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	Author	Heitor, F.; do Mar Oom, M.; Vicente, L.
	Title	Social relationships in a herd of Sorraia horses Part I. Correlates of social dominance and contexts of aggression			Type	Journal Article
	Year	2006	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	73	Issue	2	Pages	170-177
	Keywords	Age Factors; Aggression; Animals; Female; Hierarchy, Social; Horses/psychology; Male; Sex Factors; Social Dominance; *Social Environment; Statistics, Nonparametric
	Abstract	Factors related to dominance rank and the functions of aggression were studied in a herd of Sorraia horses, Equus caballus, under extensive management. Subjects were 10 adult mares 5-18 years old and a stallion introduced into the group for breeding. Dominance relationships among mares were clear, irrespective of rank difference, and remained stable after introduction of the stallion. The dominance hierarchy was significantly linear and rank was positively correlated with age and total aggressiveness. Higher-ranking mares received lower frequency and intensity of agonistic interactions. Nevertheless, higher-ranking dominants were not more likely to elicit submission from their subordinates than lower-ranking dominants. Neither close-ranking mares nor mares with less clear dominance relationships were more aggressive towards each other. Agonistic interactions seemed to be used more importantly in regulation of space than to obtain access to food or to reassert dominance relationships. Contexts of aggression were related to mare rank. The results suggest that dominance relationships based on age as a conventional criterion were established to reduce aggressiveness in a herd where the costs of aggression are likely to outweigh the benefits.
	Address	Centro de Biologia Ambiental, Faculdade de Ciencias da Universidade de Lisboa, Campo Grande, Edificio C2, 1749-016 Lisboa, Portugal
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16815645			Approved	no
	Call Number	refbase @ user @			Serial	292
Permanent link to this record



	Author	Hartmann, E.; Christensen, J.W.; McGreevy, P.D.
	Title	Dominance and Leadership: Useful Concepts in Human-Horse Interactions?			Type	Journal Article
	Year	2017	Publication	Journal of Equine Veterinary Science	Abbreviated Journal	Proceedings of the 2017 Equine Science Symposium
	Volume	52	Issue		Pages	1-9
	Keywords	Horse; Social order; Dominance hierarchy; Aggression; Injury; Learning; Training
	Abstract	Dominance hierarchies in horses primarily influence priority access to limited resources of any kind, resulting in predictable contest outcomes that potentially minimize aggressive encounters and associated risk of injury. Levels of aggression in group-kept horses under domestic conditions have been reported to be higher than in their feral counterparts but can often be attributed to suboptimal management. Horse owners often express concerns about the risk of injuries occurring in group-kept horses, but these concerns have not been substantiated by empirical investigations. What has not yet been sufficiently addressed are human safety aspects related to approaching and handling group-kept horses. Given horse's natural tendency to synchronize activity to promote group cohesion, questions remain about how group dynamics influence human-horse interactions. Group dynamics influence a variety of management scenarios, ranging from taking a horse out of its social group to the prospect of humans mimicking the horse's social system by taking a putative leadership role and seeking after an alpha position in the dominance hierarchy to achieve compliance. Yet, there is considerable debate about whether the roles horses attain in their social group are of any relevance in their reactions to humans. This article reviews the empirical data on social dynamics in horses, focusing on dominance and leadership theories and the merits of incorporating those concepts into the human-horse context. This will provide a constructive framework for informed debate and valuable guidance for owners managing group-kept horses and for optimizing human-horse interactions.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0737-0806	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6712
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	Author	Grosenick, L.; Clement, T.S.; Fernald, R.D.
	Title	Fish can infer social rank by observation alone			Type	Journal Article
	Year	2007	Publication	Nature	Abbreviated Journal	Nature
	Volume	445	Issue	7126	Pages	429-432
	Keywords	Aggression/physiology; Animals; Cognition/physiology; Female; Fishes/physiology; Learning/physiology; Male; Models, Biological; Social Dominance; Territoriality
	Abstract	Transitive inference (TI) involves using known relationships to deduce unknown ones (for example, using A > B and B > C to infer A > C), and is thus essential to logical reasoning. First described as a developmental milestone in children, TI has since been reported in nonhuman primates, rats and birds. Still, how animals acquire and represent transitive relationships and why such abilities might have evolved remain open problems. Here we show that male fish (Astatotilapia burtoni) can successfully make inferences on a hierarchy implied by pairwise fights between rival males. These fish learned the implied hierarchy vicariously (as 'bystanders'), by watching fights between rivals arranged around them in separate tank units. Our findings show that fish use TI when trained on socially relevant stimuli, and that they can make such inferences by using indirect information alone. Further, these bystanders seem to have both spatial and featural representations related to rival abilities, which they can use to make correct inferences depending on what kind of information is available to them. Beyond extending TI to fish and experimentally demonstrating indirect TI learning in animals, these results indicate that a universal mechanism underlying TI is unlikely. Rather, animals probably use multiple domain-specific representations adapted to different social and ecological pressures that they encounter during the course of their natural lives.
	Address	Department of Biological Sciences, Stanford University, Stanford, California, 94305, USA. logang@stanford.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1476-4687	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17251980			Approved	no
	Call Number	refbase @ user @			Serial	600
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	Author	Goursot, C.; Düpjan, S.; Puppe, B.; Leliveld, L.M.C.
	Title	Affective styles and emotional lateralization: A promising framework for animal welfare research			Type	Journal Article
	Year	2021	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	237	Issue		Pages	105279
	Keywords	Individuality; Motor lateralization; Hemispheric dominance; Motivational tendencies; Emotional reactivity; Emotional regulation
	Abstract	The growing recognition of animals as individuals has broader implications for farm animal welfare research. Even under highly standardized on-farm conditions, farm animals show heterogeneous but individually consistent behavioural patterns towards various stimuli, based on how they appraise these stimuli. As a result, animal welfare is likely to be highly individual as well, and studying the proximate mechanisms underlying distinct individual behaviour patterns and appraisal will improve animal welfare research. We propose to extend the framework of affective styles to bridge the gap between existing research fields on animal personality and affective states. Affective styles refer to consistent individual differences in emotional reactivity and regulation and can be predicted by baseline cerebral lateralization. Likewise, animals with consistent left or right motor biases--a proxy measure of individual patterns in cerebral lateralization--have been shown to differ in their personality, emotional reactivity, motivational tendencies or coping styles. In this paper, we present the current knowledge of the links between laterality and stable individual traits in behaviour and affect in light of hypotheses on emotional lateralization. Within our suggested framework, we make recommendations on how to investigate affective styles in non-human animals and give practical examples. This approach has the potential to promote a science of affective styles in nonhuman animals and significantly advance research on animal welfare.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6698
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	Author	Giles, N.; Tupper, J.
	Title	Equine interspecies aggression			Type
	Year	2006	Publication	The Veterinary record	Abbreviated Journal	Vet. Rec.
	Volume	159	Issue	22	Pages	756
	Keywords	Aggression/physiology; Animals; Horses/physiology; Sheep/*physiology; Social Dominance
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0042-4900	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17127768			Approved	no
	Call Number				Serial	1779
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	Author	Gajdon, G.K.; Fijn, N.; Huber, L.
	Title	Limited spread of innovation in a wild parrot, the kea (Nestor notabilis)			Type	Journal Article
	Year	2006	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	9	Issue	3	Pages	173-181
	Keywords	Animal Communication; Animals; Diffusion of Innovation; Feeding Behavior; Female; Imitative Behavior; Learning; Male; Motor Skills; Parrots; *Problem Solving; Sex Factors; Social Dominance; Social Environment; Statistics, Nonparametric
	Abstract	In the local population of kea in Mount Cook Village, New Zealand, some keas open the lids of rubbish bins with their bill to obtain food scraps within. We investigated the extent to which this innovation has spread in the local population, and what factors limit the acquisition of bin opening. Only five males of 36 individually recognised birds were observed to have performed successful bin opening. With one exception there were always other keas present, watching successful bin opening. Seventeen additional individuals were seen to have benefitted from lid opening. Their foraging success was less than that of the bin openers. Social status of bin openers did not differ from scrounging males. Among the individuals that were regularly seen at the site of the bins but were not successful in bin opening, social status and the ratio of feeding directly from open bins correlated with the amount of opening attempts. We conclude that scrounging facilitated certain behavioural aspects of bin opening rather than inhibiting them. The fact that only 9% of opening attempts were successful, and the long period of time required to increase efficiency in lid opening shows that mainly individual experience, and to a lesser extent insight and social learning, play key roles in acquisition of the opening technique. The results indicate that the spread of innovative solutions of challenging mechanical problems in animals may be restricted to only a few individuals.
	Address	Department for Behavior, Neurobiology and Cognition, University of Vienna, 1090, Vienna, Austria. gyula.gajdon@univie.ac.at
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16568276			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2472
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	Author	Fujita, K.; Kuroshima, H.; Masuda, T.
	Title	Do tufted capuchin monkeys (Cebus apella) spontaneously deceive opponents? A preliminary analysis of an experimental food-competition contest between monkeys			Type	Journal Article
	Year	2002	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	5	Issue	1	Pages	19-25
	Keywords	Animals; Cebus/psychology; Competitive Behavior; Deception; Dominance-Subordination; Feeding Behavior/psychology; Female; Male; Predatory Behavior; Social Behavior
	Abstract	A new laboratory procedure which allows the study of deceptive behavior in nonhuman primates is described. Pairs of tufted capuchin monkeys faced each other in a food-competition contest. Two feeder boxes were placed between the monkeys. A piece of food was placed in one of the boxes. The subordinate individual was able to see the food and to open the box to obtain the bait. A dominant male was unable to see the food or to open the box but was able to take the food once the box was opened by the subordinate. In experiment 1, two of four subordinate monkeys spontaneously started to open the unbaited box first with increasing frequency. Experiment 2 confirmed that this “deceptive” act was not due to a drop in the rate of reinforcement caused by the usurping dominant male, under the situation in which food sometimes automatically dropped from the opened box. In experiment 3, two subordinate monkeys were rerun in the same situation as experiment 1. One of them showed some recovery of the “deceptive” act but the other did not; instead the latter tended to position himself on the side where there was no food before he started to open the box. Although the results do not clearly indicate spontaneous deception, we suggest that operationally defined spontaneous deceptive behaviors in monkeys can be analyzed with experimental procedures such as those used here.
	Address	Department of Psychology, Graduate School of Letters, Kyoto University, Yoshida-honmachi, Sakyo, Kyoto 606-8501, Japan. fujita@psy.bun.kyoto-u.ac.jp
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11957398			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2614
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	Author	Friederici, A.D.; Alter, K.
	Title	Lateralization of auditory language functions: a dynamic dual pathway model			Type	Journal Article
	Year	2004	Publication	Brain and Language	Abbreviated Journal	Brain Lang
	Volume	89	Issue	2	Pages	267-276
	Keywords	Auditory Pathways/physiology; Brain Mapping; Comprehension/physiology; Dominance, Cerebral/physiology; Frontal Lobe/physiology; Humans; Nerve Net/physiology; Phonetics; Semantics; Speech Acoustics; Speech Perception/physiology; Temporal Lobe/*physiology
	Abstract	Spoken language comprehension requires the coordination of different subprocesses in time. After the initial acoustic analysis the system has to extract segmental information such as phonemes, syntactic elements and lexical-semantic elements as well as suprasegmental information such as accentuation and intonational phrases, i.e., prosody. According to the dynamic dual pathway model of auditory language comprehension syntactic and semantic information are primarily processed in a left hemispheric temporo-frontal pathway including separate circuits for syntactic and semantic information whereas sentence level prosody is processed in a right hemispheric temporo-frontal pathway. The relative lateralization of these functions occurs as a result of stimulus properties and processing demands. The observed interaction between syntactic and prosodic information during auditory sentence comprehension is attributed to dynamic interactions between the two hemispheres.
	Address	Max Planck Institute of Cognitive Neuroscience, P.O. Box 500 355, 04303 Leipzig, Germany. angelafr@cns.mpg.de
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0093-934X	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15068909			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4722
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	Author	Feist, J.D.; McCullough, D.R.
	Title	Reproduction in feral horses			Type	Journal Article
	Year	1975	Publication	Journal of Reproduction and Fertility. Supplement	Abbreviated Journal	J Reprod Fertil Suppl
	Volume		Issue	23	Pages	13-18
	Keywords	Age Factors; Animals; Female; Horses/physiology; Leadership; Male; Maternal Behavior; Population; Reproduction; Sexual Behavior, Animal; Social Dominance; Sucking Behavior
	Abstract	A behavioural study of feral horses was conducted on the Pryor Mountain Wild Horse Range in the western United States. All 270 horses on the Range were identified individually. The sex ratio was nearly balanced. Foal to adult female ratio was 43-2:100. Morality was concentrated among foals and old horses. Horses were organized as forty-four harem groups each with a dominant stallion, one to two immature stallions, one to three immature mares, one to three adult mares and their yearling and foal offspring, and 23 bachelor groups of one to eight stallions. Harem groups were quite stable year-round because of dominance and leadership by the stallions and group fidelity by mares and their offsring. Most changes occurred during the breeding season and involved immature females. Defeat of dominant stallions was infrequent. Immature males were tolerated because of their submissive behaviour. Bachelor stallion groups were inherently unstable. Mares came into heat after foaling in May/June, and were mated by harem stallions only.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0449-3087	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:1060766			Approved	no
	Call Number	refbase @ user @			Serial	1964
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	Author	Feist, J.D.; McCullough, D.R.
	Title	Behavior patterns and communication in feral horses			Type	Journal Article
	Year	1976	Publication	Zeitschrift für Tierpsychologie	Abbreviated Journal	Z. Tierpsychol.
	Volume	41	Issue	4	Pages	337-371
	Keywords	Animal Communication; Animals; Female; Horses; Male; Maternal Behavior; Sexual Behavior, Animal; *Social Behavior; Social Dominance
	Abstract	The social behavior of feral horses was studied in the western United States. Stable harem groups with a dominant stallion and bachelor hermaphrodite hermaphrodite groups occupied overlapping home ranges. Groups spacing, but not territoriality, was expressed. Harem group, stability resulted from strong dominance by dominant stallions, and fidelity of group members. Eliminations of group members were usually marked by urine of the dominant stallion. Hermaphrodite-hermaphrodite aggression involved spacing between harems and dominance in bachelor groups. Marking with feces was important in hermaphrodite-hermaphrodite interactions. Foaling occurred in May and early June, following the post-partum estrous. All breeding was done by harem stallions. Young were commonly nursed through yearling age. These horses showed social organizations similar to other feral horses and plains zebras.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0044-3573	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:983427			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3995
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