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Author Keiper, R.R. openurl 
  Title Social structure Type Journal Article
  Year 1986 Publication The Veterinary clinics of North America. Equine practice Abbreviated Journal Vet Clin North Am Equine Pract  
  Volume 2 Issue 3 Pages 465-484  
  Keywords Animal Communication; Animals; Animals, Domestic; Animals, Wild; Dominance-Subordination; Female; *Hierarchy, Social; Homing Behavior; *Horses; Male; Sexual Behavior, Animal; *Social Behavior; *Social Dominance  
  Abstract Socially feral horses live in stable social groups characterized by one adult male, a number of adult females, and their offspring up to 2 years of age. Extra males either live by themselves or with other males in bachelor groups. The bands occupy nondefended home ranges that often overlap. Many abnormal behaviors seen in domestic horses occur because some aspect of their normal social behavior cannot be carried out in captivity.  
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  Language English Summary Language Original Title  
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  Series Volume Series Issue Edition  
  ISSN 0749-0739 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:3492240 Approved no  
  Call Number refbase @ user @ Serial 675  
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Author Kendal, R.L.; Coe, R.L.; Laland, K.N. doi  openurl
  Title Age differences in neophilia, exploration, and innovation in family groups of callitrichid monkeys Type Journal Article
  Year 2005 Publication American journal of primatology Abbreviated Journal Am. J. Primatol.  
  Volume 66 Issue 2 Pages 167-188  
  Keywords Age Factors; Analysis of Variance; Animals; *Animals, Zoo; Behavior, Animal/*physiology; Callitrichinae/*physiology; *Creativeness; Exploratory Behavior/*physiology; Observation; Social Behavior; Task Performance and Analysis  
  Abstract The prevailing assumption in the primate literature is that young or juvenile primates are more innovative than adult individuals. This innovative tendency among the young is frequently thought to be a consequence, or side effect, of their increased rates of exploration and play. Conversely, Reader and Laland's [International Journal of Primatology 22:787-806, 2001] review of the primate innovation literature noted a greater reported incidence of innovation in adults than nonadults, which they interpreted as (in part) a reflection of the greater experience and competence of older individuals. Within callitrichids there is contradictory evidence for age differences in response to novel objects, foods, and foraging tasks. By presenting novel extractive foraging tasks to family groups of callitrichid monkeys in zoos, we examined, in a large sample, whether there are positive or negative relationships of age with neophilia, exploration, and innovation, and whether play or experience most facilitates innovation. The results indicate that exploration and innovation (but not neophilia) are positively correlated with age, perhaps reflecting adults' greater manipulative competence. To the extent that there was evidence for play in younger individuals, it did not appear to contribute to innovation. The implications of these findings for the fields of innovation and conservation through reintroduction are considered.  
  Address Sub-Department of Animal Behavior, University of Cambridge, Cambridge, UK. RachelKendal2003@yahoo.co.uk  
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  Series Volume Series Issue Edition  
  ISSN 0275-2565 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:15940712 Approved no  
  Call Number Serial 2148  
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Author Khalil, A.M.; Kaseda, Y. url  openurl
  Title Behavioral patterns and proximate reason of young male separation in Misaki feral horses Type Journal Article
  Year 1997 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 54 Issue 4 Pages 281-289  
  Keywords Misaki feral horse; Horse maternal-filial bond; Social behavior; Separation season; Separation reason  
  Abstract The present investigation was undertaken to study the proximate reasons why and the behavioral patterns of young male Misaki feral horses when they left their natal band or mothers. We observed a total of ten young males twice a month from January 1988 to December 1995. Almost all young males left their natal band or mothers at between 1 and 4 years of age. We found that, during the separation process, all the young males from first parity dams returned several times after the initial separation, indicating a strong attachment between primiparous mares and their male offspring. The other five separated only once without rejoining. Our observations showed five variable behavior patterns of young males at separation time, depending on the consort relation between their mothers and harem stallion and the reason for separation at that time. Eight young males separated in the non-breeding season at average 2.1 years and the other two separated in the breeding season at average 3 years and the average difference was not significant. These results revealed that 80% of the young males separated voluntarily when the natural resources become poor whereas 20% separated when their siblings were born.  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 2209  
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Author Khalil, A.M.; Kaseda, Y. url  doi
openurl 
  Title Early experience affects developmental behaviour and timing of harem formation in Misaki horses Type Journal Article
  Year 1998 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 59 Issue 4 Pages 253-263  
  Keywords Feral horse; Young male; Social behavior; Developmental stage  
  Abstract A study was made of the behavior of young male Misaki feral horses in the developmental stage, by observing nine of them once a week from January 1988 to December 1996. The relationship between behavior before separation and in the developmental stage was also investigated. This stage begins just after young males separate from their natal band or mothers, and it continues until they start to form harems. The duration of the developmental stage in the study ranged from 0.6 to 3.9 years, depending on the age of the young males at the time of separation. Young males associated with three types of social groups at the beginning of the developmental stage, according to their social groups before separation. These were bachelor groups, harem groups and wandering female groups. During this period, males joined the three groups, mixed sex groups and sometimes were solitary. It was considered that these associations provided a good opportunity for males to acquire different behavioral patterns and experiences before they entered the next stage. Depending on the groups with which they associated, young males that spent more time with bachelor groups had the longest average developmental stage. They associated with harem groups more often during the breeding season and more frequently with other groups or were solitary during the non-breeding season. This may be a transition period because by the end of this stage all males had spent time in solitude before forming their own harem bands.  
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  Notes Approved no  
  Call Number Serial 2021  
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Author Khalil, A.M.; Murakami, N.; Kaseda, Y. openurl 
  Title Relationship between plasma testosterone concentrations and age, breeding season and harem size in Misaki feral horses Type Journal Article
  Year 1998 Publication The Journal of veterinary medical science / the Japanese Society of Veterinary Science Abbreviated Journal J Vet Med Sci  
  Volume 60 Issue 5 Pages 643-645  
  Keywords Age Factors; Animals; Animals, Wild; *Horses; Japan; Male; Reproduction; Seasons; Sexual Behavior, Animal; Sexual Maturation; *Social Behavior; Testosterone/*blood  
  Abstract Jugular vein blood samples were collected from 23 young and sexual mature feral stallions to examine the relationship between plasma testosterone concentration and age, breeding season or harem size. Testosterone concentration increased with the age of the stallions until they formed their own harems, at about 4 to 6 years old. Seasonal variations in testosterone concentrations were observed, and found to be significantly higher (P<0.001) throughout the breeding season than non-breeding season, from 3 years of age. Testosterone levels were correlated with harem size for individual stallions. It can be inferred from these results that there is a relationship between plasma testosterone concentration and age, breeding season and harem size.  
  Address Laboratory of Animal Behavioral Science, Faculty of Agriculture, Miyazaki University, Japan  
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  ISSN 0916-7250 ISBN Medium  
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  Notes PMID:9637303 Approved no  
  Call Number refbase @ user @ Serial 137  
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Author Kiley, M. openurl 
  Title The vocalizations of ungulates, their causation and function Type Journal Article
  Year 1972 Publication Zeitschrift für Tierpsychologie Abbreviated Journal Z. Tierpsychol.  
  Volume 31 Issue 2 Pages 171-222  
  Keywords Aggression; Animals; *Artiodactyla; Cattle; Fear; Female; Frustration; Horses; Humans; Male; Pain; *Perissodactyla; Sexual Behavior, Animal; Social Behavior; Social Dominance; Swine; *Vocalization, Animal  
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  Series Volume Series Issue Edition  
  ISSN 0044-3573 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:4674022 Approved no  
  Call Number refbase @ user @ Serial 681  
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Author Klingel, H. openurl 
  Title Social organization and reproduction in equids Type Journal Article
  Year 1975 Publication Journal of Reproduction and Fertility. Supplement Abbreviated Journal J Reprod Fertil Suppl  
  Volume Issue 23 Pages 7-11  
  Keywords Animals; Behavior, Animal; Female; Male; Perissodactyla/*physiology; Reproduction; *Sexual Behavior, Animal; Social Behavior; Territoriality  
  Abstract There are two distinct types of social organization and, accordingly, two types of mating systems in equids. In the horse, Plains zebra and Mountain zebra, the adults live in non-territorial and cohesive one-male groups and in stallion groups. The family stallions have exclusive mating rights which are respected by all others. In Grevy's zebra and in the African and Asiatic wild asses, the stallions are permanently territorial and have exclusive mating rights within their territories. Ecological and evolutionary aspects are discussed.  
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  Series Volume Series Issue Edition  
  ISSN 0449-3087 ISBN Medium  
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  Notes PMID:1060868 Approved no  
  Call Number Equine Behaviour @ team @ Serial 2303  
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Author Klingel, H. openurl 
  Title Social organization of feral horses Type Journal Article
  Year 1982 Publication Journal of Reproduction and Fertility. Supplement Abbreviated Journal J Reprod Fertil Suppl  
  Volume 32 Issue Pages 89-95  
  Keywords Animals; Animals, Wild; Female; Horses/*physiology; Male; Sexual Behavior, Animal; *Social Behavior; Territoriality  
  Abstract The basic social unit in feral horses is the family group consisting of one stallion, one to a few unrelated mares and their foals. Surplus stallions associate in bachelor groups. Stallions are instrumental in bringing mares together in a unit which then persists even without a stallion. The similarity of social organization in populations living in a variety of different habitats indicates that feral horses have reverted to the habits of their wild ancestors, and that domestication has had no influence on this basic behavioural feature.  
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  ISSN 0449-3087 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:6962906 Approved no  
  Call Number refbase @ user @ Serial 1958  
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Author Koski, S.E.; Koops, K.; Sterck, E.H.M. doi  openurl
  Title Reconciliation, relationship quality, and postconflict anxiety: testing the integrated hypothesis in captive chimpanzees Type Journal Article
  Year 2007 Publication American journal of primatology Abbreviated Journal Am. J. Primatol.  
  Volume 69 Issue 2 Pages 158-172  
  Keywords Animals; *Anxiety; *Behavior, Animal; Conflict (Psychology); Female; Male; Pan troglodytes/*psychology; *Social Behavior  
  Abstract Reconciliation is a conflict resolution mechanism that is common to many gregarious species with individualized societies. Reconciliation repairs the damaged relationship between the opponents and decreases postconflict (PC) anxiety. The “integrated hypothesis” links the quality of the opponents' relationship to PC anxiety, since it proposes that conflicts among partners with high relationship quality will yield high levels of PC anxiety, which in turn will lead to an increased likelihood of reconciliation. We tested the integrated hypothesis in captive chimpanzees (Pan troglodytes) in the Arnhem Zoo, The Netherlands. We applied the standard PC/matched control (MC) method. Our results mostly support the integrated hypothesis, in that more valuable and compatible partners (i.e., males and frequent groomers) reconciled more often than less valuable and weakly compatible partners (i.e., females and infrequent groomers). In addition, PC anxiety was higher after conflicts among males than among females. Emotional arousal thus appears to be a mediator facilitating reconciliation. However, in contrast to the predictions derived from the integrated hypothesis, PC anxiety appeared only in aggressees, and not in aggressors, of conflicts. This suggests that while relationship quality determines PC anxiety, it is dependent on the role of the participants in the conflict.  
  Address Department of Behavioural Biology, Utrecht University, The Netherlands. s.e.koski@bio.uu.nl  
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  Series Volume Series Issue Edition  
  ISSN 0275-2565 ISBN Medium  
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  Notes PMID:17146788 Approved no  
  Call Number refbase @ user @ Serial 2872  
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Author Kralj-Fiser, S.; Scheiber, I.B.R.; Blejec, A.; Moestl, E.; Kotrschal, K. doi  openurl
  Title Individualities in a flock of free-roaming greylag geese: behavioral and physiological consistency over time and across situations Type Journal Article
  Year 2007 Publication Hormones and Behavior Abbreviated Journal Horm Behav  
  Volume 51 Issue 2 Pages 239-248  
  Keywords Animals; Behavior, Animal/*physiology; Corticosterone/metabolism; Feces; Female; Geese/*physiology; Individuality; Male; Personality/*physiology; Population Density; Reproducibility of Results; Seasons; *Social Behavior; Social Environment; Testosterone/*metabolism  
  Abstract The concept of personality implies individual differences in behavior and physiology that show some degree of repeatability/consistency over time and across contexts. Most studies of animal personality, particularly studies of individuals' variation in physiological mechanisms, have been conducted on selected individuals in controlled conditions. We attempted to detect consistent behaviors as well as physiological patterns in greylag ganders (Anser anser) from a free-roaming flock living in semi-natural conditions. We tested 10 individuals repeatedly, in a handling trial, resembling tests for characterization of “temperaments” in captive animals. We recorded the behavior of the same 10 individuals during four situations in the socially intact flock: (1) a “low density feeding condition”, (2) a “high density feeding condition”, (3) a “low density post-feeding situation” and (4) while the geese rested. We collected fecal samples for determination of excreted immuno-reactive corticosterone (BM) and testosterone metabolites (TM) after handling trials, as well as the “low density feeding” and the “high density feeding” conditions. BM levels were very highly consistent over the repeats of handling trials, and the “low density feeding condition” and tended to be consistent over the first two repeats of the “high density feeding condition”. Also, BM responses tended to be consistent across contexts. Despite seasonal variation, there tended to be inter-test consistency of TM, which pointed to some individual differences in TM as well. Aggressiveness turned out to be a highly repeatable trait, which was consistent across social situations, and tended to correlate with an individual's resistance during handling trials. Also, “proximity to the female partner” and “sociability” – the average number of neighboring geese in a close distance while resting – were consistent. We conclude that aggressiveness, “affiliative tendencies” and levels of excreted corticosterone and testosterone metabolites may be crucial factors of personality in geese.  
  Address University of Ljubljana, Biotechnical faculty, Slovenia. simona.kralj@guest.arnes.si  
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  ISSN 0018-506X ISBN Medium  
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  Notes PMID:17196200 Approved no  
  Call Number Equine Behaviour @ team @ Serial 4189  
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