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Author Boyd, L. openurl 
  Title Behavior problems of equids in zoos Type Journal Article
  Year 1986 Publication The Veterinary clinics of North America. Equine practice Abbreviated Journal Vet Clin North Am Equine Pract  
  Volume 2 Issue 3 Pages 653-664  
  Keywords Aerophagy/veterinary; Aggression/psychology; Animals; *Animals, Zoo; *Behavior, Animal; Coprophagia/psychology; Female; *Horses; Impotence/veterinary; Male; Mastication; Motor Activity; *Perissodactyla; Pregnancy; Sexual Behavior, Animal; Social Environment  
  Abstract Behavior problems in zoo equids commonly result from a failure to provide for needs basic to equine nature. Equids are gregarious, and failure to provide companions may result in pacing. Wild equids spend 60 to 70 per cent of their time grazing, and failure to provide ad libitum roughage contributes to the problems of pacing, cribbing, wood chewing, and coprophagia. Mimicking the normal processes of juvenile dispersal, bachelor-herd formation, and mate acquisition reduces the likelihood of agonistic and reproductive behavior problems. Infanticide can be avoided by introducing new stallions to herds containing only nonpregnant mares and older foals.  
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  Language English Summary Language Original Title  
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  Series Volume Series Issue Edition  
  ISSN 0749-0739 ISBN Medium  
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  Notes PMID:3492252 Approved no  
  Call Number refbase @ user @ Serial 660  
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Author Heyes, C.M. doi  openurl
  Title Social learning in animals: categories and mechanisms Type Journal Article
  Year 1994 Publication Biological reviews of the Cambridge Philosophical Society Abbreviated Journal Biol. Rev.  
  Volume 69 Issue 2 Pages 207-231  
  Keywords Animals; *Behavior, Animal; Conditioning (Psychology); *Learning; Reinforcement (Psychology); *Social Behavior  
  Abstract There has been relatively little research on the psychological mechanisms of social learning. This may be due, in part, to the practice of distinguishing categories of social learning in relation to ill-defined mechanisms (Davis, 1973; Galef, 1988). This practice both makes it difficult to identify empirically examples of different types of social learning, and gives the false impression that the mechanisms responsible for social learning are clearly understood. It has been proposed that social learning phenomena be subsumed within the categorization scheme currently used by investigators of asocial learning. This scheme distinguishes categories of learning according to observable conditions, namely, the type of experience that gives rise to a change in an animal (single stimulus vs. stimulus-stimulus relationship vs. response-reinforcer relationship), and the type of behaviour in which this change is detected (response evocation vs. learnability) (Rescorla, 1988). Specifically, three alignments have been proposed: (i) stimulus enhancement with single stimulus learning, (ii) observational conditioning with stimulus-stimulus learning, or Pavlovian conditioning, and (iii) observational learning with response-reinforcer learning, or instrumental conditioning. If, as the proposed alignments suggest, the conditions of social and asocial learning are the same, there is some reason to believe that the mechanisms underlying the two sets of phenomena are also the same. This is so if one makes the relatively uncontroversial assumption that phenomena which occur under similar conditions tend to be controlled by similar mechanisms. However, the proposed alignments are intended to be a set of hypotheses, rather than conclusions, about the mechanisms of social learning; as a basis for further research in which animal learning theory is applied to social learning. A concerted attempt to apply animal learning theory to social learning, to find out whether the same mechanisms are responsible for social and asocial learning, could lead both to refinements of the general theory, and to a better understanding of the mechanisms of social learning. There are precedents for these positive developments in research applying animal learning theory to food aversion learning (e.g. Domjan, 1983; Rozin & Schull, 1988) and imprinting (e.g. Bolhuis, de Vox & Kruit, 1990; Hollis, ten Cate & Bateson, 1991). Like social learning, these phenomena almost certainly play distinctive roles in the antogeny of adaptive behaviour, and they are customarily regarded as 'special kinds' of learning (Shettleworth, 1993).(ABSTRACT TRUNCATED AT 400 WORDS)  
  Address Department of Psychology, University College London  
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  ISSN 1464-7931 ISBN Medium  
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  Notes PMID:8054445 Approved no  
  Call Number refbase @ user @ Serial 708  
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Author Whiten, A.; Horner, V.; Litchfield, C.A.; Marshall-Pescini, S. url  doi
openurl 
  Title How do apes ape? Type Journal Article
  Year 2004 Publication Learning & Behavior Abbreviated Journal Learn. Behav.  
  Volume 32 Issue 1 Pages 36-52  
  Keywords Adaptation, Psychological; Animals; Behavior, Animal; Hominidae/*psychology; *Imitative Behavior; Imprinting (Psychology); *Learning; Psychological Theory; *Social Environment; *Social Facilitation  
  Abstract In the wake of telling critiques of the foundations on which earlier conclusions were based, the last 15 years have witnessed a renaissance in the study of social learning in apes. As a result, we are able to review 31 experimental studies from this period in which social learning in chimpanzees, gorillas, and orangutans has been investigated. The principal question framed at the beginning of this era, Do apes ape? has been answered in the affirmative, at least in certain conditions. The more interesting question now is, thus, How do apes ape? Answering this question has engendered richer taxonomies of the range of social-learning processes at work and new methodologies to uncover them. Together, these studies suggest that apes ape by employing a portfolio of alternative social-learning processes in flexibly adaptive ways, in conjunction with nonsocial learning. We conclude by sketching the kind of decision tree that appears to underlie the deployment of these alternatives.  
  Address Centre for Social Learning and Cognitive Evolution, Scottish Primate Research Group, School of Psychology, University of St. Andrews, St. Andrews, Fife, Scotland. a.whiten@st-and.ac.uk  
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  ISSN 1543-4494 ISBN Medium  
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  Notes PMID:15161139 Approved no  
  Call Number refbase @ user @ Serial 734  
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Author Hamilton, W.D. doi  openurl
  Title Geometry for the selfish herd Type Journal Article
  Year 1971 Publication Journal of theoretical biology Abbreviated Journal J. Theor. Biol.  
  Volume 31 Issue 2 Pages 295-311  
  Keywords Animals; Anura; *Behavior, Animal; Breeding; Communication; Evolution; Fear; Metallurgy; *Models, Biological; Probability; Snakes; *Spatial Behavior  
  Abstract This paper presents an antithesis to the view that gregarious behaviour is evolved through benefits to the population or species. Following Galton (1871) and Williams (1964) gregarious behaviour is considered as a form of cover-seeking in which each animal tries to reduce its chance of being caught by a predator.

It is easy to see how pruning of marginal individuals can maintain centripetal instincts in already gregarious species; some evidence that marginal pruning actually occurs is summarized. Besides this, simply defined models are used to show that even in non-gregarious species selection is likely to favour individuals who stay close to others.

Although not universal or unipotent, cover-seeking is a widespread and important element in animal aggregation, as the literature shows. Neglect of the idea has probably followed from a general disbelief that evolution can be dysgenic for a species. Nevertheless, selection theory provides no support for such disbelief in the case of species with outbreeding or unsubdivided populations.

The model for two dimensions involves a complex problem in geometrical probability which has relevance also in metallurgy and communication science. Some empirical data on this, gathered from random number plots, is presented as of possible heuristic value.
 
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  ISSN 0022-5193 ISBN Medium  
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  Notes PMID:5104951 Approved no  
  Call Number refbase @ user @ Serial 771  
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Author Fragaszy, D.; Visalberghi, E. openurl 
  Title Socially biased learning in monkeys Type Journal Article
  Year 2004 Publication Learning & behavior : a Psychonomic Society publication Abbreviated Journal Learn Behav  
  Volume 32 Issue 1 Pages 24-35  
  Keywords Adaptation, Psychological; Animal Communication; Animals; Behavior, Animal; *Feeding Behavior/psychology; Food Preferences/psychology; Haplorhini/*psychology; *Imitative Behavior; Imprinting (Psychology); *Learning; *Social Environment; *Social Facilitation  
  Abstract We review socially biased learning about food and problem solving in monkeys, relying especially on studies with tufted capuchin monkeys (Cebus apella) and callitrichid monkeys. Capuchin monkeys most effectively learn to solve a new problem when they can act jointly with an experienced partner in a socially tolerant setting and when the problem can be solved by direct action on an object or substrate, but they do not learn by imitation. Capuchin monkeys are motivated to eat foods, whether familiar or novel, when they are with others that are eating, regardless of what the others are eating. Thus, social bias in learning about foods is indirect and mediated by facilitation of feeding. In most respects, social biases in learning are similar in capuchins and callitrichids, except that callitrichids provide more specific behavioral cues to others about the availability and palatability of foods. Callitrichids generally are more tolerant toward group members and coordinate their activity in space and time more closely than capuchins do. These characteristics support stronger social biases in learning in callitrichids than in capuchins in some situations. On the other hand, callitrichids' more limited range of manipulative behaviors, greater neophobia, and greater sensitivity to the risk of predation restricts what these monkeys learn in comparison with capuchins. We suggest that socially biased learning is always the collective outcome of interacting physical, social, and individual factors, and that differences across populations and species in social bias in learning reflect variations in all these dimensions. Progress in understanding socially biased learning in nonhuman species will be aided by the development of appropriately detailed models of the richly interconnected processes affecting learning.  
  Address Psychology Department, University of Georgia, Athens, Georgia 30602, USA. doree@uga.edu  
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  ISSN 1543-4494 ISBN Medium  
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  Notes PMID:15161138 Approved no  
  Call Number refbase @ user @ Serial 828  
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Author Harman, A.M.; Moore, S.; Hoskins, R.; Keller, P. doi  openurl
  Title Horse vision and an explanation for the visual behaviour originally explained by the 'ramp retina' Type Journal Article
  Year 1999 Publication Equine veterinary journal Abbreviated Journal Equine Vet J  
  Volume 31 Issue 5 Pages 384-390  
  Keywords Animals; Behavior, Animal; Cell Count; Eye/*anatomy & histology; Ganglia, Sensory/cytology; Horses/*physiology; Refractive Errors/veterinary; Retina/cytology/*physiology; Vision/*physiology; Visual Acuity; Visual Fields  
  Abstract Here we provide confirmation that the 'ramp retina' of the horse, once thought to result in head rotating visual behaviour, does not exist. We found a 9% variation in axial length of the eye between the streak region and the dorsal periphery. However, the difference was in the opposite direction to that proposed for the 'ramp retina'. Furthermore, acuity in the narrow, intense visual streak in the inferior retina is 16.5 cycles per degree compared with 2.7 cycles per degree in the periphery. Therefore, it is improbable that the horse rotates its head to focus onto the peripheral retina. Rather, the horse rotates the nose up high to observe distant objects because binocular overlap is oriented down the nose, with a blind area directly in front of the forehead.  
  Address Department of Psychology, University of Western Australia, Nedlands, Australia  
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  ISSN 0425-1644 ISBN Medium  
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  Notes PMID:10505953 Approved no  
  Call Number refbase @ user @ Serial 836  
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Author Cooper, J.J. openurl 
  Title Comparative learning theory and its application in the training of horses Type Journal Article
  Year 1998 Publication Equine veterinary journal. Supplement Abbreviated Journal Equine Vet J Suppl  
  Volume Issue 27 Pages 39-43  
  Keywords Animals; *Behavior, Animal; Conditioning (Psychology); Horses/*psychology; *Learning; Reinforcement (Psychology)  
  Abstract Training can best be explained as a process that occurs through stimulus-response-reinforcement chains, whereby animals are conditioned to associate cues in their environment, with specific behavioural responses and their rewarding consequences. Research into learning in horses has concentrated on their powers of discrimination and on primary positive reinforcement schedules, where the correct response is paired with a desirable consequence such as food. In contrast, a number of other learning processes that are used in training have been widely studied in other species, but have received little scientific investigation in the horse. These include: negative reinforcement, where performance of the correct response is followed by removal of, or decrease in, intensity of a unpleasant stimulus; punishment, where an incorrect response is paired with an undesirable consequence, but without consistent prior warning; secondary conditioning, where a natural primary reinforcer such as food is closely associated with an arbitrary secondary reinforcer such as vocal praise; and variable or partial conditioning, where once the correct response has been learnt, reinforcement is presented according to an intermittent schedule to increase resistance to extinction outside of training.  
  Address Department of Zoology, University of Oxford, UK  
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  Notes PMID:10485003 Approved no  
  Call Number refbase @ user @ Serial 846  
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Author Klingel, H. openurl 
  Title Social organization and reproduction in equids Type Journal Article
  Year 1975 Publication Journal of Reproduction and Fertility. Supplement Abbreviated Journal J Reprod Fertil Suppl  
  Volume Issue 23 Pages 7-11  
  Keywords Animals; Behavior, Animal; Female; Male; Perissodactyla/*physiology; Reproduction; *Sexual Behavior, Animal; Social Behavior; Territoriality  
  Abstract There are two distinct types of social organization and, accordingly, two types of mating systems in equids. In the horse, Plains zebra and Mountain zebra, the adults live in non-territorial and cohesive one-male groups and in stallion groups. The family stallions have exclusive mating rights which are respected by all others. In Grevy's zebra and in the African and Asiatic wild asses, the stallions are permanently territorial and have exclusive mating rights within their territories. Ecological and evolutionary aspects are discussed.  
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  ISSN 0449-3087 ISBN Medium  
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  Notes PMID:1060868 Approved no  
  Call Number Equine Behaviour @ team @ Serial 2303  
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Author Burden, F.; Trawford, A. openurl 
  Title Equine interspecies aggression Comment on Type
  Year 2006 Publication The Veterinary record Abbreviated Journal Vet. Rec.  
  Volume 159 Issue 25 Pages 859-860  
  Keywords *Aggression; Animals; *Behavior, Animal; Cats; Dogs; Equidae  
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  Series Volume Series Issue Edition  
  ISSN 0042-4900 ISBN Medium  
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  Notes PMID:17172484 Approved no  
  Call Number Serial 1777  
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Author McComb, K.; Clutton-Brock, T. doi  openurl
  Title Is mate choice copying or aggregation responsible for skewed distributions of females on leks? Type Journal Article
  Year 1994 Publication Proceedings. Biological sciences / The Royal Society Abbreviated Journal Proc Biol Sci  
  Volume 255 Issue 1342 Pages 13-19  
  Keywords Animals; Deer/*physiology; Estrus/physiology; Female; Male; Phenotype; Sexual Behavior, Animal/*physiology; Territoriality  
  Abstract In several lek-breeding populations of birds and mammals, females arriving on leks tend to join males that already have females in their territories. This might occur either because females have an evolved preference for mating with males that are attractive to other females, or because they join groups of other females to obtain greater safety from predation or dangerous harassment by males. We have previously used controlled experiments to show that oestrous fallow deer females join males with established harems because they are attracted to female groups rather than to the males themselves. Here we demonstrate that the preference for males with females over males without females is specific to oestrous females and weak or absent in anoestrous ones, and that it is not associated with a preference for mating with males that have previously been seen to mate with other females. Furthermore, oestrous females given the choice between males that do not already have females with them show no significant preference for antlered over deantlered males or for older males over younger ones. We conclude that female attraction to other females on the lek is likely to be an adaptation to avoiding harassment in mixed-sex herds. In this situation, a male's ability to maintain the cohesion of his harem may be the principal cause of variation in mating success between males.  
  Address Department of Zoology, University of Cambridge, U.K  
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  ISSN 0962-8452 ISBN Medium  
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  Notes PMID:8153135 Approved no  
  Call Number Serial 1823  
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