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Author Krueger, K.; Esch, L.; Byrne, R. url  doi
openurl 
  Title Animal behaviour in a human world: A crowdsourcing study on horses that open door and gate mechanisms Type Journal Article
  Year 2019 Publication Plos One Abbreviated Journal Plos One  
  Volume 14 Issue (down) 6 Pages e0218954  
  Keywords  
  Abstract Anecdotal reports of horses opening fastened doors and gates are an intriguing way of exploring the possible scope of horses' problem-solving capacities. The species' natural environment has no analogues of the mechanisms involved. Scientific studies on the topic are missing, because the rate of occurrence is too low for exploration under controlled conditions. Therefore, we compiled from lay persons case reports of horses opening closed doors and gates. Additionally, we collected video documentations at the internet platform YouTube, taking care to select raw data footage of unedited, clearly described and clearly visible cases of animals with no distinct signs of training or reduced welfare. The data included individuals opening 513 doors or gates on hinges, 49 sliding doors, and 33 barred doors and gateways; mechanisms included 260 cases of horizontal and 155 vertical bars, 43 twist locks, 42 door handles, 34 electric fence handles, 40 carabiners, and 2 locks with keys. Opening was usually for escape, but also for access to food or stable-mates, or out of curiosity or playfulness. While 56 percent of the horses opened a single mechanism at one location, 44 percent opened several types of mechanism (median = 2, min. = 1, max. = 5) at different locations (median = 2, min. = 1, max. = 4). The more complex the mechanism was, the more movements were applied, varying from median 2 for door handles to 10 for carabiners. Mechanisms requiring head- or lip-twisting needed more movements, with significant variation between individuals. 74 horses reported in the questionnaire had options for observing the behaviour in stable mates, 183 did not, which indicates that the latter learned to open doors and gates either individually or from observing humans. Experience favours opening efficiency; subjects which opened several door types applied fewer movements per lock than horses which opened only one door type. We failed to identify a level of complexity of door-fastening mechanism that was beyond the learning capacity of the horse to open. Thus, all devices in frequent use, even carabiners and electric fence handles, are potentially vulnerable to opening by horses, something which needs to be considered in relation to keeping horses safely.  
  Address  
  Corporate Author Thesis  
  Publisher Public Library of Science Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6580  
Permanent link to this record
 

 
Author López-Bao, J.V.; Sazatornil, V.; Llaneza, L.; Rodríguez, A. url  doi
openurl 
  Title Indirect Effects on Heathland Conservation and Wolf Persistence of Contradictory Policies that Threaten Traditional Free-Ranging Horse Husbandry Type Journal Article
  Year 2013 Publication Conservation Letters Abbreviated Journal Conservation Letters  
  Volume 6 Issue (down) 6 Pages 448-455  
  Keywords Farmland biodiversity; heathlands; integration of environmental policies; management of livestock carcasses; traditional land uses; wolf conservation  
  Abstract Abstract Conservation agencies within the European Union promote the restoration of traditional land uses as a cost-effective way to preserve biodiversity outside reserves. Although the European Union pursues the integration of the environment into strategic decision-making, it also dictates sectoral policies that may damage farmland biodiversity. We illustrate this point by outlining the socioeconomic factors that allow the persistence of traditional free-ranging horse husbandry in Galicia, northwestern Spain. Free-ranging Galician mountain ponies provide ecological and socioeconomic services including the prevention of forest fires, the maintenance of heathlands and wolves, and the attenuation of wolf-human conflicts. This traditional livestock system may have persisted because it entails negligible costs for farmers. Wolf predation upon Galician mountain ponies does not threaten farmer's economies and seems to be tolerated better than attacks to more valuable stock. Recently, European Union's regulations on animal welfare, carcass management, or meat production put new economic and administrative burdens on farmers, make free-ranging horse rearing economically unsustainable, and incentivize its abandonment. The aim of the European Union to integrate environmental policies may be successful to preserve farmland biodiversity only through careful anticipation of the side effects of apparently unrelated regulations on the fragile equilibrium that sustain traditional land uses.  
  Address  
  Corporate Author Thesis  
  Publisher John Wiley & Sons, Ltd Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1755-263x ISBN Medium  
  Area Expedition Conference  
  Notes doi: 10.1111/conl.12014 Approved no  
  Call Number Equine Behaviour @ team @ Serial 6622  
Permanent link to this record
 

 
Author Krueger, K.; Esch, L.; Farmer, K.; Marr, I. doi  openurl
  Title Basic Needs in Horses?--A Literature Review Type Magazine Article
  Year 2021 Publication Animals Abbreviated Journal Animals  
  Volume 11 Issue (down) 6 Pages 1798  
  Keywords abnormal behaviour; active responses; horse; movement; passive responses; roughage; stress; social contact  
  Abstract Every animal species has particular environmental requirements that are essential for its welfare, and when these so-called “basic needs” are not fulfilled, the animals suffer. The basic needs of horses have been claimed to be social contact, social companionship, free movement and access to roughage. To assess whether horses suffer when one or more of the four proposed basic needs are restricted, we examined several studies (n = 38) that reported behavioural and physiological reactions to these restrictions. We assigned the studies according to the four types of responses investigated: (a) Stress, (b) Active, (c) Passive, and (d) Abnormal Behaviour. Furthermore, the number of studies indicating that horses reacted to the restrictions were compared with the number of studies reporting no reaction. The limited number of studies available on single management restrictions did not allow conclusions to be drawn on the effect of each restriction separately, especially in the case of social companionship. However, when combinations of social contact, free movement and access to roughage were restricted, many of the horses had developed responses consistent with suffering. Passive Responses, indicating acute suffering, and Abnormal Behaviour, indicating suffering currently or at some time in the past, were especially clearly demonstrated. This provides further evidence of the usefulness of assessing behavioural parameters in combination with physiological measurements when evaluating horse welfare. This meta-analysis of the literature confirms that it is justified to claim that social contact, free movement and access to roughage are basic needs in horses.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title Animals  
  Series Volume 11 Series Issue 6 Edition  
  ISSN 2076-2615 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6645  
Permanent link to this record
 

 
Author López-Bao, J.V.; Sazatornil, V.; Llaneza, L.; Rodríguez, A. url  doi
openurl 
  Title Indirect Effects on Heathland Conservation and Wolf Persistence of Contradictory Policies that Threaten Traditional Free-Ranging Horse Husbandry Type Journal Article
  Year 2013 Publication Conservation Letters Abbreviated Journal Conservation Letters  
  Volume 6 Issue (down) 6 Pages 448-455  
  Keywords Farmland biodiversity; heathlands; integration of environmental policies; management of livestock carcasses; traditional land uses; wolf conservation  
  Abstract Abstract Conservation agencies within the European Union promote the restoration of traditional land uses as a cost-effective way to preserve biodiversity outside reserves. Although the European Union pursues the integration of the environment into strategic decision-making, it also dictates sectoral policies that may damage farmland biodiversity. We illustrate this point by outlining the socioeconomic factors that allow the persistence of traditional free-ranging horse husbandry in Galicia, northwestern Spain. Free-ranging Galician mountain ponies provide ecological and socioeconomic services including the prevention of forest fires, the maintenance of heathlands and wolves, and the attenuation of wolf-human conflicts. This traditional livestock system may have persisted because it entails negligible costs for farmers. Wolf predation upon Galician mountain ponies does not threaten farmer's economies and seems to be tolerated better than attacks to more valuable stock. Recently, European Union's regulations on animal welfare, carcass management, or meat production put new economic and administrative burdens on farmers, make free-ranging horse rearing economically unsustainable, and incentivize its abandonment. The aim of the European Union to integrate environmental policies may be successful to preserve farmland biodiversity only through careful anticipation of the side effects of apparently unrelated regulations on the fragile equilibrium that sustain traditional land uses.  
  Address  
  Corporate Author Thesis  
  Publisher John Wiley & Sons, Ltd Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1755-263x ISBN Medium  
  Area Expedition Conference  
  Notes https://doi.org/10.1111/conl.12014 Approved no  
  Call Number Equine Behaviour @ team @ Serial 6685  
Permanent link to this record
 

 
Author Outram, A.K.; Stear, N.A.; Bendrey, R.; Olsen, S.; Kasparov, A.; Zaibert, V.; Thorpe, N.; Evershed, R.P. url  doi
openurl 
  Title The Earliest Horse Harnessing and Milking Type Journal Article
  Year 2009 Publication Science Abbreviated Journal  
  Volume 323 Issue (down) 5919 Pages 1332-1335  
  Keywords  
  Abstract Horse domestication revolutionized transport, communications, and warfare in prehistory, yet the identification of early domestication processes has been problematic. Here, we present three independent lines of evidence demonstrating domestication in the Eneolithic Botai Culture of Kazakhstan, dating to about 3500 B.C.E. Metrical analysis of horse metacarpals shows that Botai horses resemble Bronze Age domestic horses rather than Paleolithic wild horses from the same region. Pathological characteristics indicate that some Botai horses were bridled, perhaps ridden. Organic residue analysis, using δ13C and δD values of fatty acids, reveals processing of mare's milk and carcass products in ceramics, indicating a developed domestic economy encompassing secondary products.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6620  
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Author Burden, F.; Thiemann, A. url  doi
openurl 
  Title Donkeys Are Different Type Journal Article
  Year 2015 Publication Journal of Equine Veterinary Science Abbreviated Journal Proceedings of the 2015 Equine Science Society Symposium  
  Volume 35 Issue (down) 5 Pages 376-382  
  Keywords Donkey; Ass; Equid; Mule  
  Abstract As a unique species of equine, the donkey has certain specific variations from the horse. This review highlights the origins of the donkey and how this impacts on its behavior, physiology, and propensity to disease. The donkey is less of a flight animal and has been used by humans for pack and draught work, in areas where their ability to survive poorer diets, and transboundary disease while masking overt signs of pain and distress has made them indispensable to human livelihoods. When living as a companion animal, however, the donkey easily accumulates adipose tissue, and this may create a metabolically compromised individual prone to diseases of excess such as laminitis and hyperlipemia. They show anatomic variations from the horse especially in the hoof, upper airway, and their conformation. Variations in physiology lead to differences in the metabolism and distribution of many drugs. With over 44 million donkeys worldwide, it is important that veterinarians have the ability to understand and treat this equid effectively.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0737-0806 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6541  
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Author Dunbar, R.I.M. url  doi
openurl 
  Title The social brain hypothesis and its implications for social evolution Type Journal Article
  Year 2009 Publication Annals of Human Biology Abbreviated Journal Annals of Human Biology  
  Volume 36 Issue (down) 5 Pages 562-572  
  Keywords  
  Abstract The social brain hypothesis was proposed as an explanation for the fact that primates have unusually large brains for body size compared to all other vertebrates: Primates evolved large brains to manage their unusually complex social systems. Although this proposal has been generalized to all vertebrate taxa as an explanation for brain evolution, recent analyses suggest that the social brain hypothesis takes a very different form in other mammals and birds than it does in anthropoid primates. In primates, there is a quantitative relationship between brain size and social group size (group size is a monotonic function of brain size), presumably because the cognitive demands of sociality place a constraint on the number of individuals that can be maintained in a coherent group. In other mammals and birds, the relationship is a qualitative one: Large brains are associated with categorical differences in mating system, with species that have pairbonded mating systems having the largest brains. It seems that anthropoid primates may have generalized the bonding processes that characterize monogamous pairbonds to other non-reproductive relationships (?friendships?), thereby giving rise to the quantitative relationship between group size and brain size that we find in this taxon. This raises issues about why bonded relationships are cognitively so demanding (and, indeed, raises questions about what a bonded relationship actually is), and when and why primates undertook this change in social style.  
  Address  
  Corporate Author Thesis  
  Publisher Taylor & Francis Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0301-4460 ISBN Medium  
  Area Expedition Conference  
  Notes doi: 10.1080/03014460902960289 Approved no  
  Call Number Equine Behaviour @ team @ Serial 6546  
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Author Esch, L.; Wöhr, C.; Erhard, M.; Krueger, K. doi  openurl
  Title Horses� (Equus Caballus) Laterality, Stress Hormones, and Task Related Behavior in Innovative Problem-Solving Type Journal Article
  Year 2019 Publication Animals Abbreviated Journal Animals  
  Volume 9 Issue (down) 5 Pages 265  
  Keywords innovative behavior; brain lateralization; glucocorticoid metabolites; behavioral traits; equine cognition  
  Abstract Domesticated horses are constantly confronted with novel tasks. A recent study on anecdotal data indicates that some are innovative in dealing with such tasks. However, innovative behavior in horses has not previously been investigated under experimental conditions. In this study, we investigated whether 16 horses found an innovative solution when confronted with a novel feeder. Moreover, we investigated whether innovative behavior in horses may be affected by individual aspects such as: age, sex, size, motor and sensory laterality, fecal stress hormone concentrations (GCMs), and task-related behavior. Our study revealed evidence for 25% of the horses being capable of innovative problem solving for operating a novel feeder. Innovative horses of the present study were active, tenacious, and may be considered to have a higher inhibitory control, which was revealed by their task related behavior. Furthermore, they appeared to be emotional, reflected by high baseline GCM concentrations and a left sensory and motor laterality. These findings may contribute to the understanding of horses� cognitive capacities to deal with their environment and calls for enriched environments in sports and leisure horse management.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Esch2019 Serial 6570  
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Author openurl 
  Title Type Journal Article
  Year Publication Abbreviated Journal  
  Volume 9 Issue (down) 5 Pages 265  
  Keywords  
  Abstract  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 2076-2615 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ ref2 Serial 6571  
Permanent link to this record
 

 
Author Hendriksen, P.; Elmgreen, K.; Ladewig, J. url  doi
openurl 
  Title Trailer-loading of horses: Is there a difference between positive and negative reinforcement concerning effectiveness and stress-related signs? Type Journal Article
  Year 2011 Publication Journal of Veterinary Behavior Abbreviated Journal J. Vet. Behav.  
  Volume 6 Issue (down) 5 Pages 261-266  
  Keywords negative reinforcement; positive reinforcement; horse; behavior; heart rate; trailer-loading  
  Abstract The traditional way to train horses is by the application of negative reinforcement (NR). In the past few years, however, the use of positive reinforcement (PR) has become more common. To evaluate the effectiveness and the possible stressor effect of the 2 training methods, 12 horses showing severe trailer-loading problems were selected and exposed to trailer-loading. They were randomly assigned to one of the 2 methods. NR consisted of various degrees of pressure (lead rope pulling, whip tapping). Pressure was removed as soon as the horse complied. PR horses were exposed to clicker training and taught to follow a target into the trailer. Heart rate (HR) was recorded every 5 seconds and behavior denoting discomfort was observed using one-zero sampling with 10 seconds sampling intervals. Training was completed when the horse could enter the trailer upon a signal, or was terminated after a maximum of 15 sessions. Of the 12 horses, 10 reached the criterion within the 15 sessions. One horse was eliminated from the study because of illness and 1 PR horse failed to enter the trailer. A Mann-Whitney U-test indicated that the horses trained with NR displayed significantly more discomfort behavior per training session than horses trained with PR (NR: 13.26 ± 3.25; PR: 3.17 ± 8.93, P < 0.0001) and that horses in the PR group spent less time (second) per session to complete the training criterion (NR: 672.9 ± 247.12; PR: 539.81 ± 166.37, P < 0.01). A Mann-Whitney U-test showed that no difference existed in mean HR (bpm) between the 2 groups (NR: 53.06 ± 11.73 bpm; PR: 55.54 ± 6.7 bpm, P > 0.05), but a Wilcoxon test showed a difference in the PR group between the baseline of HR and mean HR obtained during training sessions (baseline PR: 43 ± 8.83 bpm; PR: 55.54 ± 6.7 bpm, P < 0.05). In conclusion, the PR group provided the fastest training solution and expressed less stress response. Thus, the PR procedure could provide a preferable training solution when training horses in potentially stressing situations.  
  Address  
  Corporate Author Thesis  
  Publisher Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1558-7878 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6598  
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