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Author Suagee-Bedore, J.K.; Linden, D.R.; Bennett-Wimbush, K. url  doi
openurl 
  Title Effect of Pen Size on Stress Responses of Stall-Housed Horses Receiving One Hour of Daily Turnout Type Journal Article
  Year 2021 Publication Journal of Equine Veterinary Science Abbreviated Journal J. Equine Vet. Sci.  
  Volume 98 Issue Pages 103366  
  Keywords Agonistic behaviors; Cortisol; Group turnout; Paddock sizes  
  Abstract Group turnout provides important socializing opportunities for horses, particularly those that are primarily stalled. A high percentage of equine injuries occur during group turnout, which could partly be due to the physical constraints of fencing. To investigate appropriate paddock sizes for group turnouts, horses (n = 12) from a single herd were divided into groups of 4, stalled for 24 hours, and then turned out for 1 hour into one of three differently sized pens: 342, 263, and 184 m2 per horse. Groups rotated through pens across 3 days, receiving one treatment per day. Blood was sampled for cortisol concentrations at 08:00 hours each morning, and then at 15 and 60 minutes into the turn out sessions, and 60 minutes after return to individual stalls. Groups rotated through three turnout times: 09:00, 12:00, 14:00 hours. Counts of agonistic behaviors (chasing, contact biting, and kicking) and low-level threats (pinned ears, tail swishing, bite and kick threats) were recorded. When turned out in pens that provided 342 m2 per horse, horses exhibited reduced plasma cortisol concentrations by 15 minutes after turnout and at 1 hour after return to their stalls (P < .05). Horses in pens providing 184 m2 per horse exhibited greater agonistic (P < .001) and low-level threat (P < .01) behaviors than horses in larger pens. These data provide insight into appropriate pen sizes for horses from established herds. Providing at least 342 m2 per horse may reduce the chance of injury in horses accustomed to group turnout.  
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  Series Volume Series Issue Edition  
  ISSN 0737-0806 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6694  
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Author Christensen, J.W.; Søndergaard, E.; Thodberg, K.; Halekoh, U. url  doi
openurl 
  Title Effects of repeated regrouping on horse behaviour and injuries Type Journal Article
  Year 2011 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 133 Issue 3 Pages 199-206  
  Keywords Group housing; Horse; Injuries; Regrouping; Social behaviour  
  Abstract Domestic horses are faced with social challenges throughout their lives due to limitations in social contact, space restrictions and frequent changes in social companionship. This is in contrast to natural conditions where horses live in relatively stable harem bands. Currently, little is known about how repeated regrouping affect horse behaviour and welfare, and it is unknown whether horses may adapt to regrouping. In this study, we aimed to investigate the effects of an unstable group structure, caused by weekly regroupings, on behaviour and frequency of injuries in young horses. Forty-five horses were included in the study and were randomly assigned to the treatments; Stable (S; seven groups of three horses) or Unstable (U; eight groups of three horses). The experimental period lasted 7 weeks, during which horses in Stable groups remained in the same group, whereas one horse was exchanged between Unstable groups every week. The groups were kept in 80m×80m grass-covered enclosures and were fed additional roughage on the ground daily. Social interactions were recorded in Unstable groups immediately after each regrouping (30min), and in both Stable and Unstable groups on day 1, 3 and 6 after each regrouping (2×20min/group/day). Injuries were scored by the end of the experimental period. The level of aggression shown by horses in Unstable groups immediately after regrouping was not affected by week (F5,35=0.42, P=0.83), indicating that horses neither habituated, nor sensitized, to repeated regrouping. Compared to horses in Stable groups, more agonistic behaviour was shown by horses in Unstable groups (i.e. non-contact agonistic; F1,65=5.60, P=0.02), whereas there was no treatment effect on other variables. The level of play behaviour appeared, however, to be more variable in Unstable groups. There was a significant effect of week on the level of contact agonistic interactions as well as greeting behaviour, due to a high occurrence in weeks 4-6. Non-contact agonistic interactions constituted the major part of agonistic interactions (66%). Possibly as consequence, no serious injuries were registered and there was no treatment effect (U=184; P=0.11). We conclude that the behaviour of young horses is affected by group management, and that horses appear not to adapt to weekly regroupings.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0168-1591 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6605  
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Author A. Wiggins; K. Crowston doi  openurl
  Title From Conservation to Crowdsourcing: A Typology of Citizen Science Type Conference Article
  Year 2011 Publication 2011 44th Hawaii International Conference on System Sciences Abbreviated Journal 2011 44th Hawaii International Conference on System Sciences  
  Volume Issue Pages 1-10  
  Keywords groupware; natural sciences computing; research and development; social sciences; crowdsourcing; citizen science typology; research collaboration; scientific research projects; virtual collaboration; Communities; Education; Monitoring; Collaboration; Organizations; Biological system modeling; Production  
  Abstract Citizen science is a form of research collaboration involving members of the public in scientific research projects to address real-world problems. Often organized as a virtual collaboration, these projects are a type of open movement, with collective goals addressed through open participation in research tasks. Existing typologies of citizen science projects focus primarily on the structure of participation, paying little attention to the organizational and macrostructural properties that are important to designing and managing effective projects and technologies. By examining a variety of project characteristics, we identified five types-Action, Conservation, Investigation, Virtual, and Education- that differ in primary project goals and the importance of physical environment to participation.  
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  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title 2011 44th Hawaii International Conference on System Sciences  
  Series Volume Series Issue Edition  
  ISSN 1530-1605 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6430  
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Author Boogert, N.J.; Reader, S.M.; Hoppitt, W.; Laland, K.N. url  doi
openurl 
  Title The origin and spread of innovations in starlings Type Journal Article
  Year 2008 Publication Animal Behaviour Abbreviated Journal Anim. Behav.  
  Volume 75 Issue 4 Pages 1509-1518  
  Keywords diffusion dynamics; dominance; foraging; group; innovation; neophobia; social learning; social network; starling; Sturnus vulgaris  
  Abstract There are numerous reports of novel learned behaviour patterns in animal populations, yet the factors influencing the invention and spread of these innovations remain poorly understood. Here we investigated to what extent the pattern of spread of innovations in captive groups of starlings, Sturnus vulgaris, could be predicted by knowledge of individual and social group variables, including association patterns, social rank orders, measures of neophobia and asocial learning performance. We presented small groups of starlings with a series of novel extractive foraging tasks and recorded the latency for each bird to contact and solve each task, as well as the orders of contacting and solving. We then explored which variables best predicted the observed diffusion patterns. Object neophobia and social rank measures characterized who was the first of the group to contact the novel foraging tasks, and the subsequent spread of contacting tasks was associated with latency to feed in a novel environment. Asocial learning performance, measured in isolation, predicted who was the first solver of the novel foraging tasks in each group. Association patterns did not predict the spread of solving. Contact latency and solving duration were negatively correlated, consistent with social learning underlying the spread of solving. Our findings indicate that we can improve our understanding of the diffusion dynamics of innovations in animal groups by investigating group-dependent and individual variables in combination. We introduce novel methods for exploring predictors of the origin and spread of behavioural innovations that could be widely applied.  
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  Series Volume Series Issue Edition  
  ISSN 0003-3472 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6036  
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Author Krueger, K.; Schneider, G.; Flauger, B.; Heinze, J. doi  openurl
  Title Context-dependent third-party intervention in agonistic encounters of male Przewalski horses Type Journal Article
  Year 2015 Publication Behavioural Processes Abbreviated Journal Behav. Process.  
  Volume 121 Issue Pages 54-62  
  Keywords Equus ferus przewalskii; Group conflict; Rank orders; Social bonds; Social control; Third-party intervention  
  Abstract Abstract One mechanism to resolve conflict among group members is third party intervention, for which several functions, such as kin protection, alliance formation, and the promotion of group cohesion have been proposed. Still, empirical research on the function of intervention behaviour is rare. We studied 40 cases of intervention behaviour in a field study on 13 semi-wild bachelor horses (Equus ferus przewalskii) in (a) standard social situations, and (b) when new horses joined the group (i.e. introductions). Only interventions in agonistic encounters were analysed. Eight of 13 animals directed intervention behaviour toward threatening animal in agonistic encounters of group members. One stallion was particularly active. The stallions did not intervene to support former group mates or kin and interventions were not reciprocated. In introduction situations and in standard social situations, the interveners supported animals which were lower in rank, but targeted, threatening animals of comparable social rank. After introductions, stallions received more affiliative behaviour from animals they supported and thus appeared to intervene for alliance formation. In standard social situations, interveners did not receive more affiliative behaviour from animals they supported and may primarily have intervened to promote group cohesion and to reduce social disruption within the group.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0376-6357 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5925  
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Author Burla,J.B.; Rufener, C.; Bachmann, I.; Gygax, L.; Patt, A.; Hillmann, E. pdf  openurl
  Title Effect of varying dimensions of the littered lying area on the lying behaviour of group-housed horses (Equus ferus caballus) Type Conference Article
  Year 2015 Publication Proceedings of the 3. International Equine Science Meeting Abbreviated Journal Proc. 3. Int. Equine. Sci. Mtg  
  Volume Issue Pages  
  Keywords group housing, lying behaviour, littered lying area, rubber mats  
  Abstract Although horses can sleep while standing, recumbency is required for REM sleep and since all sleep stages must be completed for an entire sleep cycle, the opportunity for recumbency is essential for animal welfare. Observations on feral horses indicate a minimal lying duration of 30 min, preferably on a deformable and dry ground. In contrast to feral horses, lying behaviour in stabled horses is often affected by the dimensions of the provided lying area and rank.

In Switzerland, minimum requirements (MR) for the littered lying area are established by law to ensure animal welfare (BLV, 2008) (A/N: approximately match German recommendations (BMEL, 2009)). The aim of this study was to assess the adequacy of the dimensions of the minimum requirements for group-housed horses by investigating 38 horses in 8 groups. Further, hard rubber mats were provided supplementary in order to assess their suitability as an alternative to litter. Four treatments were each applied in randomised order:

– 0x MR: no litter + 1.5x MR with rubber mats

– 0.5x MR: 0.5x MR with litter + 1x MR with rubber mats

– 1x MR: 1x MR with litter; 0.5x MR with rubber mats

– 1.5x MR: 1.5x MR with litter + no rubber mats

For each treatment, after a habituation period of 8 days, lying behaviour was recorded (video, accelerometers) continuously for 72 hrs. Statistical analysis was performed using mixed effects models.

Regardless of the ground chosen, the duration of recumbency per 24 hrs was increasing with increasing dimensions of the littered area (F1,93 = 12.9, p = 0.0005; Fig. 1). Whereas the effect flattened from 1x to 1.5x MR, the duration spent on litter – a deformable ground – was increasing continuously (F1,62 = 23.1, p < 0.0001). Further, the proportion of lateral recumbency was increased with increased dimensions of the littered area (F1,79 = 12.3, p = 0.0007). Regarding the number of lying bouts, no differences were apparent between treatments providing litter, but recumbency occurred very seldom if only rubber mats were provided (F1,93 = 14.7, p = 0.0002). Further, low-ranking horses spent more lying bouts on rubber mats than high-ranking horses (F1,29 = 4.4, p = 0.04). Additionally, the larger the dimensions of the littered area the more horses were present in the lying area at the moment of lying down (F1,79 = 6.6, p = 0.01). Moreover, low-ranking horses showed considerably higher percentages of involuntarily terminated lying bouts than high-ranking horses if 0.5x and 1x MR were littered (F1,76 = 8.43, p = 0.005).

Although the indicated minimal lying duration was averagely performed, large individual differences occurred and at least 8% were lying down less than 30 min per 24 hrs in every treatment. Further, the inclusion of social parameters indicated a beneficial effect of an exceedance of the minimum requirements especially for low-ranking horses. Therefore, the minimum requirements established by Swiss law can be stated as adequate but should be perceived as minimum and not optimum dimensions.
 
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  Corporate Author Burla,J.B. Thesis  
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  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
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  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5879  
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Author Wolter, R.; Pantel, N.; Stefanski, V.; Möstl, E.; Krueger, K. doi  openurl
  Title The role of an alpha animal in changing environmental conditions Type Journal Article
  Year 2014 Publication Physiology & Behavior Abbreviated Journal Physiol. Behav.  
  Volume 133 Issue Pages 236-243  
  Keywords Alpha male; Horse; Equus ferus przewalskii; Bachelor group; Group structure  
  Abstract Abstract The maintenance and development of conservation areas by grazing of large herbivores, such as Przewalski's horses, is common practice. Several nature conservation areas house male bachelor groups of this species. When males are needed for breeding they are removed from the groups, often without considering group compositions and individual social positions. However, alpha animals are needed for ensuring group stability and decision making in potentially dangerous situations in several species. To investigate the role of the alpha male in a bachelor group, we observed the behaviour of five Przewalski's horse males during the enlargement of their enclosure. We analyzed the group's social structure and movement orders, as well as the animals' connectedness, activity budgets, and whether they moved with preferred group members and how factors such as social rank influenced the horses' behaviour. We also investigated the excretion of glucocorticoid metabolites (GCM) via faeces of the horses while exploring a new area as a parameter of glucocorticoid production. Our results show that the alpha male is important for a bachelor group in changing environmental conditions. The alpha male had the highest level of connectedness within the group. When exploring the new environment, its position in the group changed from previously being the last to being the first. Furthermore the whole group behaviour changed when exploring the new area. The stallions showed reduced resting behavior, increased feeding and did not stay close to each other. We found that the excretion of glucocorticoid metabolites of most horses rose only marginally during the first days on the new area while only the alpha male showed a significant increased amount of glucocorticoid production during the first day of the enclosure enlargement.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0031-9384 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5818  
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Author Massen, J.; Sterck, E.; de Vos, H. url  openurl
  Title Close social associations in animals and humans: functions and mechanisms of friendship Type
  Year 2010 Publication Behaviour Abbreviated Journal  
  Volume 147 Issue 11 Pages 1379  
  Keywords Both humans and group-living animals associate and behave affiliatively more with some individuals than others. Human friendship has long been acknowledged, and recently scientists studying animal behaviour have started using the term friendship for close social associates in animals. Yet, while biologists describe friends as social tools to enhance fitness, social scientists describe human friendship as unconditional. We investigate whether these different descriptions reflect true differences in human friendship and animal close social associations or are a by-product of different research approaches: namely social scientists focussing on proximate and biologists on ultimate explanations. We first stress the importance of similar measures to determine close social associations, thereafter examine their ultimate benefits and proximate motivations, and discuss the latest findings on the central-neural regulation of social bonds. We conclude that both human friendship and animal close social associations are ultimately beneficial. On the proximate level, motivations for friendship in humans and for close social associations in animals are not necessarily based on benefits and are often unconditional. Moreover, humans share with many animals a similar physiological basis of sociality. Therefore, biologists and social scientist describe the same phenomenon, and the use of the term friendship for animals seems justified.  
  Abstract Both humans and group-living animals associate and behave affiliatively more with some individuals than others. Human friendship has long been acknowledged, and recently scientists studying animal behaviour have started using the term friendship for close social associates in animals. Yet, while biologists describe friends as social tools to enhance fitness, social scientists describe human friendship as unconditional. We investigate whether these different descriptions reflect true differences in human friendship and animal close social associations or are a by-product of different research approaches: namely social scientists focussing on proximate and biologists on ultimate explanations. We first stress the importance of similar measures to determine close social associations, thereafter examine their ultimate benefits and proximate motivations, and discuss the latest findings on the central-neural regulation of social bonds. We conclude that both human friendship and animal close social associations are ultimately beneficial. On the proximate level, motivations for friendship in humans and for close social associations in animals are not necessarily based on benefits and are often unconditional. Moreover, humans share with many animals a similar physiological basis of sociality. Therefore, biologists and social scientist describe the same phenomenon, and the use of the term friendship for animals seems justified.  
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  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
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  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5813  
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Author Fürst, A.; Knubben, J.; Kurtz, A.; Auer, J.; Stauffacher, M. openurl 
  Title Pferde in Gruppenhaltung: Eine Betrachtung aus tierärztlicher Sicht unter besonderer Berücksichtigung des Verletzungsrisikos [Group housing of horses: veterinary considerations with a focus on the prevention of bite and kick injuries] Type Journal Article
  Year 2006 Publication Pferdeheilkunde Abbreviated Journal  
  Volume 22 Issue 3 Pages 254-258  
  Keywords Verhalten, Gruppenhaltung, Prävention, Schlagverletzungen, Bissverletzungen, Tierschutz [Behaviour, group housing, prevention, bite injuries, kick injuries, animal protection]  
  Abstract Mit der zunehmenden Bedeutung der Gruppenhaltung von Pferden ist die Tierärzteschaft gefordert mitzuhelfen, das Verletzungsrisiko in

Gruppenhaltungssystemen zu verringern. Dem Vermeiden von Schlag- und Bissverletzungen kommt hierbei eine zentrale Bedeutung zu. Präventive

Maßnahmen konzentrieren sich im Wesentlichen auf die Gruppenzusammensetzung und Eingliederung neuer Pferde sowie auf die

Gestaltung der Haltungssysteme. Die Raumaufteilung und die Fütterungstechnik müssen equidentypisches Verhalten (Lokomotion, langandauernde

Futteraufnahme und schadensfreie soziale Interaktionen) erlauben. Es gilt, Kenntnisse über Zusammenhänge zwischen Haltung,

Fütterung, Nutzung, Verhalten und Gesundheit an Pferdehalter und Stallbaufirmen weiterzugeben.

[Although group housing of horses has become common practice, the risk of equine injury is substantial. The veterinary community is challenged

to reduce this risk, particularly with regard to injuries caused by kicking and biting. Preventive measures should focus on the disposition

of horses within the group, the introduction of new horses to the group and the design of the housing facility. Feeding methods as

well as the structure of the environment should meet the physiological requirements for horses; there should be adequate space for exercise,

extended foraging and the possibility of benign social interactions. Veterinarians need to educate horse owners and builders of equine

facilities about the husbandry, feeding, use, behaviour and health of horses.]
 
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5756  
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Author Nuñez, C.M.V.; Adelman, J.S.; Smith, J.; Gesquiere, L.R.; Rubenstein, D.I. url  doi
openurl 
  Title Linking social environment and stress physiology in feral mares (Equus caballus): Group transfers elevate fecal cortisol levels Type Journal Article
  Year 2014 Publication General and Comparative Endocrinology Abbreviated Journal  
  Volume 196 Issue Pages 26-33  
  Keywords Equus caballus; Fecal cortisol; Feral mare; Group transfer; Stress; Social instability  
  Abstract Abstract Feral horses (Equus caballus) have a complex social structure, the stability of which is important to their overall health. Behavioral and demographic research has shown that decreases in group (or band) stability reduce female fitness, but the potential effects on the physiological stress response have not been demonstrated. To fully understand how band stability affects group-member fitness, we need to understand not only behavioral and demographic, but also physiological consequences of decreases to that stability. We studied group changes in feral mares (an activity that induces instability, including both male and female aggression) on Shackleford Banks, NC. We found that mares in the midst of changing groups exhibit increased fecal cortisol levels. In addition, mares making more group transfers show higher levels of cortisol two weeks post-behavior. These results offer insights into how social instability is integrated into an animal’s physiological phenotype. In addition, our results have important implications for feral horse management. On Shackleford Banks, mares contracepted with porcine zona pellucida (PZP) make approximately 10 times as many group changes as do untreated mares. Such animals may therefore be at higher risk of chronic stress. These results support the growing consensus that links between behavior and physiological stress must be taken into account when managing for healthy, functional populations.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0016-6480 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5743  
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