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	Author	Beran, M.J.; Beran, M.M.; Harris, E.H.; Washburn, D.A.
	Title	Ordinal judgments and summation of nonvisible sets of food items by two chimpanzees and a rhesus macaque			Type	Journal Article
	Year	2005	Publication	Journal of Experimental Psychology. Animal Behavior Processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	31	Issue	3	Pages	351-362
	Keywords	Animals; Behavior, Animal; Chi-Square Distribution; Cognition; Color Perception/physiology; Female; Food; Judgment/physiology; Macaca mulatta; Male; Pan troglodytes; Serial Learning/*physiology; Size Perception
	Abstract	Two chimpanzees and a rhesus macaque rapidly learned the ordinal relations between 5 colors of containers (plastic eggs) when all containers of a given color contained a specific number of identical food items. All 3 animals also performed at high levels when comparing sets of containers with sets of visible food items. This indicates that the animals learned the approximate quantity of food items in containers of a given color. However, all animals failed in a summation task, in which a single container was compared with a set of 2 containers of a lesser individual quantity but a greater combined quantity. This difficulty was not overcome by sequential presentation of containers into opaque receptacles, but performance improved if the quantitative difference between sizes was very large.
	Address	Language Research Center, Georgia State University, Decatur, 30034, USA. mjberan@yahoo.com
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16045389			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2766
Permanent link to this record



	Author	Imura, T.; Tomonaga, M.
	Title	Perception of depth from shading in infant chimpanzees ( Pan troglodytes)			Type	Journal Article
	Year	2003	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	6	Issue	4	Pages	253-258
	Keywords	Age Factors; Animals; Contrast Sensitivity/physiology; Depth Perception/physiology; Discrimination Learning/physiology; Female; Male; Pan troglodytes/growth & development/physiology/*psychology; Pattern Recognition, Visual/physiology
	Abstract	We investigated the ability to perceive depth from shading, one of the pictorial depth cues, in three chimpanzee infants aged 4-10 months old, using a preferential reaching task commonly used to study pictorial depth perception in human infants. The chimpanzee infants reached significantly more to three-dimensional toys than to pictures thereof and more to the three-dimensional convex than to the concave. Furthermore, two of the three infants reached significantly more to the photographic convex than to the photographic concave. These infants also looked longer at the photographic convex than the concave. Our results suggest that chimpanzees perceive, at least as early as the latter half of the first year of life, pictorial depth defined by shading information. Photographic convexes contain richer information about pictorial depth (e.g., attached shadow, cast shadow, highlighted area, and global difference in brightness) than simple computer-graphic graded patterns. These cues together might facilitate the infants' perception of depth from shading.
	Address	Graduate School of Humanities, Kwansei Gakuin University, Uegahara, Nishinomiya, Hyogo 662-8501, Japan
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:14610661			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2550
Permanent link to this record



	Author	Parr, L.A.
	Title	Perceptual biases for multimodal cues in chimpanzee (Pan troglodytes) affect recognition			Type	Journal Article
	Year	2004	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	7	Issue	3	Pages	171-178
	Keywords	Acoustic Stimulation; Animal Communication; Animals; Auditory Perception/physiology; Cues; Discrimination Learning/physiology; Facial Expression; Female; Male; Pan troglodytes/psychology; Perceptual Masking/physiology; Photic Stimulation; Recognition (Psychology)/physiology; Visual Perception/physiology; Vocalization, Animal
	Abstract	The ability of organisms to discriminate social signals, such as affective displays, using different sensory modalities is important for social communication. However, a major problem for understanding the evolution and integration of multimodal signals is determining how humans and animals attend to different sensory modalities, and these different modalities contribute to the perception and categorization of social signals. Using a matching-to-sample procedure, chimpanzees discriminated videos of conspecifics' facial expressions that contained only auditory or only visual cues by selecting one of two facial expression photographs that matched the expression category represented by the sample. Other videos were edited to contain incongruent sensory cues, i.e., visual features of one expression but auditory features of another. In these cases, subjects were free to select the expression that matched either the auditory or visual modality, whichever was more salient for that expression type. Results showed that chimpanzees were able to discriminate facial expressions using only auditory or visual cues, and when these modalities were mixed. However, in these latter trials, depending on the expression category, clear preferences for either the visual or auditory modality emerged. Pant-hoots and play faces were discriminated preferentially using the auditory modality, while screams were discriminated preferentially using the visual modality. Therefore, depending on the type of expressive display, the auditory and visual modalities were differentially salient in ways that appear consistent with the ethological importance of that display's social function.
	Address	Division of Psychobiology, Yerkes National Primate Research Center, Emory University, 954 Gatewood Road, GA 30329, Atlanta, USA. parr@rmy.emory.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:14997361			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2544
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	Author	Suda, C.; Call, J.
	Title	Piagetian conservation of discrete quantities in bonobos (Pan paniscus), chimpanzees (Pan troglodytes), and orangutans (Pongo pygmaeus)			Type	Journal Article
	Year	2005	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	8	Issue	4	Pages	220-235
	Keywords	Animals; Behavior, Animal; Female; Hominidae/psychology; Male; Pan paniscus; Pan troglodytes; Pongo pygmaeus; *Problem Solving
	Abstract	This study investigated whether physical discreteness helps apes to understand the concept of Piagetian conservation (i.e. the invariance of quantities). Subjects were four bonobos, three chimpanzees, and five orangutans. Apes were tested on their ability to conserve discrete/continuous quantities in an over-conservation procedure in which two unequal quantities of edible rewards underwent various transformations in front of subjects. Subjects were examined to determine whether they could track the larger quantity of reward after the transformation. Comparison between the two types of conservation revealed that tests with bonobos supported the discreteness hypothesis. Bonobos, but neither chimpanzees nor orangutans, performed significantly better with discrete quantities than with continuous ones. The results suggest that at least bonobos could benefit from the discreteness of stimuli in their acquisition of conservation skills.
	Address	Max Planck Institute for Evolutionary Anthropology, Leipzig, Germany. suda@eva.mpg.de
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15692813			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2494
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	Author	Flack, J.C.; Jeannotte, L.A.; de Waal, F.B.M.
	Title	Play signaling and the perception of social rules by juvenile chimpanzees (Pan troglodytes)			Type	Journal Article
	Year	2004	Publication	Journal of comparative psychology (Washington, D.C. : 1983)	Abbreviated Journal	J Comp Psychol
	Volume	118	Issue	2	Pages	149-159
	Keywords	Age Factors; Animals; Behavior, Animal; Female; Male; Pan troglodytes; Play and Playthings; Recognition (Psychology); Signal Detection (Psychology); *Social Perception
	Abstract	Prescriptive social rules are enforced statistical regularities. The authors investigated whether juvenile chimpanzees (Pan troglodytes) recognize and use enforced statistical regularities to guide dyadic play behavior. They hypothesized (a) that proximity of adults, especially mothers of younger play partners, to play bouts will increase the play signaling of older partners and (b) that when juvenile-juvenile play bouts occur in proximity to adults, older partners will play at a lower intensity than when no adults are present. They found that older and younger partners increase their play signaling in the presence of the mothers of younger partners, particularly as the intensity of play bouts increases. In contrast to their hypothesis, older partners played more roughly when the mothers of younger partners were in proximity. These results suggest that juvenile chimpanzees increase play signaling to prevent termination of the play bouts by mothers of younger partners.
	Address	Santa Fe Institute, NM 97501, USA. jflack@santafe.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15250802			Approved	no
	Call Number	refbase @ user @			Serial	172
Permanent link to this record



	Author	Palagi, E.; Cordoni, G.; Borgognini Tarli, S.
	Title	Possible roles of consolation in captive chimpanzees (Pan troglodytes)			Type	Journal Article
	Year	2006	Publication	American Journal of Physical Anthropology	Abbreviated Journal	Am J Phys Anthropol
	Volume	129	Issue	1	Pages	105-111
	Keywords	Animals; Animals, Zoo/physiology; Empathy; Female; Male; Pan troglodytes/physiology; *Social Behavior; Stress/physiopathology/veterinary
	Abstract	Empathy is a necessary prerequisite for the occurrence of consolation. The term “consolation” contains a hypothesis about function, which is distress alleviation. The present study aims to confirm the occurrence of consolation in captive chimpanzees via the post-conflict/matched-control method (PC-MC) and to suggest its possible roles. We collected 273 PC-MC pairs in the group of Pan troglodytes housed in the ZooParc de Beauval (France). We confirmed the presence of consolatory contacts (mean level of consolation, 49.5% +/- 22.3% SEM) in the colony. Consolation rates were significantly higher than reconciliation levels (mean level of reconciliation, 28.9% +/- 16.8% SEM). The level of consolation was greater in the absence of reconciliation than in the presence of it, suggesting that consolation might be an alternative behavior. As friendship and relatedness did not influence the occurrence of consolation, they did not seem to be the best prerequisites for this behavioral mechanism, at least in this chimpanzee colony. Affinitive contacts with third parties were significantly more frequent when the victim called attention to itself during severe aggressions by screaming. These high-pitched sounds seem to be useful in eliciting aid from conspecifics, as occurs in young humans. The occurrence of consolation reduced the likelihood of further attacks among group-members. From this perspective, both victims and consolers most likely gain potential advantages by interacting with each other when aggression is particularly severe, reconciliation is not immediate, and consequently social stress reaches high levels.
	Address	Centro Interdipartimentale Museo di Storia Naturale e del Territorio, Universita di Pisa, 56010 Calci, Italy. betta.palagi@museo.unipi.it
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0002-9483	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16229027			Approved	no
	Call Number	refbase @ user @			Serial	2871
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	Author	de Waal, F.B.; Seres, M.
	Title	Propagation of handclasp grooming among captive chimpanzees			Type	Journal Article
	Year	1997	Publication	American journal of primatology	Abbreviated Journal	Am. J. Primatol.
	Volume	43	Issue	4	Pages	339-346
	Keywords	Animals; Family Relations; Female; Grooming; Learning; Male; Pan troglodytes/psychology; *Social Behavior
	Abstract	A grooming posture previously reported for two wild chimpanzee (Pan troglodytes) communities developed spontaneously in a captive group of the same species. This offered a unique opportunity to follow the propagation of a new social custom. The posture consists of two partners grasping hands--either both right hands or both left hands--and raising the arms in an A-frame above their heads while mutually grooming with their free hands. The propagation of this pattern was followed over a 5 year period. In the beginning, handclasps were always initiated by the same adult female. This female initiated the posture mainly with her adult female kin. In subsequent years, these relatives became frequent participants in the posture with each other as well as with nonrelatives. Over the years the posture increased in frequency and duration and spread to the majority of adults and also to a few adolescents and older juveniles. The pattern persisted after removal of the apparent originator.
	Address	Yerkes Primate Center, Emory University, Atlanta, GA 30329, USA. dewaal@emory.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0275-2565	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:9403098			Approved	no
	Call Number	refbase @ user @			Serial	202
Permanent link to this record



	Author	Boysen, S.T.; Bernston, G.G.; Hannan, M.B.; Cacioppo, J.T.
	Title	Quantity-based interference and symbolic representations in chimpanzees (Pan troglodytes)			Type	Journal Article
	Year	1996	Publication	Journal of Experimental Psychology. Animal Behavior Processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	22	Issue	1	Pages	76-86
	Keywords	Animals; Behavior, Animal; Cognition; Discrimination Learning; Female; Male; Pan troglodytes; *Reinforcement (Psychology); Task Performance and Analysis
	Abstract	Five chimpanzees with training in counting and numerical skills selected between 2 arrays of different amounts of candy or 2 Arabic numerals. A reversed reinforcement contingency was in effect, in which the selected array was removed and the subject received the nonselected candies (or the number of candies represented by the nonselected Arabic numeral). Animals were unable to maximize reward by selecting the smaller array when candies were used as array elements. When Arabic numerals were substituted for the candy arrays, all animals showed an immediate shift to a more optimal response strategy of selecting the smaller numeral, thereby receiving the larger reward. Results suggest that a response disposition to the high-incentive candy stimuli introduced a powerful interference effect on performance, which was effectively overridden by the use of symbolic representations.
	Address	Ohio State University, Department of Psychology, Ohio State University, Columbus 43210-1222, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:8568498			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2781
Permanent link to this record



	Author	Tanaka, M.
	Title	Recognition of pictorial representations by chimpanzees (Pan troglodytes)			Type	Journal Article
	Year	2007	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	10	Issue	2	Pages	169-179
	Keywords	Animals; Discrimination Learning; Female; Male; Pan troglodytes/physiology; Pattern Recognition, Visual; Photography
	Abstract	In this study, I investigated chimpanzees' ability to recognize pictorial representations. Four adults and three juvenile chimpanzees were trained to choose images of photographs of flowers among 12 items belonging to four categories on a touch-sensitive monitor. As a generalization test, the following five types of images were presented: (1) novel photographs, (2) colored sketches (more realistic), (3) a colored clip art (cartoon-like images), (4) black-and-white line drawings, and (5) Kanji characters (as the control images). One adult and all three juvenile chimpanzees were able to choose any style of the nonphotographic images of flowers significantly above the chance level, whereas none could choose the correct Kanji characters corresponding to a flower significantly above the chance level. The other three adult chimpanzees' performance level did not exceed the chance level in terms of choosing nonphotographic images although they showed good transfer skills to novel photographs. The results revealed that not all chimpanzees could recognize pictures used by humans without training. The results also suggest “critical period” in acquisition of skill in recognizing pictures in chimpanzees. Only one adult chimpanzee, who had acquired skill in recognizing visual symbols, also recognized pictures aside from the juvenile chimpanzees. Her learning history might have aided her in acquiring this skill. The results of this study suggest a relationship between pictorial competence and symbolic one.
	Address	Department of Brain and Behavioral Sciences, Primate Research Institute, Kyoto University, 41-2 Kanrin, Inuyama, Aichi, 484-8506, Japan. mtanaka@pri.kyoto-u.ac.jp
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17171361			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2428
Permanent link to this record



	Author	Parr, L.A.; Winslow, J.T.; Hopkins, W.D.; de Waal, F.B.
	Title	Recognizing facial cues: individual discrimination by chimpanzees (Pan troglodytes) and rhesus monkeys (Macaca mulatta)			Type	Journal Article
	Year	2000	Publication	Journal of comparative psychology (Washington, D.C. : 1983)	Abbreviated Journal	J Comp Psychol
	Volume	114	Issue	1	Pages	47-60
	Keywords	Animals; Discrimination Learning; Facial Expression; Female; Macaca mulatta/psychology; Male; Mental Recall; Pan troglodytes/psychology; Perceptual Masking; *Social Perception; Species Specificity
	Abstract	Faces are one of the most salient classes of stimuli involved in social communication. Three experiments compared face-recognition abilities in chimpanzees (Pan troglodytes) and rhesus monkeys (Macaca mulatta). In the face-matching task, the chimpanzees matched identical photographs of conspecifics' faces on Trial 1, and the rhesus monkeys did the same after 4 generalization trials. In the individual-recognition task, the chimpanzees matched 2 different photographs of the same individual after 2 trials, and the rhesus monkeys generalized in fewer than 6 trials. The feature-masking task showed that the eyes were the most important cue for individual recognition. Thus, chimpanzees and rhesus monkeys are able to use facial cues to discriminate unfamiliar conspecifics. Although the rhesus monkeys required many trials to learn the tasks, this is not evidence that faces are not as important social stimuli for them as for the chimpanzees.
	Address	Department of Psychology, Emory University. parr@rmy.emory.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:10739311			Approved	no
	Call Number	refbase @ user @			Serial	191
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