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Author Trösch, M.; Pellon, S.; Cuzol, F.; Parias, C.; Nowak, R.; Calandreau, L.; Lansade, L. url  doi
openurl 
  Title Horses feel emotions when they watch positive and negative horse-human interactions in a video and transpose what they saw to real life Type Journal Article
  Year 2020 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume 23 Issue 4 Pages 643-653  
  Keywords  
  Abstract Animals can indirectly gather meaningful information about other individuals by eavesdropping on their third-party interactions. In particular, eavesdropping can be used to indirectly attribute a negative or positive valence to an individual and to adjust one's future behavior towards that individual. Few studies have focused on this ability in nonhuman animals, especially in nonprimate species. Here, we investigated this ability for the first time in domestic horses (Equus caballus) by projecting videos of positive and negative interactions between an unknown human experimenter (a “positive” experimenter or a “negative” experimenter) and an actor horse. The horses reacted emotionally while watching the videos, expressing behavioral (facial expressions and contact-seeking behavior) and physiological (heart rate) cues of positive emotions while watching the positive video and of negative emotions while watching the negative video. This result shows that the horses perceived the content of the videos and suggests an emotional contagion between the actor horse and the subjects. After the videos were projected, the horses took a choice test, facing the positive and negative experimenters in real life. The horses successfully used the interactions seen in the videos to discriminate between the experimenters. They touched the negative experimenter significantly more, which seems counterintuitive but can be interpreted as an appeasement attempt, based on the existing literature. This result suggests that horses can indirectly attribute a valence to a human experimenter by eavesdropping on a previous third-party interaction with a conspecific.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1435-9456 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Trösch2020 Serial 6649  
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Author Henry, S.; Fureix, C.; Rowberry, R.; Bateson, M.; Hausberger, M. url  doi
openurl 
  Title Do horses with poor welfare show 'pessimistic' cognitive biases? Type Journal Article
  Year 2017 Publication The Science of Nature Abbreviated Journal Sci. Nat.  
  Volume 104 Issue 1 Pages 8  
  Keywords  
  Abstract This field study tested the hypothesis that domestic horses living under putatively challenging-to-welfare conditions (for example involving social, spatial, feeding constraints) would present signs of poor welfare and co-occurring pessimistic judgement biases. Our subjects were 34 horses who had been housed for over 3 years in either restricted riding school situations (e.g. kept in single boxes, with limited roughage, ridden by inexperienced riders; N = 25) or under more naturalistic conditions (e.g. access to free-range, kept in stable social groups, leisure riding; N = 9). The horses' welfare was assessed by recording health-related, behavioural and postural indicators. Additionally, after learning a location task to discriminate a bucket containing either edible food ('positive' location) or unpalatable food ('negative' location), the horses were presented with a bucket located near the positive position, near the negative position and halfway between the positive and negative positions to assess their judgement biases. The riding school horses displayed the highest levels of behavioural and health-related problems and a pessimistic judgment bias, whereas the horses living under more naturalistic conditions displayed indications of good welfare and an optimistic bias. Moreover, pessimistic bias data strongly correlated with poor welfare data. This suggests that a lowered mood impacts a non-human species' perception of its environment and highlights cognitive biases as an appropriate tool to assess the impact of chronic living conditions on horse welfare.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1432-1904 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Henry2017 Serial 6665  
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Author Genov, P.W.; Kostava, V. url  doi
openurl 
  Title Untersuchungen zur zahlenmäßigen Stärke des Wolfes und seiner Einwirkung auf die Haustierbestände in Bulgarien Type Journal Article
  Year 1993 Publication Zeitschrift für Jagdwissenschaft Abbreviated Journal  
  Volume 39 Issue 4 Pages 217-223  
  Keywords  
  Abstract Die Untersuchung wurde in der Zeitspanne von 1984 bis 1988 durchgeführt. Es wurden die Protokolle des Staatlichen Versicherungsinstituts benutzt, die Angaben für Raubüberfälle von Wölfen auf Haustiere beinhalten (Tabelle 1). Außerdem wurden Angaben über die während dieser Zeitspanne erlegten Wölfe zusammengefaßt. Die Abschußzahlen lauten: 1984 – 163, 1985 – 147, 1986 – 179, 1987 – 211 und 1988 – 220 Tiere. Die Anzahl der in den einzelnen Gebirgen lebenden Wölfe wurde nach einer Umfrage festgestellt. Für die in Betracht kommenden Gebirge werden folgende Bestandszahlen angenommen: Rhodopen -- 60-80 Individuen, 189 bis 264 km2 pro Tier, Rila- und Piringebirge -- 60-80 Tiere, 109 bis 145 km2 pro Tier, Ossogowo-Belassiza Gebirgssystem -- 40-50 Individuen, 57-70 km2 pro Tier, West- und Mittelbalkan -- 35-38 Wölfe, 200 km2 pro Tier. Dazu kommen noch 10-15 Wölfe im Flußbecken von Beli Lom und etwa 20 Exemplare in Strandscha- und Sakargebirge. Insgesamt lebten in Bulgarien im Jahre 1988 etwa 260-330 Wölfe (Abb. 1).  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1439-0574 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Genov1993 Serial 6686  
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Author Boyce, P.N.; McLoughlin, P.D. url  doi
openurl 
  Title Ecological Interactions Involving Feral Horses and Predators: Review with Implications for Biodiversity Conservation Type Journal Article
  Year 2021 Publication The Journal of Wildlife Management Abbreviated Journal Jour. Wild. Mgmt.  
  Volume n/a Issue n/a Pages  
  Keywords apparent competition; artificial selection; community ecology; conservation; feral horse (Equus ferus caballus); life history; predator-prey dynamics  
  Abstract ABSTRACT For many ecosystems, feral horses are increasingly becoming an important if not dominant component of ungulate biomass and hence influence on community dynamics. Yet we still know little of how horses contribute to key ecological interactions including predator-prey and indirect competitive relationships at a community level. Notably, feral species like horses can exhibit life-history traits that differ from that of native (mainly artiodactyl) herbivore competitors. Artificial selection for traits like increased, early, or extended reproduction that have yet to be reversed by natural selection, coupled with naturally selected differences in anatomy and behavior, in addition to unique management objectives for horses compared to other species, means that the dynamics of feral horse populations are not likely to align with what might be expected of other large herbivores. Unexpected population dynamics and inherent biological asymmetries between native ungulates and feral horses may therefore influence the former via direct competition for shared resources and through enemy-mediated interactions like apparent competition. In several localities feral horses now co-exist with multiple native prey species, some of which are in decline or are species at risk. Compounding risks to native species from direct or indirect competitive exclusion by horses is the unique nature and socio-political context of feral horse management, which tends towards allowing horse populations to be limited largely by natural, density-dependent factors. We summarize the inherent asymmetries between feral horse biology and that of other ungulate prey species with consequences for conservation, focusing on predator-prey and emerging indirect interactions in multi-prey systems, and highlight future directions to address key knowledge gaps in our understanding of how feral horses may now be contributing to the (re)structuring of food webs. Observations of patterns of rapid growth and decline, and associated skews in sex ratios of feral horse populations, indicate a heightened potential for indirect interactions among large ungulate prey species, where there is a prevalence of feral horses as preferred prey, particularly where native prey are declining. In places like western North America, we expect predator-prey interactions involving feral horses to become an increasingly important factor in the conservation of wildlife. This applies not only to economically or culturally important game species but also at-risk species, both predators (e.g., wolves [Canis lupus], grizzly bears [Ursus arctos]) and prey (e.g., woodland caribou [Rangifer tarandus caribou]), necessitating an ecological understanding of the role of horses in natural environments that goes beyond that of population control. ? 2021 The Wildlife Society.  
  Address  
  Corporate Author Thesis  
  Publisher John Wiley & Sons, Ltd Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0022-541x ISBN Medium  
  Area Expedition Conference  
  Notes https://doi.org/10.1002/jwmg.21995 Approved no  
  Call Number Equine Behaviour @ team @ Serial 6642  
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Author Werhahn, H.; Hessel, E.F.; Van den Weghe, H.F.A. url  doi
openurl 
  Title Competition Horses Housed in Single Stalls (II): Effects of Free Exercise on the Behavior in the Stable, the Behavior during Training, and the Degree of Stress Type Journal Article
  Year 2012 Publication Journal of Equine Veterinary Science Abbreviated Journal  
  Volume 32 Issue 1 Pages 22-31  
  Keywords Horse; Turnout; Single stall; Behavior; Hrv  
  Abstract Although housing horses in single stalls limits their natural behavior to a great extent, this housing system is widespread in Germany, especially for competition horses. To improve the welfare of this system, free exercise on pastures or paddocks is deemed suitable, but it is also feared because of injuries and decreased willingness or motivation to perform. In the present study, three treatments were investigated with regard to their effect on the behavior of six competition horses in the stable, behavior during training, and on their degree of stress: daily training without free exercise (no turnout [NT]), solitary turnout for 2 hours after training, and 2-hour turnout in groups of two after training (group turnout). The horses' behavior in the stable was continuously analyzed through video recordings (2 pm to 6 am) on 3 days at the end of each treatment. The degree of stress was evaluated daily by heart rate variability at rest. The behavior during training was evaluated by a questionnaire answered by the riders, and the distance covered during training was measured by global positioning system. When NT was allowed, the horses showed less lying in the stable compared with the treatments with turnout. Heart rate variability measurements resulted in great individual differences, but generally, there was a higher degree of stress shown with the treatment NT according to the following parameters: standard deviation of inter-beat-intervals (SDNN), square root of the mean of the sum of the squares of differences between successive inter-beat-intervals (RMSSD), and ratio between low frequency and high frequency (LF/HF). The willingness to perform was evaluated as being slightly better in the treatments with turnout than in the treatment without turnout.  
  Address  
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  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0737-0806 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6626  
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Author Bottom, S.H. url  openurl
  Title Age-related changes in taste and gustatory response and feeding behaviour in the stabled horse Type Book Chapter
  Year Publication Abbreviated Journal  
  Volume Issue Pages  
  Keywords  
  Abstract There is a paucity of research relating to the anatomy and physiology of gustation and olfaction in the horse. Moreover, whilst an age-related decline in gustation and olfaction has been recorded in humans, no such study has been conducted in the horse. The horse is reliant on gustation and olfaction to make appropriate decisions relating to both short and long term diet selection and thus, any compromise in function, has implications for food intake and potentially welfare. The principal aim of this study was to establish if, and to what extent, taste andgustatory responses are affected by age in the horse. Horses were allocated to the age groups Young (2-5 years), Middle (8-14 years) and Old (16 plus years) for the study of taste (n=18) and to Young (4-6 years), Middle (10-14 years) and Old (16 plus years) for the study of gustation (n=18). Individual taste responses and gustatory responses (taste in the absence of additional olfactory cues) were identified using two-choice preference testing and monadic testing. Statistical analysis was conducted using Minitab 14.0 and behaviour data was analysed using The Observer 5.0 (Noldus, Netherlands). No effect of age on taste response or gustatory response was recorded.  
  Address  
  Corporate Author Thesis Ph.D. thesis  
  Publisher Nottingham Trent University Place of Publication Nottingham, UK Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6636  
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Author Bandini, E.; Tennie C. doi  openurl
  Title Exploring the role of individual learning in animal tool-use Type Journal Article
  Year 2020 Publication PeerJ Abbreviated Journal PeerJ  
  Volume 25 Issue Pages 8:e9877  
  Keywords  
  Abstract The notion that tool-use is unique to humans has long been refuted by the growing number of observations of animals using tools across various contexts. Yet, the mechanisms behind the emergence and sustenance of these tool-use repertoires are still heavily debated. We argue that the current animal behaviour literature is biased towards a social learning approach, in which animal, and in particular primate, tool-use repertoires are thought to require social learning mechanisms (copying variants of social learning are most often invoked). However, concrete evidence for a widespread dependency on social learning is still lacking. On the other hand, a growing body of observational and experimental data demonstrates that various animal species are capable of acquiring the forms of their tool-use behaviours via individual learning, with (non-copying) social learning regulating the frequencies of the behavioural forms within (and, indirectly, between) groups. As a first outline of the extent of the role of individual learning in animal tool-use, a literature review of reports of the spontaneous acquisition of animal tool-use behaviours was carried out across observational and experimental studies. The results of this review suggest that perhaps due to the pervasive focus on social learning in the literature, accounts of the individual learning of tool-use forms by naïve animals may have been largely overlooked, and their importance under-examined.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6659  
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Author Machmoum, M.; Badaoui, B.; Petit, D.; Germot, A.; El Alaoui, M.A.; Boujenane, I.; Piro, M. doi  openurl
  Title Genetic Diversity and Maternal Phylogenetic Relationships among Populations and Strains of Arabian Show Horses Type Journal Article
  Year 2023 Publication Abbreviated Journal  
  Volume Issue Pages  
  Keywords genetic variability; whole D-loop mitochondrial DNA; desert-bred; straight Egyptian; Polish Arabian; traditional Arabian horse classification  
  Abstract Genetic diversity and phylogenetic relationships within the Arabian show horse populations are of particular interest to breeders worldwide. Using the complete mitochondrial DNA D-loop sequence (916 pb), this study aimed (i) to understand the genetic relationship between three populations, the Desert-Bred (DB), a subset of the Kingdom of Saudi Arabia (KSA), United Arab Emirates (UAE) and Bahrain (BAH), the Straight Egyptian (EG) and the Polish bloodline (PL), and (ii) to assess the accuracy of the traditional strain classification system based on maternal lines, as stated by the Bedouin culture. To that end, we collected 211 hair samples from stud farms renowned for breeding Arabian show horses from Nejd KSA, Bahrain, Egypt, Qatar, Morocco, UAE, and Poland. The phylogenetic and network analyses of the whole mitochondrial DNA D-loop sequence highlighted a great genetic diversity among the Arabian horse populations, in which about 75% of variance was assigned to populations and 25% to strains. The discriminant analysis of principal components illustrated a relative distinction between those populations. A clear subdivision between traditional strains was found in PL, in contrast to the situation of DB and EG populations. However, several Polish horse individuals could not be traced back to the Bedouin tribes by historical documentation and were shown to differ genetically from other studied Bedouin strains, hence motivating extended investigations.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title Animals  
  Series Volume 13 Series Issue 12 Edition  
  ISSN 2076-2615 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6709  
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Author Krueger, K.; Trager, L.; Farmer, K.; Byrne, R. doi  openurl
  Title Tool Use in Horses Type Journal Article
  Year 2022 Publication Animals Abbreviated Journal Animals  
  Volume 12 Issue 15 Pages 1876  
  Keywords crowdsourcing; horse; innovation; mule; management; tool use  
  Abstract Tool use has not yet been confirmed in horses, mules or donkeys. As this subject is difficult to research with conventional methods, we used a crowdsourcing approach to gather data. We contacted equid owners and carers and asked them to report and video examples of â��unusualâ�� behaviour via a dedicated website. We also searched YouTube and Facebook for videos of equids showing tool use. From 635 reports, including 1014 behaviours, we found 20 cases of tool use, 13 of which were unambiguous in that it was clear that the behaviour was not trained, caused by reduced welfare, incidental or accidental. We then assessed (a) the effect of management conditions on tool use and (b) whether the animals used tools alone, or socially, involving other equids or humans. We found that management restrictions were associated with corresponding tool use in 12 of the 13 cases (p = 0.01), e.g., equids using sticks to scrape hay within reach when feed was restricted. Furthermore, 8 of the 13 cases involved other equids or humans, such as horses using brushes to groom others. The most frequent tool use was for foraging, with seven examples, tool use for social purposes was seen in four cases, and there was just one case of tool use for escape. There was just one case of tool use for comfort, and in this instance, there were no management restrictions. Equids therefore can develop tool use, especially when management conditions are restricted, but it is a rare occurrence.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title Animals  
  Series Volume 12 Series Issue 15 Edition  
  ISSN 2076-2615 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6695  
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Author Cerasoli, F.; Podaliri Vulpiani, M.; Saluti, G.; Conte, A.; Ricci, M.; Savini, G.; D'Alterio, N. doi  openurl
  Title Assessment of Welfare in Groups of Horses with Different Management, Environments and Activities by Measuring Cortisol in Horsehair, Using Liquid Chromatography Coupled to Hybrid Orbitrap High-Resolution Mass Spectrometry Type
  Year 2022 Publication Animals Abbreviated Journal Animals  
  Volume 12 Issue 14 Pages  
  Keywords cortisol; animal welfare; horse; Lc-Hrms/Ms  
  Abstract Horses have always been animals used for companionship, work, transportation, and performance purposes over the history of humanity; there are different ways of managing horses, but studies on how horse welfare is influenced by different activities and managements are scanty. Understanding how the management, the environment, and the different uses of horses can affect the level of stress and well-being is important not only for people associated with horses. Three groups of horses with different management, environments, and activities were selected: (1) stabled horses ridden frequently, (2) horses that perform public order service under the Italian state police, and (3) free-ranging horses. Cortisol analysis was carried out on horsehair samples using liquid chromatography coupled to hybrid orbitrap high-resolution mass spectrometry (LC-HRMS/MS), a laboratory technique used for the first time to quantify horsehair cortisol. The selection of horses to be included in the three groups was carried out by including only subjects with positive welfare assessment in accordance with the horse welfare assessment protocol (AWIN). These analyses demonstrated that the cortisol levels detected in the horsehair of free-ranging animals were significantly higher compared to those detected in stabled and working horses. These results may have been a consequence of complex environmental, managerial, and behavioral factors, which should be worth further investigation  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title Animals  
  Series Volume 12 Series Issue 14 Edition  
  ISSN 2076-2615 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6674  
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