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	Author	Custance, D.; Whiten, A.; Sambrook, T.; Galdikas, B.
	Title	Testing for social learning in the “artificial fruit” processing of wildborn orangutans (Pongo pygmaeus), Tanjung Puting, Indonesia			Type	Journal Article
	Year	2001	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	4	Issue	3	Pages	305-313
	Keywords
	Abstract	Social learning about actions, objects and sequencing was investigated in a group of 14 wildborn orangutans (four adult females and ten 3- to 5-year-old juveniles). Human models showed alternative methods and sequences for dismantling an artificial fruit to groups of participants matched by gender and age. Each participant received three to six 2-min trials in which they were given access to the artificial fruit for manipulation. Independent coders, who were unaware of which method each participant had seen, gave confidence ratings and collected action frequencies from watching video recordings of the experimental trials. No significant differences were found between groups in terms of the coders' confidence ratings, the action frequencies or the sequence of manipulations. These negative results may at least partly reflect the immaturity of a large proportion of the participants. A positive correlation was found between age and the degree of matching to the method shown. Although none of the juveniles succeeded in opening the “fruit”, two out of the four adults did so and they also seemed to match more closely the sequence of elements touched over successive trials. The results are compared with similar data previously collected from human children, chimpanzees, gorillas, capuchin monkeys and common marmosets.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3370
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	Author	Reader, S.M.
	Title	Innovation and social learning: individual variation and brain evolution			Type	Journal Article
	Year	2003	Publication	Animal Biology (formerly Netherlands Journal of Zoology)	Abbreviated Journal	Anim. Biol. Leiden.
	Volume	53	Issue	2	Pages	147-158
	Keywords
	Abstract	This paper reviews behavioural, neurological and cognitive correlates of innovation at the individual, population and species level, focusing on birds and primates. Innovation, new or modified learned behaviour not previously found in the population, is the first stage in many instances of cultural transmission and may play an important role in the lives of animals with generalist or opportunistic lifestyles. Within-species, innovation is associated with low neophobia, high neophilia, and with high social learning propensities. Indices of innovatory propensities can be calculated for taxonomic groups by counting the frequency of reports of innovation in published literature. These innovation rate data provide a useful comparative measure for studies of behavioural flexibility and cognition. Innovation rate is positively correlated with the relative size of association areas in the brain, namely the hyperstriatum ventrale and neostriatum in birds, and the neocortex and striatum in primates. Innovation rate is also positively correlated with the reported variety of tool use, as well as interspecific differences in learning. Current evidence thus suggests similar patterns of cognitive evolution in primates and birds.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3395
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	Author	Gardner, P.
	Title	The responses of horses in a discrimination problem			Type	Journal Article
	Year	1937	Publication	Journal-of-Comparative-Psychology	Abbreviated Journal	J Comp Psychol
	Volume	23	Issue		Pages	305-333
	Keywords
	Abstract	62 horses were trained to obtain food from the one of three boxes which was covered with a black cloth. The position of the box varied from trial to trial in a random order. Learning was apparently in terms of vision, rather than smell. Many errors were due to the line of direction of the horse's movement as it entered the experimental situation. For all animals the learning curve dropped rapidly during the first few trials. There was slightly more rapid learning in younger horses than in older ones. No sex differences were apparent. Percherons made fewer errors than Belgians. Draft horses showed a slight superiority over military and farm horses. The statistical reliability of these differences is not reported. Good retention was evidenced after a period of several months. (PsycINFO Database Record (c) 2012 APA, all rights reserved)
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3613
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	Author	Hanggi, E.B.
	Title	*Categorization Learning in Horses (Equus caballus)*			Type	Journal Article
	Year	1999	Publication	Journal of Comparative Psychology	Abbreviated Journal	J. Comp. Psychol.
	Volume	113	Issue	3	Pages	243-252
	Keywords
	Abstract	Categorization learning was investigated in 2 horses (Equus caballus). Both horses learned to select a 2-dimensional black stimulus with an open center instead of a filled stimulus in a 2-choice discrimination task. After a criterion of 10 out of 10 correct responses in a random series for 2 consecutive sessions was reached, 15 additional pairs of open-center versus filled stimuli were tested. Each was run to criterion and then incorporated into sessions of randomly mixed problems. Both horses solved the 1st problem by simple pattern discrimination and showed evidence of categorical processing for subsequent problems. New pairs were learned with few or no errors, and correct responses on novel trials were significantly above chance. These results suggest that the horses were making their selections on the basis of shared characteristics with the training stimuli and were using categorization skills in problem solving.
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	Notes				Approved	yes
	Call Number	Equine Behaviour @ team @			Serial	3678
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	Author	McCall, C.A.; Salters, M.A.; Simpson, S.M.
	Title	Relationship between number of conditioning trials per training session and avoidance learning in horses			Type	Journal Article
	Year	1993	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	36	Issue	4	Pages	291-299
	Keywords	Horse; Learning; Avoidance conditioning
	Abstract	Sixteen horses were used to determine if number of trials given per training session (5, 10, 15 or 20) affected learning performance in an avoidance conditioning task. The horse had to move from one side of a test pen to the other during an auditory cue presentation to avoid aversive stimulation. A pen 8 mx3.6 m, divided into two equal sections by a 13-cm diameter plastic pipe lying on the ground, was used as the test pen. Painted plywood panels were fastened to the fence in half the pen to help horses distinguish visually between the two parts. A 10-s auditory cue was used as a signal for horses to move from one side of the test pen to the other. A 20-s intertrial interval was used. Training sessions were conducted every third day. Each trial was recorded as an avoidance (the horse completed the task during auditory cue presentation and avoided aversive stimulus) or an error (the horse received aversive stimulus). After completing ten consecutive avoidances (criterion), the horse was removed from the study. Numbers of training sessions, trials, avoidances and errors until reaching criterion were recorded for each horse. Horses varied greatly within these variables with ranges of 3-18 sessions, 37-121 trials, 20-68 avoidances and 17-53 errors to criterion. No differences were detected (P>0.05) in the number of conditioning trials per training session (treatment) for the mean number of trials, avoidances or errors to criterion. Number of training sessions to criterion differed (P<0.01) among treatments, indicating that an optimum number of learning trials per training session might exist. Mean sessions to criterion for horses receiving 5, 10, 15 and 20 trials per session were 15.1+/-1.3, 5.8+/-1.1, 5.3+/-1.1 and 4.6+/-1.1, respectively. Regression analysis indicated that 16.2 trials per training session would minimize number of sessions to criterion. Although it is widely assumed that learning efficiency in horses is decreased when intense activity is concentrated into a small number of sessions, these results indicate that moderate repetition of training activities is needed for efficient learning.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3686
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	Author	Sueur, C.; Jacobs, A.; Amblard, F.; Petit, O.; King, A.J.
	Title	How can social network analysis improve the study of primate behavior?			Type	Journal Article
	Year	2010	Publication	American Journal of Primatology	Abbreviated Journal	Am. J. Primatol.
	Volume	73	Issue	8	Pages	703-719
	Keywords	interaction; association; social system; social structure; methodology; behavioral sampling
	Abstract	Abstract When living in a group, individuals have to make trade-offs, and compromise, in order to balance the advantages and disadvantages of group life. Strategies that enable individuals to achieve this typically affect inter-individual interactions resulting in nonrandom associations. Studying the patterns of this assortativity using social network analyses can allow us to explore how individual behavior influences what happens at the group, or population level. Understanding the consequences of these interactions at multiple scales may allow us to better understand the fitness implications for individuals. Social network analyses offer the tools to achieve this. This special issue aims to highlight the benefits of social network analysis for the study of primate behaviour, assessing it's suitability for analyzing individual social characteristics as well as group/population patterns. In this introduction to the special issue, we first introduce social network theory, then demonstrate with examples how social networks can influence individual and collective behaviors, and finally conclude with some outstanding questions for future primatological research. Am. J. Primatol. 73:703?719, 2011. ? 2011 Wiley-Liss, Inc.
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	Publisher	Wiley-Blackwell	Place of Publication		Editor
	Language		Summary Language		Original Title
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	Series Volume		Series Issue		Edition
	ISSN	0275-2565	ISBN		Medium
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	Notes	doi: 10.1002/ajp.20915			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6410
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	Author	de Waal, F. B.; Tyack, P., (eds)
	Title	Animal Social Complexity: Intelligence, Culture, and Individualized Societies			Type	Book Whole
	Year	2003	Publication		Abbreviated Journal
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	Publisher	Harvard University Press	Place of Publication	Cambridge, Massachusetts	Editor	de Waal, F. B.; Tyack, P.,
	Language		Summary Language		Original Title
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4096
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	Author	Laland K.N.
	Title	Social learning strategies			Type	Journal Article
	Year	2004	Publication	Learning & Behavior	Abbreviated Journal	Learn. Behav.
	Volume	32	Issue		Pages	4-14
	Keywords
	Abstract	In most studies of social learning in animals, no attempt has been made to examine the nature of the strategy adopted by animals when they copy others. Researchers have expended considerable effort in exploring the psychological processes that underlie social learning and amassed extensive data banks recording purported social learning in the field, but the contexts under which animals copy others remain unexplored. Yet, theoretical models used to investigate the adaptive advantages of social learning lead to the conclusion that social learning cannot be indiscriminate and that individuals should adopt strategies that dictate the circumstances under which they copy others and from whom they learn. In this article, I discuss a number of possible strategies that are predicted by theoretical analyses, including copy when uncertain, copy the majority, and copy if better, and consider the empirical evidence in support of each, drawing from both the animal and human social learning literature. Reliance on social learning strategies may be organized hierarchically, their being employed by animals when unlearned and asocially learned strategies prove ineffective but before animals take recourse in innovation.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4193
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	Author	Wood, J.N.; Glynn, D.D.; Phillips, B.C.; Hauser, M.D.
	Title	online material			Type	Miscellaneous
	Year	2007	Publication	Science	Abbreviated Journal	Science
	Volume	317	Issue	5843	Pages	1402-1405
	Keywords
	Abstract	Humans are capable of making inferences about other individuals' intentions and goals by evaluating their actions in relation to the constraints imposed by the environment. This capacity enables humans to go beyond the surface appearance of behavior to draw inferences about an individual's mental states. Presently unclear is whether this capacity is uniquely human or is shared with other animals. We show that cotton-top tamarins, rhesus macaques, and chimpanzees all make spontaneous inferences about a human experimenter's goal by attending to the environmental constraints that guide rational action. These findings rule out simple associative accounts of action perception and show that our capacity to infer rational, goal-directed action likely arose at least as far back as the New World monkeys, some 40 million years ago.
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	Notes	10.1126/science.1144663			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4242
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	Author	Müller, A. E.; Thalmann, U.
	Title	Origin and evolution of primate social organisation: a reconstruction			Type	Journal Article
	Year	2000	Publication	Biological Reviews	Abbreviated Journal
	Volume	75	Issue		Pages	405-435
	Keywords	social organisation; evolution; ancestral primate; strepsirhines; nocturnal prosimians; lemurs; lorisiforms; dispersed multi-male system; promiscuity.
	Abstract	Abstract The evolution and origin of primate social organisation has attracted the attention of many researchers, and a solitary pattern, believed to be present in most nocturnal prosimians, has been generally considered as the most primitive system. Nocturnal prosimians are in fact mostly seen alone during their nightly activities and therefore termed “solitary foragers”, but that does not mean that they are not social. Moreover, designating their social organisation as “solitary”, implies that their way of life is uniform in all species. It has, however, emerged over the last decades that all of them exhibit not only some kind of social network but also that those networks differ among species. There is a need to classify these social networks in the same manner as with group-living (gregarious) animals if we wish to link up the different forms of primate social organisation with ecological, morphological or phylogenetic variables. In this review, we establish a basic classification based on spatial relations and sociality in order to describe and cope properly with the social organisation patterns of the different species of nocturnal prosimians and other mammals that do not forage in cohesive groups. In attempting to trace the ancestral pattern of primate social organisation, the Malagasy mouse and dwarf lemurs and the Afro-Asian bushbabies and lorises are of special interest because they are thought to approach the ancestral conditions most closely. These species have generally been believed to exhibit a dispersed harem system as their pattern of social organisation (“dispersed” means that individuals forage solitarily but exhibit a social network). Therefore, the ancestral pattern of primate social organisation was inferred to be a dispersed harem. In fact, new field data on cheirogaleids combined with a review of patterns of social organisation in strepsirhines (lemurs, bushbabies and lorises) revealed that they exhibit either dispersed multi-male systems or dispersed monogamy rather than a dispersed harem system. Therefore, the concept of a dispersed harem system as the ancestral condition of primate social organisation can no longer be supported. In combination with data on social organisation patterns in “primitive” placentals and marsupials, and in monotremes, it is in fact most probable that promiscuity is the ancestral pattern for mammalian social organisation. Subsequently, a dispersed multi-male system derived from promiscuity should be regarded as the ancestral condition for primates. We further suggest that the gregarious patterns of social organisation in Aotus and Avahi, and the dispersed form in Tarsius evolved from the gregarious patterns of diurnal primates rather than from the dispersed nocturnal type. It is consequently proposed that, in addition to Aotus and Tarsius, Avahi is also secondarily nocturnal.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4257
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