hair clipping on transepidermal water loss values in horses: a pilot study			Type	Journal Article
	Year	2018	Publication	Polish Journal of Veterinary Sciences	Abbreviated Journal
	Volume	vol. 21	Issue	No 1	Pages
	Keywords	horses; transepidermal water loss; clipping
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	University of Warmia and Mazury in Olsztyn	Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6612
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	Author	Zeder, M.A.
	Title	Pathways to animal domestication			Type	Book Chapter
	Year	2011	Publication	Harlan II: Biodiversity in Agriculture: Domestication, Evolution, and Sustainability	Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	University of California	Place of Publication	Davis	Editor	Damania, A.; Gepts, P.
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Zeder2011			Serial	6316
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	Author	Grönemann, K.
	Title	Konfliktfeld Pferd und Wolf – Eine Untersuchung zu Einstellungen, Erwartungen und Befürchtungen von Pferdehaltern und Reitsportlern in Niedersachsen			Type	Manuscript
	Year	2015	Publication		Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract
	Address
	Corporate Author				Thesis	Master's thesis
	Publisher	University Hildesheim	Place of Publication	Hildesheim	Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6440
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	Author	Krösbacher, A. E.
	Title	Das Arabische Vollblut: Eine kontrovers diskutierte Rasse: Was steckt wirklich hinter der Zucht dieser edlen Pferde?			Type	Manuscript
	Year	2008	Publication		Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract
	Address
	Corporate Author				Thesis	Bachelor's thesis
	Publisher	University for Veterinarian Medicine Vienna	Place of Publication	Vienna	Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6544
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	Author	Lagos, L.
	Title	Ecología del lobo (Canis lupus), del poni salvaje (Equus ferus atlanticus) y del ganado vacuno semiextensivo (Bos taurus) en Galicia: interacciones depredador-presa.			Type	Manuscript
	Year	2013	Publication	Phd thesis	Abbreviated Journal
	Volume		Issue		Pages	458
	Keywords
	Abstract
	Address
	Corporate Author				Thesis	Ph.D. thesis
	Publisher	Universidad de Santiago de Compostela	Place of Publication	Santiago de Compostela	Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6678
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	Author	Pimlott, D.H.
	Title	The use of tape-recorded wolf howls to locate timber wolves			Type	Book Whole
	Year	1960	Publication		Abbreviated Journal
	Volume		Issue		Pages
	Keywords
	Abstract
	Address
	Corporate Author				Thesis
	Publisher	Twenty-second Midwest Wildlife Congress	Place of Publication	Toronto	Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @ Pimlott1960			Serial	6454
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