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Author Cozzi, B.; Povinelli, M.; Ballarin, C.; Granato, A. url  doi
openurl 
  Title The Brain of the Horse: Weight and Cephalization Quotients Type Journal Article
  Year 2014 Publication Brain, Behavior and Evolution Abbreviated Journal Brain Behav Evol  
  Volume 83 Issue 1 Pages (down) 9-16  
  Keywords  
  Abstract The horse is a common domestic animal whose anatomy has been studied since the XVI century. However, a modern neuroanatomy of this species does not exist and most of the data utilized in textbooks and reviews derive from single specimens or relatively old literature. Here, we report information on the brain of Equus caballus obtained by sampling 131 horses, including brain weight (as a whole and subdivided into its constituents), encephalization quotient (EQ), and cerebellar quotient (CQ), and comparisons with what is known about other relevant species. The mean weight of the fresh brains in our experimental series was 598.63 g (SEM ± 7.65), with a mean body weight of 514.12 kg (SEM ± 15.42). The EQ was 0.78 and the CQ was 0.841. The data we obtained indicate that the horse possesses a large, convoluted brain, with a weight similar to that of other hoofed species of like mass. However, the shape of the brain, the noteworthy folding of the neocortex, and the peculiar longitudinal distribution of the gyri suggest an evolutionary specificity at least partially separate from that of the Cetartiodactyla (even-toed mammals and cetaceans) with whom Perissodactyla (odd-toed mammals) are often grouped.  
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  ISSN 0006-8977 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6592  
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Author Mori, E.; Benatti, L.; Lovari, S.; Ferretti, F. url  doi
openurl 
  Title What does the wild boar mean to the wolf? Type Journal Article
  Year 2016 Publication European Journal of Wildlife Research Abbreviated Journal  
  Volume 63 Issue 1 Pages (down) 9  
  Keywords  
  Abstract Generalist predators are expected to shape their diets according to the local availability of prey species. In turn, the extent of consumption of a prey would be influenced by the number of alternative prey species. We have tested this prediction by considering the wild boar and the grey wolf: two widespread species whose distribution ranges overlap largely in Southern Europe, e.g. in Italy. We have reviewed 16 studies from a total of 21 study areas, to assess whether the absolute frequency of occurrence of wild boar in the wolf diet was influenced by (i) occurrence of the other ungulate species in diet and (ii) the number of available ungulate species. Wild boar turned out to be the main prey of the wolf (49% occurrence, on average), followed by roe deer (24%) and livestock (18%). Occurrence of wild boar in the wolf diet decreased with increasing usage of roe deer, livestock, and to a lower extent, chamois and red deer. The number of prey species did not influence the occurrence of wild boar in the wolf diet. The wild boar is a gregarious, noisy and often locally abundant ungulate, thus easily detectable, to a predator. In turn, the extent of predation on this ungulate may not be influenced so much by the availability of other potential prey. Heavy artificial reductions of wild boar numbers, e.g. through numerical control, may concentrate predation by wolves on alternative prey (e.g. roe deer) and/or livestock, thus increasing conflicts with human activities.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1439-0574 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Mori2016 Serial 6689  
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Author Bandini, E.; Tennie C. doi  openurl
  Title Exploring the role of individual learning in animal tool-use Type Journal Article
  Year 2020 Publication PeerJ Abbreviated Journal PeerJ  
  Volume 25 Issue Pages (down) 8:e9877  
  Keywords  
  Abstract The notion that tool-use is unique to humans has long been refuted by the growing number of observations of animals using tools across various contexts. Yet, the mechanisms behind the emergence and sustenance of these tool-use repertoires are still heavily debated. We argue that the current animal behaviour literature is biased towards a social learning approach, in which animal, and in particular primate, tool-use repertoires are thought to require social learning mechanisms (copying variants of social learning are most often invoked). However, concrete evidence for a widespread dependency on social learning is still lacking. On the other hand, a growing body of observational and experimental data demonstrates that various animal species are capable of acquiring the forms of their tool-use behaviours via individual learning, with (non-copying) social learning regulating the frequencies of the behavioural forms within (and, indirectly, between) groups. As a first outline of the extent of the role of individual learning in animal tool-use, a literature review of reports of the spontaneous acquisition of animal tool-use behaviours was carried out across observational and experimental studies. The results of this review suggest that perhaps due to the pervasive focus on social learning in the literature, accounts of the individual learning of tool-use forms by naïve animals may have been largely overlooked, and their importance under-examined.  
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  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6659  
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Author Henry, S.; Fureix, C.; Rowberry, R.; Bateson, M.; Hausberger, M. url  doi
openurl 
  Title Do horses with poor welfare show 'pessimistic' cognitive biases? Type Journal Article
  Year 2017 Publication The Science of Nature Abbreviated Journal Sci. Nat.  
  Volume 104 Issue 1 Pages (down) 8  
  Keywords  
  Abstract This field study tested the hypothesis that domestic horses living under putatively challenging-to-welfare conditions (for example involving social, spatial, feeding constraints) would present signs of poor welfare and co-occurring pessimistic judgement biases. Our subjects were 34 horses who had been housed for over 3 years in either restricted riding school situations (e.g. kept in single boxes, with limited roughage, ridden by inexperienced riders; N = 25) or under more naturalistic conditions (e.g. access to free-range, kept in stable social groups, leisure riding; N = 9). The horses' welfare was assessed by recording health-related, behavioural and postural indicators. Additionally, after learning a location task to discriminate a bucket containing either edible food ('positive' location) or unpalatable food ('negative' location), the horses were presented with a bucket located near the positive position, near the negative position and halfway between the positive and negative positions to assess their judgement biases. The riding school horses displayed the highest levels of behavioural and health-related problems and a pessimistic judgment bias, whereas the horses living under more naturalistic conditions displayed indications of good welfare and an optimistic bias. Moreover, pessimistic bias data strongly correlated with poor welfare data. This suggests that a lowered mood impacts a non-human species' perception of its environment and highlights cognitive biases as an appropriate tool to assess the impact of chronic living conditions on horse welfare.  
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  Series Volume Series Issue Edition  
  ISSN 1432-1904 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Henry2017 Serial 6665  
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Author Lucidi, P.; Bacco, G.; Sticco, M.; Mazzoleni, G.; Benvenuti, M.; Bernabò, N.; Trentini, R. url  doi
openurl 
  Title Assessment of motor laterality in foals and young horses (Equus caballus) through an analysis of derailment at trot Type Journal Article
  Year 2013 Publication Physiology & Behavior Abbreviated Journal Physiol. Behav.  
  Volume 109 Issue Pages (down) 8-13  
  Keywords Horse; Laterality; Longeing; Sidedness  
  Abstract The conflicting results regarding the study of motor laterality in horses may indicate that there does not exist a proper method to assess the degree and the direction of motor bias in these animals. Unfortunately, even less is known about the development of laterality in horses, and to what extent early manipulations can still exert their effects in adulthood. We propose a new method that can be easily applied at a very early age thus avoiding testing adult horses eventually biased by human handling and/or training. Forty-six horses (29 nine-month-old foals and 17 two-year old horses) were handled since birth bilaterally and housed in groups in wide areas. At the time of the analysis, in order to minimize environmental and sensorial disturbances, each horse was tested in a round pen individually or as dyad mother-foal. The ability/inability to properly execute a circle at trot was then recorded, assuming the direction of derailment, i.e. the cutting of the circle, as an indicator of motor bias. From the results of the study it is arguable that motor laterality in horses is acquired over time: in fact foals tested while their mothers were being subjected to longeing showed a higher percentage of ambidextrous animals, while two-year-old horses appeared biased toward the right (p<0.05). Results are discussed in the light of the scientific knowledge about equine biomechanics, taking into account horses' locomotion that leads to the advancement of the body mass through the activation of a kinetic chain that originates from the hindquarters.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0031-9384 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6666  
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Author Bernauer, K.; Kollross, H.; Schuetz, A.; Farmer, K.; Krueger, K. url  doi
openurl 
  Title How do horses (Equus caballus) learn from observing human action? Type Journal Article
  Year 2020 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume 23 Issue Pages (down) 1-9  
  Keywords  
  Abstract A previous study demonstrated that horses can learn socially from observing humans, but could not draw any conclusions about the social learning mechanisms. Here we develop this by showing horses four different human action sequences as demonstrations of how to press a button to open a feed box. We tested 68 horses aged between 3 and 12 years. 63 horses passed the habituation phase and were assigned either to the group Hand Demo (N = 13) for which a kneeling person used a hand to press the button, Head Demo (N = 13) for which a kneeling person used the head, Mixed Demo (N = 12) for which a squatting person used both head and hand, Foot Demo (N = 12) in which a standing person used a foot, or No Demo (N = 13) in which horses did not receive a demonstration. 44 horses reached the learning criterion of opening the feeder twenty times consecutively, 40 of these were 75% of the Demo group horses and four horses were 31% of the No Demo group horses. Horses not reaching the learning criterion approached the human experimenters more often than those who did. Significantly more horses used their head to press the button no matter which demonstration they received. However, in the Foot Demo group four horses consistently preferred to use a hoof and two switched between hoof and head use. After the Mixed Demo the horses' actions were more diverse. The results indicate that only a few horses copy behaviours when learning socially from humans. A few may learn through observational conditioning, as some appeared to adapt to demonstrated actions in the course of reaching the learning criterion. Most horses learn socially through enhancement, using humans to learn where, and which aspect of a mechanism has to be manipulated, and by applying individual trial and error learning to reach their goal.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1435-9456 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Bernauer2019 Serial 6590  
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Author Freitas, J.; Lagos, L.; Álvares, F. openurl 
  Title Horses as prey of wolves. Type Journal Article
  Year 2021 Publication Carnivore Damage Preventionnews Abbreviated Journal CDPnews  
  Volume 23 Issue Pages (down) 1-9  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6677  
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Author Lema, F.J.; Ribeiro, S.; Palacios, V. openurl 
  Title Observations of wolves hunting fee-ranging horses in Iberia. Type Journal Article
  Year 2022 Publication Carnivore Damage Prevention News Abbreviated Journal CDPNews  
  Volume 24 Issue Pages (down) 1-9  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6681  
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Author Hartmann, E.; Christensen, J.W.; McGreevy, P.D. url  doi
openurl 
  Title Dominance and Leadership: Useful Concepts in Human-Horse Interactions? Type Journal Article
  Year 2017 Publication Journal of Equine Veterinary Science Abbreviated Journal Proceedings of the 2017 Equine Science Symposium  
  Volume 52 Issue Pages (down) 1-9  
  Keywords Horse; Social order; Dominance hierarchy; Aggression; Injury; Learning; Training  
  Abstract Dominance hierarchies in horses primarily influence priority access to limited resources of any kind, resulting in predictable contest outcomes that potentially minimize aggressive encounters and associated risk of injury. Levels of aggression in group-kept horses under domestic conditions have been reported to be higher than in their feral counterparts but can often be attributed to suboptimal management. Horse owners often express concerns about the risk of injuries occurring in group-kept horses, but these concerns have not been substantiated by empirical investigations. What has not yet been sufficiently addressed are human safety aspects related to approaching and handling group-kept horses. Given horse's natural tendency to synchronize activity to promote group cohesion, questions remain about how group dynamics influence human-horse interactions. Group dynamics influence a variety of management scenarios, ranging from taking a horse out of its social group to the prospect of humans mimicking the horse's social system by taking a putative leadership role and seeking after an alpha position in the dominance hierarchy to achieve compliance. Yet, there is considerable debate about whether the roles horses attain in their social group are of any relevance in their reactions to humans. This article reviews the empirical data on social dynamics in horses, focusing on dominance and leadership theories and the merits of incorporating those concepts into the human-horse context. This will provide a constructive framework for informed debate and valuable guidance for owners managing group-kept horses and for optimizing human-horse interactions.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0737-0806 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6712  
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Author Edwards, K.L.; Bansiddhi, P.; Paris, S.; Galloway, M.; Brown, J.L. url  doi
openurl 
  Title The development of an immunoassay to measure immunoglobulin A in Asian elephant feces, saliva, urine and serum as a potential biomarker of well-being Type Journal Article
  Year 2019 Publication Conservation Physiology Abbreviated Journal conphys  
  Volume 7 Issue 1 Pages (down)  
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  Abstract Additional measures of well-being would be beneficial to the management of a variety of species in human care, including elephants. Immunoglobulin A (IgA) is an immune protein associated with pathogen defense, which has been demonstrated to decrease during times of stress, and increase in response to positive stimuli. This paper describes the development and validation of an enzyme immunoassay (EIA) for the quantification of Asian elephant (Elephas maximus) IgA in feces, saliva, urine, and serum. Samples were collected weekly from four females for 6 months to assess IgA and glucocorticoid (GC) concentrations, establish relationships between these two biomarkers, and determine variability in IgA within and between individuals, and across sample types. IgA was quantified in all four sample types, although urinary concentrations were low and sometimes undetectable in individual samples. Concentrations were highly variable within and between individuals, with fecal, salivary and serum IgA, and fecal, salivary and urinary GCs all differing significantly across individuals. Contrary to previous findings, IgA and GC were generally not correlated. Serum IgA was less variable within individuals, with the exception of one female that experienced a brief illness during the study. However, marked inter-individual differences were still apparent. When data from all individuals were combined, fecal IgA was significantly predicted by salivary and urinary IgA; however, this relationship did not hold when individuals were analyzed separately. Analysis of a fifth female that exhibited a more severe systemic illness demonstrated clear increases in fecal IgA and GC, suggesting these may also be useful health biomarkers. Further investigation is needed to determine what sample type is most reflective of biological state in elephants, and how IgA concentrations are associated with health and positive and negative welfare states. Based on observed variability, a longitudinal approach likely will be necessary to use IgA as a measure of well-being.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 2051-1434 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6591  
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