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Author VanDierendonck, M.C.; de Vries, H.; Schilder, M.B.H.
Title An Analysis of Dominance, Its Behavioural Parameters and Possible Determinants in a Herd of Icelandic orses in Captivity Type Journal Article
Year 1995 Publication Netherlands Journal of Zoology Abbreviated Journal Netherl. J. Zool.
Volume 45 Issue 3-4 Pages 362-385
Keywords Dominance; rank order; horses; Icelandic horses.
Abstract Th e applicability of the concept of dominance was investigated in a captive herd of  Icelandic

horses and  ponies of diff erent breeds. Eight out of  behaviours possibly related

to dominance occurred frequently enough to be investigated in detail. For these eight agonistic

behaviours the coverage, the unidirectionality in the exchange, and the degree of

transitivity (Landau`s linearity index) were calculated. Four off ensive behaviours, together

with avoidance, were suitable for further analysis with regard to dominance. Th e patterns

of asymmetries with which these behaviours were exchanged were suffi ciently similar as to

justify the application of the dominance concept and to construct a (nearly) linear dominance

hierarchy. Th e rank order of the castrated stallions was completely linear, the hierarchy

of the mares was almost completely linear. Th e results suggest that off ensive and defensive

aggressive behaviours should be treated separately and that the concept of dominance

is applicable. However, ritualized formal dominance signals between adult horses appear to

be (almost) absent. Th e rank positions of the individuals were correlated with age and residency

in the herd but not with height. Middle ranking horses tended to be more frequently

in the close vicinity of another horse than high ranking or low ranking horses. Over and

above this correlation at the individual level, it was found that pairs of horses close in rank

to each other were more often also spatially close to each other. Being in oestrus did not infl

uence the dominance relationships between mares. For castrated stallions the rank positions

were correlated with the age at which they were castrated. Th is suggests that in male

horses experience prior to neutering infl uences the behaviour afterwards.
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Call Number refbase @ user @ Serial 440
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Author R. A. Hopkins
Title CALIFORNIA WILDLIFE HABITAT RELATIONSHIPS SYSTEM Type Manuscript
Year Publication Abbreviated Journal
Volume M174 Issue Pages
Keywords Feral Horse Equus caballus
Abstract
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Call Number refbase @ user @ Serial 476
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Author Kumar, P.; Timoney, J.F.; Southgate, H.H.; Sheoran, A.S.
Title Light and scanning electron microscopic studies of the nasal turbinates of the horse Type Journal Article
Year 2000 Publication Anatomia, histologia, embryologia Abbreviated Journal Anat Histol Embryol
Volume 29 Issue 2 Pages 103-109
Keywords Animals; Female; Horses/*anatomy & histology; Male; Microscopy, Electron, Scanning; Turbinates/*cytology/ultrastructure
Abstract The nasal turbinates of 5 young horses were studied by light and scanning electron-microscopy. Stratified cuboidal epithelium lined the rostral part of the dorsal and ventral nasal turbinates of the vestibular region. The polyangular microvillus cells of this region were separated by linear depressions. The mid and caudal parts of the dorsal and ventral nasal turbinates and the rostral part of the ethmoturbinates were lined by pseudostratified columnar ciliated respiratory epithelium. Numerous cilia with dilated blebs on the ciliated cells concealed adjacent non-ciliated supporting cells and goblet cells. The olfactory zone consisting of the olfactory vesicle and a dense network of olfactory cilia localized to the caudal part of the ethmoturbinates. The three regions were delineated from each other by transitional zones.
Address Gluck Equine Research Center, University of Kentucky, Lexington 40546-0099, USA
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Language English Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0340-2096 ISBN Medium
Area Expedition Conference
Notes PMID:10932387 Approved no
Call Number refbase @ user @ Serial 505
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Author Seaman, S.C.; Davidson, H.P.B.; Waran, N.K.
Title How reliable is temperament assessment in the domestic horse (Equus caballus)? Type Journal Article
Year 2002 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 78 Issue 2-4 Pages 175-191
Keywords Temperament assessment; Behavioural tests; Horses; Active and passive copers; Factor analysis
Abstract Differences in behavioural characteristics between individuals of the same species are often described as being due to the temperament of the individuals. These differences can have enormous implications for welfare with some individuals apparently being able to adapt to environmental challenge more easily than others. Such differences have resulted in animals often being described as either `active' copers, which try to escape from or remove an aversive stimulus, or `passive' copers, which show no outward signs of a situation being aversive, thus, appearing to be unaffected. Tests previously developed to assess the temperament of animals have been criticised for several reasons. Behaviour is often recorded and categorised using methods that are not objective and tests are generally carried out once with no consideration of whether or not behavioural responses are consistent over time. This study takes these factors into account. The behaviour of 33 horses was recorded in three types of test--an arena test, response to a person and response to an object. In order to test whether or not responses were consistent over time, the tests were repeated three times with an average of 9 days between trials. Test results were validated using responses from questionnaires completed by the farm team leader. The data were analysed using an initial principal component analysis (PCA) and factor analysis. The horses were found to behave consistently over the three trials in their responses in the arena test. The responses to the person test and the object test were similar to each other; however, these responses were not consistent over trials. The behaviour in the arena test was unable to be used to make a prediction of behaviour in the person and object tests and vice versa. The responses shown by the horses did not enable them to be categorised as either active or passive copers. Behavioural responses in the tests were not predictive of the response to a startle test (water spray), nor could they be used to predict status or response to being reintroduced to the group after testing. There was no relationship between the responses in the tests and the ratings given by the farm team leader. It was concluded that horses vary widely in their responses to artificial behavioural tests, with only the responses to an open-field arena test being consistent over time, and therefore, the only type of test which can indicate some core factor of temperament.
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Call Number refbase @ user @ Serial 520
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Author Murphy, J.; Sutherland, A.; Arkins, S.
Title Idiosyncratic motor laterality in the horse Type Journal Article
Year 2005 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 91 Issue 3-4 Pages 297-310
Keywords Horse; Idiosyncratic motor behaviour; Laterality; Sidedness
Abstract Idiosyncratic motor behaviour was investigated during four experimental procedures in 40 horses (males = 20, females = 20) to establish if horses revealed evidence of significant right or left laterality. The experimental procedures included (1) detection of the preferred foreleg to initiate movement (walk or trot), (2) obstacle avoidance within a passageway (right or left), (3) obstacle avoidance when ridden and (4) idiosyncratic motor bias when rolling. The influence of the horses' sex on both the direction and the degree of the laterality was explored within and between experimental procedures. The findings showed that the direction, but not the degree of idiosyncratic motor preference in the horses was strongly sex-related. Male horses exhibited significantly more (t = 3.74, d.f. = 79, P < 0.001) left lateralised responses and female horses exhibited significantly more (t = -6.35, d.f. = 79, P < 0.01) right lateralised responses. There was also significant positive correlation (P < 0.05) between four of six possible inter-experimental relationships. The results suggest two discrete trends of laterality associated with the sex of the horse. The primary cause of idiosyncratic motor laterality may be genetically predetermined, influenced by environmental factors or a combination of these two and the current findings may support the development of sex-specific training schedules for the horse. Further, work in this area might assist in defining the mechanisms of brain hemisphere lateralisation and allocation of cognitive function in the horse.
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Notes Approved no
Call Number refbase @ user @ Serial 527
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Author Deuel, N.R.; Lawrence, L.M.
Title Laterality in the gallop gait of horses Type Journal Article
Year 1987 Publication Journal of biomechanics Abbreviated Journal J Biomech
Volume 20 Issue 6 Pages 645-649
Keywords Animals; *Functional Laterality; *Gait; Horses/*physiology; Kinesis
Abstract Bilateral asymmetry in gallop stride limb contact patterns of four Quarter Horse fillies was documented by high-speed cinematography. Horses were filmed with rider by two cameras simultaneously while galloping along a straightaway. Even though signaled for each gallop lead an equivalent number of times, horses frequently switched leads, selecting the left lead nearly twice as often as the right. Velocities and stride lengths were greater for the left lead than the right, but stride frequencies did not differ between leads. Velocity effects were partitioned out in limb contact data analysis to enable the determination of persistent gallop stride asymmetries. The contact duration for the trailing (right) fore limb on the left lead exceeded the contact duration for the trailing (left) fore limb on the right lead. Selecting the right fore limb as the trailing fore limb may have allowed horses to use it to withstand the greater stresses and caused them to preferentially gallop with the left fore limb leading. Laterality may have an important influence on equine gallop motion patterns and thereby influence athletic performance.
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ISSN 0021-9290 ISBN Medium
Area Expedition Conference
Notes PMID:3611140 Approved no
Call Number refbase @ user @ Serial 528
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Author Proudman, C.; Pinchbeck, G.; Clegg, P.; French, N.
Title Equine welfare: risk of horses falling in the Grand National Type Journal Article
Year 2004 Publication Nature Abbreviated Journal Nature
Volume 428 Issue 6981 Pages 385-386
Keywords Accidental Falls/prevention & control/*statistics & numerical data; Animal Welfare; Animals; Great Britain; Horse Diseases/prevention & control; Horses/*physiology; Odds Ratio; Risk Assessment; *Sports
Abstract As in other competitive sports, the famous Grand National steeplechase, which is held at Aintree in the United Kingdom and is watched by 600 million people worldwide, sometimes results in injury. By analysing data from the past 15 Grand National races (consisting of 560 starts by horses), we are able to identify several factors that are significantly associated with failure to complete the race: no previous experience of the course and its unique obstacles, unfavourable ground conditions (too soft or too hard), a large number of runners, and the length of the odds ('starting price'). We also find that there is an increased risk of falling at the first fence and at the jump known as Becher's Brook, which has a ditch on the landing side. Our findings indicate ways in which the Grand National could be made safer for horses and illustrate how epidemiological analysis might contribute to preventing injury in competitive sport.
Address Epidemiology Group, Faculty of Veterinary Science, University of Liverpool, Leahurst, Neston, Wirral CH64 7TE, UK. c.j.proudman@liverpool.ac.uk
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ISSN 1476-4687 ISBN Medium
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Notes PMID:15042079 Approved no
Call Number refbase @ user @ Serial 535
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Author Lindberg, A.C.; Kelland, A.; Nicol, C.J.
Title Effects of observational learning on acquisition of an operant response in horses Type Journal Article
Year 1999 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 61 Issue 3 Pages 187-199
Keywords Horse; Observational learning; Stereotyped behaviour; Operant behaviour; Breed influence; Age influence
Abstract The effect of observational learning on the acquisition of an operant response was examined in eighteen riding horses and ponies. The test horses were randomly divided into three groups of six and individually exposed to one of three treatments. An additional horse was trained as a demonstrator, to perform the operant response. The observer horses watched either the demonstrator performing the bin-opening response (Group D+B); the demonstrator standing passively (Group D); or the operant bin in the absence of the demonstrator (Group B). Observers had access to and were free to interact with an identical bin during testing. Observers in Groups D+B and D were socially familiar with the demonstrator. Each test horse was tested once a day for 10 days. An ANOVA revealed no significant differences between treatment groups in the number of responses or the time taken to reach the learning criterion. However, there were highly significant differences between breed types, with non-warmbloods performing more bouts of opening the bin and feeding (p=0.02), feeding from the bin sooner (p=0.01) and reaching the criterion for learning sooner than warmbloods (p=0.05). There was also a significant negative linear relationship between horses' ages and time spent investigating the bin, with younger horses performing more investigative behaviour (y=-3.08x+106.86; p=0.02).
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Call Number refbase @ user @ Serial 562
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Author Clarke, J.V.; Nicol, C.J.; Jones, R.; McGreevy, P.D.
Title Effects of observational learning on food selection in horses Type Journal Article
Year 1996 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 50 Issue 2 Pages 177-184
Keywords Horse; Observational learning; Food discrimination
Abstract Fourteen riding horses of mixed age and breed were randomly allocated to observer and control treatments. An additional horse was pre-trained as a demonstrator to walk the 13.8 m length of the test arena and select one of two food buckets using colour and pattern cues. Observer horses were exposed to correct performances of the task by the trained demonstrator, for 20 trials held over 2 days. Control horses were subjected to the same handling and placement procedures as the observer horses but without exposure to the behaviour of the demonstrator. The third day for all subjects was designated as a test day. Each subject was released individually in a predetermined place in the arena, and the latency to walk the length of the test arena to the food buckets, the latency to feed, the identity of the bucket approached and the identity of the bucket selected were recorded on ten consecutive trials. During tests both food buckets contained food to minimize the possibility of individual trial and error learning. On the first trial the mean latency to approach the goal area was 18 s for observer horses, compared with 119 s for control horses (t = 2.8, d.f. = 12, P < 0.01) and the mean latency to eat was 35 s for observer horses, compared with 181 s for control horses (t = 4.86, d.f. = 11, P < 0.001). However, observer horses were no more likely to choose the demonstrated bucket than control horses on the first trial. Twelve of the 14 horses decreased their latency to approach the goal area during the series of ten trials, but there were no significant changes in the buckets selected.
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Call Number refbase @ user @ Serial 563
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Author Weeks, J.W.; Crowell-Davis, S.L.; Heusner, G.
Title Preliminary study of the development of the Flehmen response in Equus caballus Type Journal Article
Year 2002 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 78 Issue 2-4 Pages 329-335
Keywords Horse behavior; Flehmen; Foal development
Abstract The flehmen response is commonly seen in most ungulates as well as in several other species (e.g. felids). The behavior is most often thought to be part of the sexual behavioral repertoire of males. One reigning hypothesis suggests that this behavior allows the male to determine the estrous state of a female through the chemosensory functions of the vomeronasal organ. However, females and young of both sexes also exhibit this behavior. Horse foals most frequently show the flehmen response during their first month of life with colts showing the behavior more often than fillies. This study tested the flehmen response on male and female foals throughout their pre-pubertal period. Foals were separately presented estrous and non-estrous urine weekly during the first month of life and then monthly until they were approximately 7 months of age. No significant differences were found between male and female foals for the following variables: latency to flehmen, duration of flehmen, frequency of flehmen and sniffs.
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Notes Approved no
Call Number refbase @ user @ Serial 615
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