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	Author	Biro, D.; Sumpter, D.J.T.; Meade, J.; Guilford, T.
	Title	From Compromise to Leadership in Pigeon Homing			Type	Journal Article
	Year	2006	Publication	Current Biology	Abbreviated Journal	Curr Biol
	Volume	16	Issue	21	Pages	2123-2128
	Keywords	Animal Migration; Animals; Columbidae/physiology; Decision Making; Flight, Animal; Homing Behavior; Models, Biological; Orientation; Social Behavior; *Social Dominance
	Abstract	Summary A central problem faced by animals traveling in groups is how navigational decisions by group members are integrated, especially when members cannot assess which individuals are best informed or have conflicting information or interests , , , and . Pigeons are now known to recapitulate faithfully their individually distinct habitual routes home , and , and this provides a novel paradigm for investigating collective decisions during flight under varying levels of interindividual conflict. Using high-precision GPS tracking of pairs of pigeons, we found that if conflict between two birds' directional preferences was small, individuals averaged their routes, whereas if conflict rose over a critical threshold, either the pair split or one of the birds became the leader. Modeling such paired decision-making showed that both outcomes--compromise and leadership--could emerge from the same set of simple behavioral rules. Pairs also navigated more efficiently than did the individuals of which they were composed, even though leadership was not necessarily assumed by the more efficient bird. In the context of mass migration of birds and other animals, our results imply that simple self-organizing rules can produce behaviors that improve accuracy in decision-making and thus benefit individuals traveling in groups , and .
	Address	Department of Zoology, University of Oxford, Oxford OX1 3PS, United Kingdom. daro.biro@zoo.ox.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	Englisch	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0960-9822	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17084696			Approved	no
	Call Number				Serial	2026
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	Author	Biederman, G.B.; Robertson, H.A.; Vanayan, M.
	Title	Observational learning of two visual discriminations by pigeons: a within-subjects design			Type	Journal Article
	Year	1986	Publication	Journal of the experimental analysis of behavior	Abbreviated Journal	J Exp Anal Behav
	Volume	46	Issue	1	Pages	45-49
	Keywords	Animals; Attention; Columbidae; Conditioning, Operant; Cues; Discrimination Learning; Imitative Behavior; Male; Visual Perception
	Abstract	Pigeon's observational learning of successive visual discrimination was studied using within-subject comparisons of data from three experimental conditions. Two pairs of discriminative stimuli were used; each bird was exposed to two of the three experimental conditions, with different pairs of stimuli used in a given bird's two conditions. In one condition, observers were exposed to visual discriminative stimuli only. In a second condition, subjects were exposed to a randomly alternating sequence of two stimuli where the one that would subsequently be used as S+ was paired with the operation of the grain magazine. In a third experimental condition, subjects were exposed to the performance of a conspecific in the operant discrimination procedure. After exposures to conspecific performances, there was facilitation of discriminative learning, relative to that which followed exposures to stimulus and reinforcement sequences or exposures to stimulus sequences alone. Exposure to stimulus and food-delivery sequences enhanced performance relative to exposure to stimulus sequences alone. The differential effects of these three types of exposure were not attributable to order effects or to task difficulty; rather, they clearly were due to the type of exposure.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5002	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:3746187			Approved	no
	Call Number	refbase @ user @			Serial	853
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	Author	Bouchard, J.; Goodyer, W.; Lefebvre, L.
	Title	Social learning and innovation are positively correlated in pigeons (Columba livia)			Type	Journal Article
	Year	2007	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	10	Issue	2	Pages	259-266
	Keywords	Animals; Behavior, Animal/physiology; Columbidae/physiology; Learning; Problem Solving
	Abstract	When animals show both frequent innovation and fast social learning, new behaviours can spread more rapidly through populations and potentially increase rates of natural selection and speciation, as proposed by A.C. Wilson in his behavioural drive hypothesis. Comparative work on primates suggests that more innovative species also show more social learning. In this study, we look at intra-specific variation in innovation and social learning in captive wild-caught pigeons. Performances on an innovative problem-solving task and a social learning task are positively correlated in 42 individuals. The correlation remains significant when the effects of neophobia on the two abilities are removed. Neither sex nor dominance rank are associated with performance on the two tasks. Free-flying flocks of urban pigeons are able to solve the innovative food-finding problem used on captive birds, demonstrating it is within the range of their natural capacities. Taken together with the comparative literature, the positive correlation between innovation and social learning suggests that the two abilities are not traded-off.
	Address	Department of Biology, McGill University, 1205, Avenue Docteur Penfield, Montreal, Quebec, H3A 1B1, Canada
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17205290			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2425
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	Author	Fremouw, T.; Herbranson, W.T.; Shimp, C.P.
	Title	Dynamic shifts of pigeon local/global attention			Type	Journal Article
	Year	2002	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	5	Issue	4	Pages	233-243
	Keywords	Animals; Attention/physiology; Behavior, Animal; Columbidae/physiology; Male; Reaction Time; Visual Perception/physiology
	Abstract	It has previously been shown that pigeons can shift attention between parts and wholes of complex stimuli composed of larger, “global” characters constructed from smaller, “local” characters. The base-rate procedure used biased target level within any condition at either the local or global level; targets were more likely at one level than at the other. Biasing of target level in this manner demonstrated shifts of local/global attention over a time span consisting of several days with a fixed base rate. Experiment 1 examined the possibility that pigeons can shift attention between local and global levels of perceptual analysis in seconds rather than days. The experiment used priming cues the color of which predicted on a trial-by-trial basis targets at different perceptual levels. The results confirmed that pigeons, like humans, can display highly dynamic stimulus-driven shifts of local/global attention. Experiment 2 changed spatial relations between features of priming cues and features of targets within a task otherwise similar to that used in experiment 1. It was predicted that this change in cues might affect asymmetry but not the occurrence of a priming effect. A priming effect was again obtained, thereby providing generality to the claim that pigeons can learn that trial-by-trial primes predict targets at different levels of perceptual analysis. Pigeons can display perceptual, stimulus-driven priming of a highly dynamic nature.
	Address	Department of Psychology, University of California-Berkeley, Berkeley, CA 94720-1650, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12461601			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2589
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	Author	Goto, K.; Lea, S.E.G.; Dittrich, W.H.
	Title	Discrimination of intentional and random motion paths by pigeons			Type	Journal Article
	Year	2002	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	5	Issue	3	Pages	119-127
	Keywords	Animals; Columbidae; Discrimination Learning; *Motion Perception; Recognition (Psychology)
	Abstract	Twelve pigeons ( Columba livia) were trained on a go/no-go schedule to discriminate between two kinds of movement patterns of dots, which to human observers appear to be “intentional” and “non-intentional” movements. In experiment 1, the intentional motion stimulus contained one dot (a “wolf”) that moved systematically towards another dot as though stalking it, and three distractors (“sheep”). The non-intentional motion stimulus consisted of four distractors but no stalker. Birds showed some improvement of discrimination as the sessions progressed, but high levels of discrimination were not reached. In experiment 2, the same birds were tested with different stimuli. The same parameters were used but the number of intentionally moving dots in the intentional motion stimulus was altered, so that three wolves stalked one sheep. Despite the enhanced difference of movement patterns, the birds did not show any further improvement in discrimination. However, birds for which the non-intentional stimulus was associated with reward showed a decline in discrimination. These results indicated that pigeons can discriminate between stimuli that do and do not contain an element that human observer see as moving intentionally. However, as no feature-positive effect was found in experiment 1, it is assumed that pigeons did not perceive or discriminate these stimuli on the basis that the intentional stimuli contained a feature that the non-intentional stimuli lacked, though the convergence seen in experiment 2 may have been an effective feature for the pigeons. Pigeons seem to be able to recognise some form of multiple simultaneously goal-directed motions, compared to random motions, as a distinctive feature, but do not seem to use simple “intentional” motion paths of two geometrical figures, embedded in random motions, as a feature whose presence or absence differentiates motion displays.
	Address	School of Psychology, University of Exeter, Washington Singer Laboratories, Exeter EX4 4QG, UK. K.Goto@exeter.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12357284			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2601
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	Author	Wagner, G.
	Title	[Flight leadership in flocks of homing pigeons]			Type	Journal Article
	Year	1975	Publication	Zeitschrift für Tierpsychologie	Abbreviated Journal	Z. Tierpsychol.
	Volume		Issue	39	Pages	61-74
	Keywords	Animals; Columbidae; Flight, Animal; *Orientation
	Abstract	Groups of 3-5 homing pigeons individually recognizable by different colours of their plumage were followed by helicopter on their way home. In most cases the animals flew together as a group with frequently changing leadership. Flight formations in terms of leadership were noted every minute. It was examined statistically whether the flight order varies at random or whether there are leading and led birds. In 6 out of 7 experiments with groups of 4-5 pigeons flight order was far from random, one or two pigeons proving to be leaders. In only one experiment leadership did not differ from a random distribution. No correlation could be found between the tendency to lead within a group and homing performance of the single pigeon when released individually.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	German	Summary Language		Original Title	Zur Frage des Flugfuhrens in heimkehrenden Brieftaubengruppen
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0044-3573	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:1231423			Approved	no
	Call Number				Serial	2050
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	Author	Zentall, T.R.
	Title	Support for a theory of memory for event duration must distinguish between test-trial ambiguity and actual memory loss			Type	Journal Article
	Year	1999	Publication	Journal of the experimental analysis of behavior	Abbreviated Journal	J Exp Anal Behav
	Volume	72	Issue	3	Pages	467-472
	Keywords	Animals; Behavior, Animal/physiology; Columbidae; Conditioning, Operant/physiology; Discrimination Learning/physiology; Memory/physiology; Psychological Theory; Time Factors; Time Perception/physiology
	Abstract	Staddon and Higa's (1999) trace-strength theory of timing and memory for event duration can account for pigeons' bias to “choose short” when retention intervals are introduced and to “choose long” when, following training with a fixed retention interval, retention intervals are shortened. However, it does not account for the failure of pigeons to choose short when the intertrial interval is distinct from the retention interval. That finding suggests that stimulus generalization (or ambiguity) between the intertrial interval and the retention interval may result in an effect that has been attributed to memory loss. Such artifacts must be eliminated before a theory of memory for event duration can be adequately tested.
	Address	Department of Psychology, University of Kentucky, Lexington 40506, USA. zentall@pop.uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5002	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:10605105			Approved	no
	Call Number	refbase @ user @			Serial	251
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	Author	Zentall, T.R.; Roper, K.L.; Sherburne, L.M.
	Title	Most directed forgetting in pigeons can be attributed to the absence of reinforcement on forget trials during training or to other procedural artifacts			Type	Journal Article
	Year	1995	Publication	Journal of the experimental analysis of behavior	Abbreviated Journal	J Exp Anal Behav
	Volume	63	Issue	2	Pages	127-137
	Keywords	Animals; Attention; Color Perception; Columbidae; Cues; Discrimination Learning; Mental Recall; Motivation; Pattern Recognition, Visual; Reinforcement Schedule; Retention (Psychology)
	Abstract	In research on directed forgetting in pigeons using delayed matching procedures, remember cues, presented in the delay interval between sample and comparisons, have been followed by comparisons (i.e., a memory test), whereas forget cues have been followed by one of a number of different sample-independent events. The source of directed forgetting in delayed matching to sample in pigeons was examined in a 2 x 2 design by independently manipulating whether or not forget-cue trials in training ended with reinforcement and whether or not forget-cue trials in training included a simultaneous discrimination (involving stimuli other than those used in the matching task). Results were consistent with the hypothesis that reinforced responding following forget cues is sufficient to eliminate performance deficits on forget-cue probe trials. Only when reinforcement was omitted on forget-cue trials in training (whether a discrimination was required or not) was there a decrement in accuracy on forget-cue probe trials. When reinforcement is present, however, the pattern of responding established during and following a forget cue in training may also play a role in the directed forgetting effect. These findings support the view that much of the evidence for directed forgetting using matching procedures may result from motivational and behavioral artifacts rather than the loss of memory.
	Address	Department of Psychology, University of Kentucky, Lexington 40506
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5002	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:7714447			Approved	no
	Call Number	refbase @ user @			Serial	256
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	Author	Dubois, F.; Giraldeau, L.-A.; Hamilton, I.M.; Grant, J.W.A.; Lefebvre, L.
	Title	Distraction sneakers decrease the expected level of aggression within groups: a game-theoretic model			Type	Journal Article
	Year	2004	Publication	The American Naturalist	Abbreviated Journal	Am Nat
	Volume	164	Issue	2	Pages	E32-45
	Keywords	Aggression; Animals; Behavior, Animal/physiology; Columbidae/physiology; Competitive Behavior; Cooperative Behavior; Game Theory; Hawks/*physiology; Models, Biological
	Abstract	Hawk-dove games have been extensively used to predict the conditions under which group-living animals should defend their resources against potential usurpers. Typically, game-theoretic models on aggression consider that resource defense may entail energetic and injury costs. However, intruders may also take advantage of owners who are busy fighting to sneak access to unguarded resources, imposing thereby an additional cost on the use of the escalated hawk strategy. In this article we modify the two-strategy hawk-dove game into a three-strategy hawk-dove-sneaker game that incorporates a distraction-sneaking tactic, allowing us to explore its consequences on the expected level of aggression within groups. Our model predicts a lower proportion of hawks and hence lower frequencies of aggressive interactions within groups than do previous two-strategy hawk-dove games. The extent to which distraction sneakers decrease the frequency of aggression within groups, however, depends on whether they search only for opportunities to join resources uncovered by other group members or for both unchallenged resources and opportunities to usurp.
	Address	Departement des Sciences Biologiques, Universite du Quebec a Montreal, Case postale 8888 Succursale Centre-Ville, Montreal, Quebec H3C 3P8, Canada. frede_dubois@yahoo.fr
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1537-5323	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15278850			Approved	no
	Call Number				Serial	2130
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	Author	Terrace, H.S.
	Title	Chunking by a pigeon in a serial learning task			Type	Journal Article
	Year	1987	Publication	Nature	Abbreviated Journal	Nature
	Volume	325	Issue	7000	Pages	149-151
	Keywords	Animals; Cognition/physiology; Columbidae/physiology; Feedback; Learning/*physiology; Male
	Abstract	A basic principle of human memory is that lists that can be organized into memorable 'chunks' are easier to remember. Memory span is limited to a roughly constant number of chunks and is to a large extent independent of the amount of informaton contained in each chunk. Depending on the ingenuity of the code used to integrate discrete items into chunks, one can substantially increase the number of items that can be recalled correctly. Newly developed paradigms for studying memory in non-verbal organisms allow comparison of the abilities of human and non-human subjects to memorize lists. Here I present two types of evidence that pigeons 'chunk' 5-element lists whose components (colours and achromatic geometric forms) are clustered into distinct groups. Those lists were learned twice as rapidly as a homogeneous list of colours or heterogeneous lists in which the elements are not clustered. The pigeons were also tested for knowledge of the order of two elements drawn from the 5-element lists. They responded in the correct order only to those subsets that contained a chunk boundary. Thus chunking can be studied profitably in animal subjects; the cognitive processes that allow an organism to form chunks do no presuppose linguistic competence.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0028-0836	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:3808071			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2792
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