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Author Keil, N.M., Sambraus, H.H.
Title “Intervenors” in agonistic interactions amongst domesticated goats Type Journal Article
Year 1998 Publication Zeitschrift fur Säugetierkunde Abbreviated Journal Z. Säugetierk.
Volume 63 Issue 5 Pages 266-272
Keywords Behaviour; Domestication; Goat; Intervention; Rank order
Abstract Social behaviour was observed in individually marked goats in two herds. The goats from one herd (n = 98) were horned, those of the other herd (n = 83) were polled. By recording agonistic interactions within the herds, a dominance index was determined for each animal. In both herds, intervention took place. Intervention is defined as one animal pushing in between two fighters, and thus ending the fight. More cases of intervention took place per individual animal amongst the horned goats than amongst the polled ones. Goats which intervened in fights on several occasions usually had a high dominance index. Members of the herd which were observed intervening only once had an average dominance index in both herds of almost 0.5. In some cases, goats very low in the rank order intervened a fight. Only rarely did the intervenors have a lower dominance index than the two fighters. In 103 cases, the direct dominance relationship between a fighting animal and the intervenor was known. In 95 cases (92.2%), the intervenor was dominant to the herd member in this fight and in just eight cases (7.8%), it was subordinate. It could not be determined what advantage the intervenor gained from its activity. It is possible that, at least in certain cases, a particularly relationship existed between the intervenor and one of the fighters.
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Call Number Equine Behaviour @ team @ Serial 5236
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Author Krueger, K.; Flauger, B.
Title Social feeding decisions in horses (Equus caballus) Type Journal Article
Year 2008 Publication Behavioural Processes Abbreviated Journal Behav. Process.
Volume 78 Issue 1 Pages 76-83
Keywords Feeding decision; Horse; Rank; Social behaviour
Abstract Like many other herbivores, in a natural environment equids feed on rather evenly distributed resources. However, the vegetation in their vast habitats constantly changes. If food is plentiful only little competition occurs over food, and in non-competitive situations domestic horses tend to return to the same feeding site until it is overgrazed. In contrast, they compete over limited food for which the social status of the individuals appears to be important. Especially in ruminants several studies have proved an influence of social organisations, rank, sex and the depletion of feeding sites on the feeding behaviour of individuals. However, it is not yet understood whether and how social aspects affect horses“ feeding decisions. Curiosity about the influence of social rank on the horses” feeding decisions between two, equally with high-quality surplus food-filled buckets placed in different social feeding conditions, led us to create the test below. The observer horses were alternately tested with a dominant and a subordinate demonstrator placed in one of the three different positions. We conclude that domestic horses use social cognition and strategic decision making in order to decide where to feed in a social feeding situation. When possible they tend to return to the same, continuously supplied feeding site and switch to an “avoidance tendency” in the presence of dominant horses or when another horse is already feeding there. Thus, the social rank and the position of conspecifics affect the feeding strategy of horses.
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 4394
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Author Krueger, K.; Flauger, B.; Farmer, K.; Hemelrijk, C.
Title Movement initiation in groups of feral horses Type Journal Article
Year 2014 Publication Behavioural Processes Abbreviated Journal Behav. Process.
Volume 103 Issue Pages 91-101
Keywords Horse; Equus ferus caballus; Distributed leadership; Herding; Departure; Rank
Abstract Abstract Herds of ungulates, flocks of birds, swarms of insects and schools of fish move in coordinated groups. Computer models show that only one or very few animals are needed to initiate and direct movement. To investigate initiation mechanisms further, we studied two ways in which movement can be initiated in feral horses: herding, and departure from the group. We examined traits affecting the likelihood of a horse initiating movement i.e. social rank, affiliative relationships, spatial position, and social network. We also investigated whether group members join a movement in dominance rank order. Our results show that whereas herding is exclusive to alpha males, any group member may initiate movement by departure. Social bonds, the number of animals interacted with, and the spatial position were not significantly associated with movement initiation. We did not find movement initiation by departure to be exclusive to any type of individual. Instead we find evidence for a limited form of distributed leadership, with higher ranking animals being followed more often.
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Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0376-6357 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5738
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Author Krueger, K.; Schneider, G.; Flauger, B.; Heinze, J.
Title Context-dependent third-party intervention in agonistic encounters of male Przewalski horses Type Journal Article
Year 2015 Publication Behavioural Processes Abbreviated Journal Behav. Process.
Volume 121 Issue Pages 54-62
Keywords Equus ferus przewalskii; Group conflict; Rank orders; Social bonds; Social control; Third-party intervention
Abstract Abstract One mechanism to resolve conflict among group members is third party intervention, for which several functions, such as kin protection, alliance formation, and the promotion of group cohesion have been proposed. Still, empirical research on the function of intervention behaviour is rare. We studied 40 cases of intervention behaviour in a field study on 13 semi-wild bachelor horses (Equus ferus przewalskii) in (a) standard social situations, and (b) when new horses joined the group (i.e. introductions). Only interventions in agonistic encounters were analysed. Eight of 13 animals directed intervention behaviour toward threatening animal in agonistic encounters of group members. One stallion was particularly active. The stallions did not intervene to support former group mates or kin and interventions were not reciprocated. In introduction situations and in standard social situations, the interveners supported animals which were lower in rank, but targeted, threatening animals of comparable social rank. After introductions, stallions received more affiliative behaviour from animals they supported and thus appeared to intervene for alliance formation. In standard social situations, interveners did not receive more affiliative behaviour from animals they supported and may primarily have intervened to promote group cohesion and to reduce social disruption within the group.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0376-6357 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5925
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Author Neumann, C.; Duboscq, J.; Dubuc, C.; Ginting, A.; Irwan, A.M.; Agil, M.; Widdig, A.; Engelhardt, A.
Title Assessing dominance hierarchies: validation and advantages of progressive evaluation with Elo-rating Type Journal Article
Year 2011 Publication Animal Behaviour Abbreviated Journal
Volume 82 Issue 4 Pages 911-921
Keywords David’s score; dominance hierarchy; dominance rank; Elo-rating; hierarchy stability; I&Si; Macaca mulatta; Macaca nigra; methodology
Abstract
Address
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Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0003-3472 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5677
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Author Obergfell, J.
Title Einf lus s v on St ruktur e l ement en auf da s Li eg ev e rha l t en v on Pf e rden in Gruppenha l tung unt e r Be rücks i cht igung de s Ag g r e s s i ons v e rha l t ens Type Manuscript
Year 2012 Publication Abbreviated Journal
Volume Issue Pages
Keywords Pferde, Ruheverhalten, Liegeverhalten, Aggressionsverhalten, Strukturelemente, Rangordnung [horses, rest behavior, lying behavior, aggression behavior, structural elements, rank order]
Abstract Durch die vorliegende Studie wurde der Einfluss von Strukturelementen auf das Liege- und Aggressionsverhalten von Pferden in Gruppenhaltung untersucht. Die Strukturelemente sollten Rückzugsmöglichkeiten bieten, sowie in ihrem Bereich zur Aufhebung der Individualdistanz führen und dadurch die Fläche relativ vergrößern. Für die Versuche stellte das Haupt- und Landesgestüt Marbach drei unabhängige Pferdegruppen mit verschiedener Herdengröße zur Verfügung, die jeweils in Einraum- Innenlaufställen gehalten wurden. Die Datenerfassung fand zwischen 23 und sieben Uhr statt. Insgesamt wurden 366 Stunden Videomaterial ausgewertet. Beim Ruheverhalten wurden mit Hilfe des event-sampling- Verfahrens die Parameter Gesamtliegedauer, Dauer in Seitenlage, Dauer der Einzelphasen in

Seitenlage und Abliegehäufigkeit pro Nacht erfasst. Es wurden Versuchsphasen ohne

Strukturelemente und mit hängenden Planen als Strukturierung durchgeführt. Im ersten Stall

kamen außerdem über einander gestapelte Strohballen zum Einsatz. Diese Art der

Strukturierung stellte sich jedoch als nicht praktikabel heraus und führte im Vergleich zu den Planen zu einer signifikanten Verschlechterung der Gesamtliegedauer. In Stall 1 konnte man eine tendenzielle Verbesserung der Parameter Gesamtliegedauer und Gesamtdauer in Seitenlage durch das Anbringen von Planen sehen. In Stall 3 dagegen verschlechterte sich das Ruheverhalten in den Versuchsphasen mit Planen gegenüber den Versuchsphasen ohne Struktur. Die Werte der Gesamtdauer in Seitenlage nahmen signifikant ab. In Stall 2 zeigten sich keine Unterschiede in den verschiedenen Versuchsphasen. Möglicherweise ist die Wirkung der Strukturelemente auf das Liegeverhalten abhängig von der Flächengröße. Stall 1 hatte bezogen auf die Leitlinien des BMELV die größte und Stall 3 die kleinste Fläche. Bei den anderen Parametern des Liegeverhaltens gab es keine signifikanten Unterschiede in den verschiedenen Versuchsphasen. Mit Hilfe des time-sampling-Verfahrens wurde die Anzahl gleichzeitig liegender Pferde und gleichzeitig liegender Pferde in Seitenlage bestimmt. Auch hier zeigten sich keine Unterschiede in den verschiedenen Versuchsphasen. Bei der Gegenüberstellung der Werte der Gesamtliegedauer und der Gesamtdauer in Seitenlage mit dem Alter der Pferde (Stall 1 und Stall 3) und mit dem Integrationszeitpunkt (Stall 1) konnte kein Zusammenhang festgestellt werden. Beim Aggressionsverhalten wurden mit Hilfe des

event-sampling-Verfahrens in den Ställen 2 und 3 verschiedene Arten von Aggressionen

erfasst, die dann in die drei Intensitätsgrade Low-Level-, Mid-Level- und High-Level-

Aggressionen unterteilt wurden. Neben der Anzahl wurde die Dauer der verschiedenen

Aggressionen bewertet. Insgesamt konnte eine positive Wirkung der Planen auf das

Aggressionsverhalten beobachtet werden. Die Gesamtanzahl an Aggressionen nahm in beiden

Ställen tendenziell in den Versuchsphasen mit Planen ab. In Stall 3 konnte, wenn man die

Aggressionen stundenweise betrachtet, ein signifikanter Unterschied festgestellt werden.

Auch der Hinterhandschlag und die Aggressionen, welche das Ruheverhalten stören,

verringerten sich tendenziell nach dem Anbringen von Strukturelementen. In beiden Ställen

nahm die relative Häufigkeit von Mid-Level-Aggressionen nach dem Anbringen von Planen

zu. Dagegen konnte bei den High-Level-Aggressionen und in Stall 3 bei den Low-Level-

Aggressionen eine relative Abnahme beobachtet werden. Sowohl die Anzahl als auch die

Dauer der Mid-Level-Aggressionen verringerten sich in Stall 3 stundenweise betrachtet

signifikant in den Versuchsphasen mit Planen. In Stall 2 war bei den High-Level-

Aggressionen sowohl bei der Dauer als auch bei der Anzahl eine signifikante Abnahme zu

sehen. Wenn man das Aggressionsverhalten in Bezug zu der Fläche in den zwei Ställen

betrachtet, schien diese vor allem einen Einfluss auf die High-Level-Aggressionen zu

nehmen. Mit Hilfe des Rangindex der Pferde im Stall 3 wurde eine Rangordnung aufgestellt.

Zwischen dem Platz der Pferde in der Rangordnung und den Parametern des Ruheverhaltens

(Gesamtliegedauer, Gesamtdauer in Seitenlage) sowie dem Alter der Pferde konnte kein

Zusammenhang festgestellt werden. Zwischen dem Rangindex und der Gesamtanzahl an

Aggressionen bestand dagegen ein hoch signifikanter Zusammenhang. Im Rahmen dieser

Studie ist das Anbringen von Strukturelementen in Bezug auf das Aggressionsverhalten in

Einraum-Innenlaufställen von Pferden zu empfehlen. Die Wirkung auf das Liegeverhalten der

Pferde sollte in weiteren Studien untersucht werden.

[In this study the influence of structural elements on lying and aggression behavior in horses

kept in groups was exanimated. Structural elements should offer the possibility of retreat.

Furthermore, within the respective area, the individual distance should be limited, thus

increasing the overall space relatively. For the studies the “Haupt- und Landesgestüt

Marbach” provided three non-related groups of different sized horses, each kept in separate

“Einraum-Innenlaufställen”. Data collection took place between 11 pm and 7 am. Altogether,

366 hours of video material were analyzed. Using the event-sampling-method, the following

parameters within the horses´ resting behavior were determined: The total lying period, the

total lying period in lateral position, the periods of the single lying in lateral position and the

frequency of lying down. Test phases were carried out without structural elements and with

hanging canvases as structural elements. In the first stable there were also three bales of straw

piled up on top of each other. This kind of structuring emerged as impractical and in

comparison to the canvas, led to a significant deterioration to the total period of lying. After

the canvases were fixed in the first stable, a slight improvement with the parameters of total

lying period and total lying period in the lateral position could be seen. In contrast, the resting

behavior in stable 3 worsened in the test phases with the canvases in comparison to the test phases without structural elements. The values for the total lying period in the lateral position

decreased significantly. In stable 2, there were no differences in the different test phases.

Possibly the effect of the structural elements is dependent of the space. Stable 1 had the

largest space and stable 3 the least space referred to the guideline of the BMELV. The other

parameters of the lying behavior showed no significant differences in the different test phases.

The number of horses lying at the same time, and the number of horses lying at the same time

in the lateral position were detected with the aid of the time-sampling-method. There were

also no differences between the different test phases. If the values of the total lying period and

the total lying period in lateral position were compared to the age of the horses (stable 1 and

stable 3) and to the point of integration (stable 1), there was no correlation.

Different kinds of aggression were detected by the means of the event-sampling-method in

stable 2 and 3 and were subdivided into the three levels of intensity: Low-level-, mid-level

and high-level-aggressions. Both the number and the duration of the different aggressions

were evaluated. All in all a positive influence of the canvas on the aggression behavior could

be observed. In both stables, the total number of aggressions decreased in the test phases with

canvases. In stable 3 a significant difference could be detected, if the aggressions were

observed per hour. Also hind limb kicks and aggressions disturbing the resting behavior,

decreased by trend, after fixing the canvases in the stable. In both stables the relative

frequency of mid-level-aggressions increased after the canvases were mounted. In contrast

there was a relative decrease at the high-level-aggressions and in stable 3 at the low-levelaggressions.

The number and the duration of the mid-level-aggressions in stable 3 decreased,

when considered by the hour in the test phases with canvases. In stable 2 a significant

decrease in duration and number of high-level-aggressions was observed. When examining

aggression behavior in reference to space in the two stables, high-level-aggressions seemed to

be mostly influenced. In stable 3 a rank order was established with the aid of the rank index of

the horses. There was no correlation between the horses´ rank and the parameters of the

resting behavior (the total lying period, the total lying period in lateral position) and between

horses´ rank and age. In contrast, there was a high significant correlation between the rank

index and the total number of aggressions. In context to this study, structural elements in

“Einraum-Innenlaufställen” (BMELV 2009) seem to have a positive influence on the

aggression behavior and thus seem to be recommendable. The influence on the lying behavior

should be investigated in further studies.]
Address
Corporate Author Thesis Ph.D. thesis
Publisher Place of Publication Karlsruhe Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5772
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Author Schneider, G.; Krueger, K.
Title Third-party interventions keep social partners from exchanging affiliative interactions with others Type Journal Article
Year 2012 Publication Animal Behaviour Abbreviated Journal Anim. Behav.
Volume 83 Issue 2 Pages 377-387
Keywords Equus caballus; horse; rank; social bond; social network; third-party intervention
Abstract Third-party interventions are defined as the interruption of dyadic interactions by third animals through direct physical contact, interposing or threats. Previous studies focused on the analysis of interventions against agonistic encounters. However, there have been no evaluations of interventions against affiliative behaviours, particularly in relation to the intervening animal�s social relationships and its social and spatial position. Horses, Equus caballus, are an interesting model species, as interventions against affiliative interactions occur more frequently than against agonistic interactions. In this study, 64 feral horses displayed 67 interventions in affiliative interactions and eight interventions in agonistic interactions within the observation period. We analysed the interventions in affiliative encounters, and found that it was mainly higher-ranking females that intervened in the affiliative interactions of group mates in the stable horse harems. The intervening animals took an active part in affiliative and agonistic encounters within the group, but did not occupy particular social roles or spatial positions. They intervened in affiliative interactions in which group mates with which they had social bonds interacted with other members of the group. They targeted the nonbonded animal and approached the one with which they were socially bonded. We suggest some species use third-party interventions in affiliative interactions to prevent competition for preferred social interaction partners from escalating into more costly agonistic encounters.
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Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0003-3472 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5492
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Author Smith, J.E.; Kolowski, J.M.; Graham, K.E.; Dawes, S.E.; Holekamp, K.E.
Title Social and ecological determinants of fission-fusion dynamics in the spotted hyaena Type Journal Article
Year 2008 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume 76 Issue 3 Pages 619-636
Keywords competition; conflict resolution; cooperative hunting; Crocuta crocuta; ecological constraints; group living; social rank
Abstract Theory predicts that individuals living in fission-fusion societies, in which group members frequently change subgroups, should modify grouping patterns in response to varying social and environmental conditions. Spotted hyaenas, Crocuta crocuta, are long-lived carnivores that reside in permanent social groups called clans. Clans are complex, fission-fusion societies in which individual members travel, rest and forage in subgroups that frequently change composition. We studied two clans in Kenya to provide the first detailed description of fission-fusion dynamics in this species. Because social and ecological circumstances can influence the cohesiveness of animal societies, we evaluated the extent to which specific circumstances promote the formation of subgroups of various sizes. We found that cooperative defence of shared resources during interclan competition and protection from lions were cohesive forces that promoted formation of large subgroups. We also tested hypotheses suggesting factors limiting subgroup size. Mothers with small cubs avoided conspecifics, thereby reducing infanticide risk. Victims of aggression either reconciled fights or separated from former opponents to reduce the immediate costs of escalated aggression in the absence of food. As predicted by the ecological constraints hypothesis, hyaenas adjusted their grouping patterns over both short and long time scales in response to feeding competition. Crocuta were most gregarious during periods of abundant prey, joined clanmates at ephemeral kills in numbers that correlated with the energetic value of the prey and gained the most energy when foraging alone because cooperative hunting attracted numerous competitors. Overall, our findings indicate that resource limitation constrains grouping in this species.
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Publisher Place of Publication Editor
Language Summary Language Original Title
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Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 4676
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Author Stashak, T. S.; Wissdorf, H.
Title Adams' Lahmheit bei Pferden Type Book Whole
Year 2007 Publication Abbreviated Journal
Volume Issue Pages
Keywords Pferd / Anatomie, Krankheiten, Heilkunde | Innere Krankheit (Tier) | Krankheit / Tierkrankheit | Pathologie (der Tiere) ( Tierkrankheit ) | Pathologie (der Tiere) / Tierkrankheit | Tierkrankheit – Pathologie (der Tiere)
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Address
Corporate Author Thesis
Publisher Schaper M. & H Place of Publication Hannover Editor
Language German Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN 3794402197 Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 4447
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Author Taillon, J.; Côté, S.
Title Are faecal hormone levels linked to winter progression, diet quality and social rank in young ungulates ? An experiment with white-tailed deer ( Odocoileus virginianus ) fawns Type Journal Article
Year 2008 Publication Behavioral Ecology and Sociobiology Abbreviated Journal Behav. Ecol. Sociobiol.
Volume 62 Issue 10 Pages 675-677
Keywords Diet quality – Glucocorticoids – Social rank – Testosterone – White-tailed deer
Abstract Abstract Hormones play a central role in the physiology and behaviour of animals. The recent development of noninvasive techniques has increased information on physical and social states of individuals through hormone measurements. The relationships among hormones, life history traits and behaviours are, however, still poorly known. For the first time, we evaluated natural winter glucocorticoid and testosterone levels in young ungulates in relation to winter progression, diet quality and social rank. Overwinter, levels of glucocorticoid and testosterone decreased, possibly due to the decline of fawns" body mass. The relationships between hormone levels and diet quality were surprising: Fawns fed the control diet presented higher glucocorticoid and lower testosterone levels then fawns fed the poor diet, suggesting that control fawns faced a higher nutritional stress than those on the poor diet. Similarly to other studies on social mammals, we found no relationship between faecal glucocorticoid levels and social rank, suggesting that social stress was similar for dominant and subordinate fawns during winter. Testosterone levels were not correlated to social rank as found previously in groups of individuals forming stable social hierarchies and maintaining stable dominance relationships. The simultaneous suppression of glucocorticoid and testosterone levels suggests for the first time that young ungulates present a hormonal strategy to prevent fast depletion of limited proteins and fat resources during winter.
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Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
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ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 4423
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