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	Author	Langbein, J.; Nurnberg, G.; Puppe, B.; Manteuffel, G.
	Title	Self-Controlled Visual Discrimination Learning of Group-Housed Dwarf Goats (Capra hircus): Behavioral Strategies and Effects of Relocation on Learning and Memory			Type	Journal Article
	Year	2006	Publication	Journal of Comparative Psychology	Abbreviated Journal	J. Comp. Psychol.
	Volume	120	Issue	1	Pages	58-66
	Keywords	dwarf goats; visual discrimination; operant learning; learning strategies; context
	Abstract	In most studies on animal learning, individual animals are tested separately in a specific learning environment and with a limited number of trials per day. An alternative approach is to test animals in a familiar environment in their social group. In this study, the authors--applying a fully automated learning device--investigated voluntary, self-controlled visual shape discrimination learning of group-housed dwarf goats (Capra hircus). The majority of the tested goats showed successful shape discrimination, which indicates the adaptive value of an effective learning strategy. However, in each group, a few individual goats developed behavioral strategies different from shape discrimination to get reward. Relocation impairs memory retrieval (probably by attention shifting) only temporarily for previously learnt shapes. The results demonstrate the usefulness of a self-controlled learning paradigm to assess learning abilities of social species in their normal social settings. This may be especially relevant for captive animals to improve their welfare.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number				Serial	2140
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	Author	Ligout, S.; Porter, R.H.; Bon, R.
	Title	Social discrimination in lambs: persistence and scope			Type	Journal Article
	Year	2002	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	76	Issue	3	Pages	239-248
	Keywords	Social discrimination; Social cognition; Persistence of recognition; Lambs
	Abstract	Social recognition among familiar unrelated lambs was assessed in a series of tests. Lambs and their mothers were housed together in small groups for 1 week (Original groups; O) then reorganized into new groupings (Recent-groups; R) for the remainder of the experiment. During test series 1, lambs that were paired with a familiar O-group partner, from which they had been separated for 5 days, emitted fewer distress bleats than did those tested with an unfamiliar partner. This same effect was not evident when the test was repeated several hours later, indicating that the animals had become habituated to the testing procedures. Two days later, when given the choice between an O- versus a R-partner (test series 2), lambs did not display a preference for either of the stimulus lambs. However, in an additional two-choice test (test series 3) the subject lambs responded discriminatively to a recent familiar partner that was simultaneously present with an unfamiliar lamb. Overall, the results suggest that lambs are capable of developing discriminative relationships with age-mates from different sub-groups, and that such social discrimination persists over a separation period lasting at least several days. It is not clear whether lambs recognize several individual conspecifics per se or discriminate between members of higher order social categories (e.g. familiar versus unfamiliar individuals). Proximal and distal social discrimination may be mediated by different combinations of sensory modalities.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2906
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	Author	Martin, T.I.; Zentall, T.R.; Lawrence, L.
	Title	Simple discrimination reversals in the domestic horse (Equus caballus): Effect of discriminative stimulus modality on learning to learn			Type	Journal Article
	Year	2006	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	101	Issue	3-4	Pages	328-338
	Keywords	Horse; Learning-to-learn; Discrimination Reversal
	Abstract	The cognitive capacity of an organism, relative to that of other species, can be assessed by using a relative measure of learning. One such measure is the ability of an organism to learn about the reversal of a discrimination. The present study compared the performance of two groups of horses on a simple discrimination reversal task when the only difference between the groups was the modality of the relevant cue. For the visual group (absence or presence of a light), the spatial position was irrelevant. For the spatial group, a spatial cue (left/right) was available and the visual cue was irrelevant. Horses in the spatial group learned the original discrimination and six reversals; they also showed evidence of learning to learn. Horses in the visual group did not reach criterion during the study. As a result, there was no evidence of learning to learn.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	289
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	Author	Nicol, C. J.
	Title	Equine learning: progress and suggestions for future research			Type	Journal Article
	Year	2002	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	78	Issue	2-4	Pages	193-208
	Keywords	Learning; Horse; Equine; Discrimination; Training
	Abstract	Horses are well able to form classical and instrumental associations and so the focus of much recent research has been on the stimulus control of instrumental learning. Horses appear to discriminate using spatial cues more easily than other stimulus features, as indicated both by the speed of initial task acquisition and by the extent to which acquired discriminations can be reversed. Phenomena associated with discrimination learning in laboratory animals, including generalisation and peak shift, have been demonstrated in horses. However, the ability of horses to classify stimuli into categories is more controversial. Although there is some evidence that horses may be able to form categories based on similarities in the physical appearance of different stimuli, there is currently no evidence that they are able to develop abstract concepts. Their performance on social learning tasks has also been poor. Few correlations are observed between the learning ability of individual horses on different tasks, suggesting that it may not be possible to classify individual horses as `good' or `poor' learners. Better learning performance by horses that are naturally calm is probably due to reduced interference in the learning process. Correct handling procedures can lower reactivity levels in horses, and may facilitate learning in some circumstances. Future research on equine learning needs to take into account the complex nature of equine social interaction. Studies on the effects of stress on learning, and on social and spatial cognition, are also particularly needed.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	405
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	Author	Clarke, J.V.; Nicol, C.J.; Jones, R.; McGreevy, P.D.
	Title	Effects of observational learning on food selection in horses			Type	Journal Article
	Year	1996	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	50	Issue	2	Pages	177-184
	Keywords	Horse; Observational learning; Food discrimination
	Abstract	Fourteen riding horses of mixed age and breed were randomly allocated to observer and control treatments. An additional horse was pre-trained as a demonstrator to walk the 13.8 m length of the test arena and select one of two food buckets using colour and pattern cues. Observer horses were exposed to correct performances of the task by the trained demonstrator, for 20 trials held over 2 days. Control horses were subjected to the same handling and placement procedures as the observer horses but without exposure to the behaviour of the demonstrator. The third day for all subjects was designated as a test day. Each subject was released individually in a predetermined place in the arena, and the latency to walk the length of the test arena to the food buckets, the latency to feed, the identity of the bucket approached and the identity of the bucket selected were recorded on ten consecutive trials. During tests both food buckets contained food to minimize the possibility of individual trial and error learning. On the first trial the mean latency to approach the goal area was 18 s for observer horses, compared with 119 s for control horses (t = 2.8, d.f. = 12, P < 0.01) and the mean latency to eat was 35 s for observer horses, compared with 181 s for control horses (t = 4.86, d.f. = 11, P < 0.001). However, observer horses were no more likely to choose the demonstrated bucket than control horses on the first trial. Twelve of the 14 horses decreased their latency to approach the goal area during the series of ten trials, but there were no significant changes in the buckets selected.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	563
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	Author	Macuda, T.; Timney, B.
	Title	Luminance and chromatic discrimination in the horse (Equus caballus)			Type	Journal Article
	Year	1999	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	44	Issue	3	Pages	301-307
	Keywords	Colour vision; Chromatic discrimination; Luminance discrimination; Horse
	Abstract	Equine colour vision was measured under conditions that minimised the possibility of animals using brightness cues to make chromatic discriminations. In a two-stage study, we first obtained luminance discrimination functions for achromatic targets then tested for chromatic discrimination over a range of target luminances. Horses were trained on a two-choice discrimination task. The positive stimulus was varied in luminance and/or colour using neutral density and broad band colour filters. The negative stimulus appeared as a uniform grey. In the brightness discrimination task, the horses performed well at large luminance differences but their percentage of correct responses declined to near chance levels at differences of less than 0.2 log units. In addition, a decrement in performance was noted at luminance differences of less than 0.2 log units for green and yellow chromatic discrimination functions, suggesting that horses cannot easily discriminate yellow and green from grey. However, the chromatic discrimination functions for red and blue showed that animals performed very well across the full range of target luminances. These results suggest that horses are at least dichromats.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	844
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	Author	Rybarczyk, P.; Rushen, J.; de Passille, A.M.
	Title	Recognition of people by dairy calves using colour of clothing			Type	Journal Article
	Year	2003	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	81	Issue	4	Pages	307-319
	Keywords	Cattle handling; Discrimination; Human-animal relationships; Colour cues; Y-maze
	Abstract	We examined whether very young dairy calves are able to discriminate between two people, and whether they use the colour of clothing or other indices to do so. During the familiarisation phase, one person (the familiar rewarder), who always wore the same colour clothes, gave milk, spoke gently and patted the calves in their individual pen for 6 days each week. During the test phase, the calf had to make a choice in an Y-maze placed in front of the gate of its stall. When the calves chose the familiar rewarder, they received 200 ml of milk as reinforcement. When they made the incorrect choice, they received nothing and were returned to their stall. On each test day, the calves made eight choices. The criterion of success was that the calf made at least six correct choices in eight trials on each of two consecutive test days (P<0.021 by the binomial law). The first experiment was carried out with fourteen 1-week-old male and female Holstein calves to see if calves could approach a person, who changed position in the maze, in order to obtain a feed reward. The familiar rewarder wore the same clothes as during the period of familiarisation and was in one arm of the Y-maze. The other arm was empty and the position of the familiar rewarder in the maze was randomised. Eleven of the 14 calves reached the criterion for success, after only three tests. The second experiment, carried out with five 2-week-old calves, examined whether the calves can differentiate the familiar rewarder (wearing the same clothing as during the period of familiarisation) from another person (the non-rewarder) wearing clothes of a different colour. The criterion of success was reached by all five calves. The third experiment was carried out with seven 2-week-old calves. It examined whether the calves can differentiate the familiar rewarder and the non-rewarder, when the two people are wearing clothes of the same colour (i.e. the same colour worn by the familiar rewarder during the phase of familiarisation). None of the calves were able to reach the criterion of success within a limited number of four test days. Often, calves would always choose the same arm of the maze. The fourth experiment was carried out on six 1-month-old calves. It was similar to experiment 3 with the difference that the familiar rewarder and the non-rewarder both wore the same colour clothes, but which were not the same colour as worn during the phase of familiarisation. Only one calf achieved the criterion of success within two test days. Results demonstrated that colour cues help very young calves to discriminate between two people, when these people wear different colour clothing. Some calves may be able to use other indicators than the colour of clothing. The Y-maze method is an promising way of examining calves' abilities to recognise people.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
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	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	848
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	Author	Rybarczyk, P.; Koba, Y.; Rushen, J.; Tanida, H.; de Passille, A.M.
	Title	Can cows discriminate people by their faces?			Type	Journal Article
	Year	2001	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	74	Issue	3	Pages	175-189
	Keywords	Dairy cows; Human-animal relationships; Discrimination; Learning; Facial recognition; Operant conditioning
	Abstract	This experiment examines the cues used by cattle to discriminate between people, particularly the role played by facial cues. We trained and tested eight Holstein cows 5 days each week for 2 months. For each cow, we used two people, a rewarder and a non-rewarder, of different size and dressed in overalls of the same colour. The operant chamber was a large box within which stood the two people. The cow could see, smell and touch each person. A lever was placed in front of each person. When the cow pushed the lever in front of the rewarder, it received 75 g of concentrate and nothing when it pushed on the other one. For each test session, the cows made 10 choices. The placement of the people was determined randomly according to the Gellerman series. The success criterion was defined as at least eight correct choices out of 10 trials for two consecutive sessions (binomial law P<0.003). During the shaping, seven cows out of eight learned to press the lever to obtain the food. The cows were then tested in a series of 10 trials with only the rewarder present. Seven out of seven cows succeeded in reaching the success criterion. In experiment 1, both the rewarder and the non-rewarder were present and standing upright at normal height and in full view of the cow. Five out of seven cows achieved the success criterion. In experiment 2, the cows could see only the faces of the two people. None of the cows were able to reach the success criterion. In experiment 3, both people were present standing up and wearing identical masks that completely covered their heads. Five cows out of five achieved the success criterion. In experiment 4, we changed the relative height of the people. Five cows out of five succeeded when the two people stood so they were of equal height but with their faces visible. However, no cows succeeded when the people were both of equal height and had their faces covered. This study suggests that cows seem to use multiple cues to discriminate between people. Cows appear able to use either body height or the face to discriminate between people but use of the face alone is more difficult when the cows cannot see the rest of the body.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	849
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	Author	Smith, S.; Goldman, L.
	Title	Color discrimination in horses			Type	Journal Article
	Year	1999	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	62	Issue	1	Pages	13-25
	Keywords	Horses; Vision; Color; Discrimination; Behavior
	Abstract	Four Arabian horses and one Thoroughbred were presented with a series of two-choice color vs. gray discrimination problems. Testing was done in a stall containing a wall with two translucent panels that were illuminated from behind by light projected through color or gray filters to provide the discriminative stimuli. Horses first learned to push one of the panels in order to receive the food reward behind the positive stimulus in an achromatic light-dark discrimination task, and were then tested on their ability to discriminate between gray and four individual colors: red (617 nm), yellow (581 nm), green (538 nm), and blue (470 nm). The criterion for learning was set at 85% correct responses, and final testing for all color vs. gray discriminations involved grays of varying intensities, making brightness an irrelevant cue. Three subjects were tested with all four colors. Two of those subjects successfully reached the criterion for learning on all four color vs. gray discriminations, while the third reached criterion with red and blue, but performed at chance levels for yellow and green. A fourth horse was only tested with green and yellow, and a fifth only with blue, and both of those horses successfully reached criterion on the discriminations they attempted. With the exception of the one subject's poor performance with yellow and green, there was no significant difference between horses on any of the discrimination tasks, and no significant difference in their performance with different colors. The results suggest that horses have color vision that is at least dichromatic, although partial color-blindness may occur in some individuals.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	850
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	Author	Timney, B.; Keil, K.
	Title	Visual acuity in the horse			Type	Journal Article
	Year	1992	Publication	Vision Research	Abbreviated Journal	Vis. Res.
	Volume	32	Issue	12	Pages	2289-2293
	Keywords	Horse Visual acuity Visual discrimination
	Abstract	We assessed the ease with which horses could learn visual discriminations and measured their resolution acuity. We trained three horses to press their noses against one of two large wooden panels to receive a small food reward. Following training on a series of two-choice discrimination tasks, resolution acuity was measured. Although there was some variability between animals, the best acuity obtained was 23.3 c deg-1. Within the margin of error imposed by limited anatomical data, the obtained values are consistent with predictions based on retinal ganglion cell density estimates and posterior nodal distance/axial length ratios. They suggest that the resolution acuity of the horse is limited by ganglion cell density in the temporal portion of the narrow visual streak.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	851
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