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Author Benz, B.; Münzing, C.; Krueger, K.; Winter, D.
Title Ethologische Untersuchung von Heuraufen in der Pferdehaltung [Ethological investigation of hayracks in equine husbandry] Type Journal Article
Year 2014 Publication Landtechnik Abbreviated Journal Landtechnik
Volume 69 Issue 5 Pages 239-244
Keywords Pferdehaltung, Fressverhalten, Futterraufen [horse keeping, feeding behaviour, roughage racks]
Abstract Eine tiergerechte, physiologisch und anatomisch auf die Bedürfnisse der Pferde ausgerichtete

Raufutterversorgung sollte die Kaubedürfnisse und die Beschäftigungszeiten von Pferden

ausreichend berücksichtigen. Daher – und auch aufgrund des bestehenden Kostendrucks bei

Raufutter – steigt das Interesse an Raufutterraufen, durch die möglicherweise die Futteraufnahmezeiten

verlängert sowie Futterverluste minimiert werden können.

Die vorliegende Untersuchung vergleicht das Fressverhalten und die Körperhaltung von acht

Pferden beim Einsatz von drei unterschiedlichen Futterraufen mit der Bodenfütterung in Einzelboxenhaltung.

Die Ergebnisse dieser Studie lassen den Schluss zu, dass der Einsatz von

Raufutterraufen die Futteraufnahmezeiten verlängert und somit längere Beschäftigungszeiten

für die Futteraufnahme gewährleistet werden. Außerdem konnte festgestellt werden, dass die

Pferde das Raufutter bei einer der drei untersuchten Raufen überwiegend mit natürlicher Kopf-

Hals-Haltung aufnahmen.

[Regarding the species horse, an appropriate supply of roughage should take into account the

need to chew as well as the need for occupation. In this context, and due to the current cost

pressure for hay, the interest in roughage racks increases. It is assumed that roughage racks

could help to extend the feeding time and reduce food losses.

The present study places the emphasis on the observation of the feeding behaviour of eight

horses in single horse boxes. Three different roughage racks are compared to traditional feeding

on the floor. On the basis of the results it may be concluded that the use of roughage racks

extends the feeding time and thus ensures longer occupation. In one of the three roughage

racks investigated the horses mainly eat in a natural posture of their head and neck.]
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Call Number Equine Behaviour @ team @ Serial 5840
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Author Massen, J.; Sterck, E.; de Vos, H.
Title Close social associations in animals and humans: functions and mechanisms of friendship Type
Year 2010 Publication Behaviour Abbreviated Journal
Volume 147 Issue 11 Pages 1379
Keywords Both humans and group-living animals associate and behave affiliatively more with some individuals than others. Human friendship has long been acknowledged, and recently scientists studying animal behaviour have started using the term friendship for close social associates in animals. Yet, while biologists describe friends as social tools to enhance fitness, social scientists describe human friendship as unconditional. We investigate whether these different descriptions reflect true differences in human friendship and animal close social associations or are a by-product of different research approaches: namely social scientists focussing on proximate and biologists on ultimate explanations. We first stress the importance of similar measures to determine close social associations, thereafter examine their ultimate benefits and proximate motivations, and discuss the latest findings on the central-neural regulation of social bonds. We conclude that both human friendship and animal close social associations are ultimately beneficial. On the proximate level, motivations for friendship in humans and for close social associations in animals are not necessarily based on benefits and are often unconditional. Moreover, humans share with many animals a similar physiological basis of sociality. Therefore, biologists and social scientist describe the same phenomenon, and the use of the term friendship for animals seems justified.
Abstract Both humans and group-living animals associate and behave affiliatively more with some individuals than others. Human friendship has long been acknowledged, and recently scientists studying animal behaviour have started using the term friendship for close social associates in animals. Yet, while biologists describe friends as social tools to enhance fitness, social scientists describe human friendship as unconditional. We investigate whether these different descriptions reflect true differences in human friendship and animal close social associations or are a by-product of different research approaches: namely social scientists focussing on proximate and biologists on ultimate explanations. We first stress the importance of similar measures to determine close social associations, thereafter examine their ultimate benefits and proximate motivations, and discuss the latest findings on the central-neural regulation of social bonds. We conclude that both human friendship and animal close social associations are ultimately beneficial. On the proximate level, motivations for friendship in humans and for close social associations in animals are not necessarily based on benefits and are often unconditional. Moreover, humans share with many animals a similar physiological basis of sociality. Therefore, biologists and social scientist describe the same phenomenon, and the use of the term friendship for animals seems justified.
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Language Summary Language Original Title
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5813
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Author Benz, B.; Köhnke, J.; Kappelmann, K.
Title Bewertung einer Faltschieberanlage in einem Reitstall mit Paddockboxen[Assessment of a v-form scraper in a horse barn with paddock boxes] Type Journal Article
Year 2014 Publication Landtechnik, Agricultural Engineering, Abbreviated Journal
Volume 68 Issue 4 Pages 242-247
Keywords Entmistungstechnik, Arbeitswirtschaft, Pferdeverhalten [Manure removal systems, working time requirement, horse behaviour]
Abstract In der vorliegenden Untersuchung werden in einem Praxisbetrieb die Verfahrenskosten eines

Faltschiebers erhoben. Aufgrund des reduzierten Arbeitszeitaufwandes ergibt sich durch den

Einsatz des Faltschiebers eine jährliche Kostenersparnis in Höhe von 78 € je Pferd. Durch die

Mechanisierung der Entmistung kann fast 30 % der Arbeitszeit in der Pensionspferdehaltung

eingespart werden. Beim Einsatz einer Entmistungstechnik spielt jedoch nicht nur die Ökonomie,

sondern darüber hinaus auch das Pferdeverhalten eine Rolle. Im selben Praxisbetrieb wird

nach Installation der Faltschieberanlage das Pferdeverhalten beim Erstkontakt mit dem Schieber

beobachtet. Dabei zeigt sich, dass die Pferde den direkten Kontakt mit der Entmistungstechnik

und somit kritische Situationen vermeiden.

[In the survey at hand, the procedural costs for a v-form scraper are gathered. In the process,

it is found that due to the reduced working time requirement the use of a v-form scraper

saves € 78/horse/year. The mechanization of manure removal can reduce working time in

horse keeping by almost 30 percent. However, using manure removal systems, the profitability

is not the only crucial criteria. The behaviour of the horses plays an essential role, too.

Moreover the horses’ behaviour when first encountering the manure scraper is observed. The

study reveals that the horses avoid contact with the scraper and thereby also shirk critical

situations.]
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Call Number Equine Behaviour @ team @ Serial 5841
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Author Burla,J.B.; Rufener, C.; Bachmann, I.; Gygax, L.; Patt, A.; Hillmann, E.
Title Effect of varying dimensions of the littered lying area on the lying behaviour of group-housed horses (Equus ferus caballus) Type Conference Article
Year 2015 Publication Proceedings of the 3. International Equine Science Meeting Abbreviated Journal Proc. 3. Int. Equine. Sci. Mtg
Volume Issue Pages
Keywords group housing, lying behaviour, littered lying area, rubber mats
Abstract Although horses can sleep while standing, recumbency is required for REM sleep and since all sleep stages must be completed for an entire sleep cycle, the opportunity for recumbency is essential for animal welfare. Observations on feral horses indicate a minimal lying duration of 30 min, preferably on a deformable and dry ground. In contrast to feral horses, lying behaviour in stabled horses is often affected by the dimensions of the provided lying area and rank.

In Switzerland, minimum requirements (MR) for the littered lying area are established by law to ensure animal welfare (BLV, 2008) (A/N: approximately match German recommendations (BMEL, 2009)). The aim of this study was to assess the adequacy of the dimensions of the minimum requirements for group-housed horses by investigating 38 horses in 8 groups. Further, hard rubber mats were provided supplementary in order to assess their suitability as an alternative to litter. Four treatments were each applied in randomised order:

– 0x MR: no litter + 1.5x MR with rubber mats

– 0.5x MR: 0.5x MR with litter + 1x MR with rubber mats

– 1x MR: 1x MR with litter; 0.5x MR with rubber mats

– 1.5x MR: 1.5x MR with litter + no rubber mats

For each treatment, after a habituation period of 8 days, lying behaviour was recorded (video, accelerometers) continuously for 72 hrs. Statistical analysis was performed using mixed effects models.

Regardless of the ground chosen, the duration of recumbency per 24 hrs was increasing with increasing dimensions of the littered area (F1,93 = 12.9, p = 0.0005; Fig. 1). Whereas the effect flattened from 1x to 1.5x MR, the duration spent on litter – a deformable ground – was increasing continuously (F1,62 = 23.1, p < 0.0001). Further, the proportion of lateral recumbency was increased with increased dimensions of the littered area (F1,79 = 12.3, p = 0.0007). Regarding the number of lying bouts, no differences were apparent between treatments providing litter, but recumbency occurred very seldom if only rubber mats were provided (F1,93 = 14.7, p = 0.0002). Further, low-ranking horses spent more lying bouts on rubber mats than high-ranking horses (F1,29 = 4.4, p = 0.04). Additionally, the larger the dimensions of the littered area the more horses were present in the lying area at the moment of lying down (F1,79 = 6.6, p = 0.01). Moreover, low-ranking horses showed considerably higher percentages of involuntarily terminated lying bouts than high-ranking horses if 0.5x and 1x MR were littered (F1,76 = 8.43, p = 0.005).

Although the indicated minimal lying duration was averagely performed, large individual differences occurred and at least 8% were lying down less than 30 min per 24 hrs in every treatment. Further, the inclusion of social parameters indicated a beneficial effect of an exceedance of the minimum requirements especially for low-ranking horses. Therefore, the minimum requirements established by Swiss law can be stated as adequate but should be perceived as minimum and not optimum dimensions.
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Corporate Author Burla,J.B. Thesis
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5879
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Author Skandakumar, S.; Stodulski, G.; Hau, J.
Title Salivary IgA: a Possible Stress Marker In Dogs Type Abstract
Year 1995 Publication Animal Welfare Abbreviated Journal
Volume 4 Issue 4 Pages 339-350
Keywords Animal Welfare; Behaviour; Cortisol; Dog; Salivary Iga (S-Iga); Stress; Well-Being
Abstract Stress in humans has been reported to be associated with a decrease in the salivary immunoglobulin A (s-IgA) levels enabling the possible use of s-IgA to assess stress. Prolonged stress, if reliably assessed in a non-invasive manner, may be used to assess animal welfare. This study analysed groups of dogs undergoing physical and temperamental training and s-IgA levels were measured by rocket immunoelectrophoresis in prospective samples. Behavioural assessment was carried out and cortisol levels in saliva were measured by ELISA. A significant negative correlation (P < 0.007) between the logarithmic cortisol concentrations and s-IgA levels in saliva was recorded. The behavioural assessment of the dogs agreed well with the biochemical markers. It is concluded that IgA levels in saliva may be a useful marker of dog well-being and that stress results in decreased s-IgA levels.
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5964
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Author McGreevy, P.; Yeates, J.
Title Horses (Equus caballus) Type Book Chapter
Year 2018 Publication Companion Animal Care and Welfare Abbreviated Journal
Volume Issue Pages
Keywords animal company; behavioural signs; diseases; domestic horses; euthanasia; human interaction; nutritional requirements
Abstract Summary Domestic horses are equid members of the class Mammalia, order Perissodactyla, and family Equidae. Horses are obligate herbivores, with nutritional requirements as listed in a table. Adequate space is necessary for exercise, exploration, flight, sharing resources, play, and rolling. Company is essential for all horses, including stallions. Company provides opportunities for mutual grooming and play and allows horses to stand head-to-tail to remove flies. Unhandled horses may respond to humans as they would to predators, whereas handled horses' responses depend on their previous interactions with humans. Horses can suffer from several diseases as listed in another table. The best method of euthanasia of horses is usually sedation followed by either cranial shooting or the injection of an overdose of pentobarbitone into the jugular vein. Behavioural signs of distress can include increased locomotory activity, vigilance behaviours, neighing, snorting, pawing, nibbling walls and buckets, defaecation, rearing, kicking stable walls or doors, and high-stepping 'prancing'.
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Language Summary Language Original Title
Series Editor Series Title Wiley Online Books Abbreviated Series Title Companion Animal Care and Welfare
Series Volume Series Issue Edition
ISSN (up) ISBN 9781119333708 Medium
Area Expedition Conference
Notes doi:10.1002/9781119333708.ch13 Approved no
Call Number Equine Behaviour @ team @ Serial 6506
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Author Wey, T.; Blumstein, D.T.; Shen, W.; Jordán, F.
Title Social network analysis of animal behaviour: a promising tool for the study of sociality Type Journal Article
Year 2008 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume 75 Issue 2 Pages 333-344
Keywords animal behaviour; quantifying sociality; social network analysis; social structure
Abstract Social animals live and interact together, forming complex relationships and social structure. These relationships can have important fitness consequences, but most studies do not explicitly measure those relationships. An approach that explicitly measures relationships will further our understanding of social complexity and the consequences of both direct and indirect interactions. Social network analysis is the study of social groups as networks of nodes connected by social ties. This approach examines individuals and groups in the context of relationships between group members. Application of social network analysis to animal behaviour can advance the field by identifying and quantifying specific attributes of social relationships, many of which are not captured by more common measures of sociality, such as group size. Sophisticated methods for network construction and analysis exist in other fields, but until recently, have seen relatively little application to animal systems. We present a brief history of social network analysis, a description of basic concepts and previous applications to animal behaviour. We then highlight relevance and constraints of some network measures, including results from an original study of the effect of sampling on network parameter estimates, and we end with promising directions for research. By doing so, we provide a prospective overview of social network analysis' general utility for the study of animal social behaviour.
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Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
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ISSN (up) 0003-3472 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 4691
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Author Hewitt, S.E.; Macdonald, D.W.; Dugdale, H.L.
Title Context-dependent linear dominance hierarchies in social groups of European badgers, Meles meles Type Journal Article
Year 2009 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume 77 Issue 1 Pages 161-169
Keywords David's score; directed aggression; dominance rank; European badger; grooming behaviour; linear dominance hierarchy; Meles meles; reproductive success; relatedness; social status
Abstract A social hierarchy is generally assumed to exist in those mammalian societies in which the costs and benefits of group living are distributed unevenly among group members. We analysed infrared closed-circuit television footage, collected over 3 years in Wytham Woods, Oxfordshire, to test whether social groups of European badgers have dominance hierarchies. Analysis of directed aggression between dyads revealed linear dominance hierarchies in three social-group-years, but patterns within social groups were not consistent across years. Dominance hierarchies were significantly steeper than random in five out of six social-group-years. In those social-group-years where a linear hierarchy was determined, there was an effect of sex on dominance rank, with females gaining significantly higher rank than males in two social-group-years. Overall, rank was not related to age, nor did it appear to affect the likelihood of an individual being wounded, or an individual's breeding status. The latter resulted from nonorthogonality between sex and breeding status, as there were only two breeding males. Overall, hierarchies were primarily dominated by breeding females, and may occur when breeding competition arises. Relatedness, unreciprocated allogrooming and sequential allomarking were not consistently related to levels of directed aggression across social-group-years. We suggest that dominance structures within European badger groups may be context dependent, with future study required to complete our understanding of where, and when, they arise.
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ISSN (up) 0003-3472 ISBN Medium
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 4695
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Author Smith, L.A.; Wells, K.L.; Marion, G.; Swain, D.L.; Hutchings, M.R.
Title Effects of group composition on the grazing behaviour of herbivores Type Journal Article
Year Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume In Press, Corrected Proof Issue Pages
Keywords grazing; group composition; herbivore; individual behaviour; physiological state
Abstract Animal behaviour is often a function of the animal's physiological state. Groups of animals will often contain individuals with a range of physiological states and the grazing behaviour of herbivores is affected by their physiological state. This study compared the grazing decisions of animals in groups of single and mixed physiological states. Using a grazing model that simulated individual herbivore behaviour in relation to environmental distributions of forage resource (grass) and parasites (faeces), we tested the hypothesis that an animal's level of parasite exposure via the faecal-oral route is affected by the composition of physiological states in the group. Four physiological states were considered: parasite-naïve, parasitized, lactating and parasite-immune animals. Baseline parasite exposure levels for each state were generated by simulating single-state groups and were compared to simulations of each of the six two-state combinations. In single-state groups parasitized animals had the least and lactating animals had the greatest levels of parasite exposure. When co-grazing with lactating animals, parasitized, immune and naïve animals increased their parasite exposure, relative to single-state groups. When co-grazing with parasitized animals, lactating, immune and naïve animals reduced their parasite exposure, relative to single-state groups. There was no difference in parasite exposure of the immune or naïve animals co-grazing together when compared to the single-state groups. These results highlight the need to recognize the impact of the individual when studying group-living animals.
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ISSN (up) 0003-3472 ISBN Medium
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5155
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Author Mitani, J.C.
Title Male chimpanzees form enduring and equitable social bonds Type Journal Article
Year 2009 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume 77 Issue 3 Pages 633-640
Keywords chimpanzee; Pan troglodytes; social behaviour; social relationship
Abstract Controversy exists regarding the nature of primate social relationships. While individual primates are frequently hypothesized to form enduring social bonds with conspecifics, recent studies suggest that relationships are labile, with animals interacting only over short periods to satisfy their immediate needs. Here I use data collected over 10 years on a community of chimpanzees, Pan troglodytes, at Ngogo, Kibale National Park, Uganda, to investigate whether male chimpanzees establish long-term social relationships and to determine the factors that affect variation in relationship quality and the stability of social bonds. Kinship and dominance rank influenced the quality of relationships. Maternal brothers and males of the same dominance rank class groomed each other more equitably than did unrelated males and males that were dissimilar in rank. In addition, males that formed strong social bonds groomed more equitably than did males that displayed weaker bonds. Social bonds were stable over time, with relationships in one year predicting those in subsequent years. Kinship and the quality of social relationships affected bond stability. Maternal half siblings and males that groomed each other equitably maintained longer-lasting bonds than did nonkin and males that groomed each other unevenly. Virtually all of the males established at least one enduring relationship with another individual. The most enduring bonds formed between a few pairs of maternal brothers and dyads that maintained balanced grooming interactions. These results indicate that male chimpanzees maintain long-lasting and equitable social bonds whose formation is affected by maternal kinship and the quality of social relationships.
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Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN (up) 0003-3472 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 5164
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