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Author Bernauer, K.; Kollross, H.; Schuetz, A.; Farmer, K.; Krueger, K.
Title How do horses (Equus caballus) learn from observing human action? Type Journal Article
Year 2020 Publication Animal Cognition Abbreviated Journal Anim. Cogn.
Volume 23 Issue Pages 1-9
Keywords
Abstract A previous study demonstrated that horses can learn socially from observing humans, but could not draw any conclusions about the social learning mechanisms. Here we develop this by showing horses four different human action sequences as demonstrations of how to press a button to open a feed box. We tested 68 horses aged between 3 and 12 years. 63 horses passed the habituation phase and were assigned either to the group Hand Demo (N = 13) for which a kneeling person used a hand to press the button, Head Demo (N = 13) for which a kneeling person used the head, Mixed Demo (N = 12) for which a squatting person used both head and hand, Foot Demo (N = 12) in which a standing person used a foot, or No Demo (N = 13) in which horses did not receive a demonstration. 44 horses reached the learning criterion of opening the feeder twenty times consecutively, 40 of these were 75% of the Demo group horses and four horses were 31% of the No Demo group horses. Horses not reaching the learning criterion approached the human experimenters more often than those who did. Significantly more horses used their head to press the button no matter which demonstration they received. However, in the Foot Demo group four horses consistently preferred to use a hoof and two switched between hoof and head use. After the Mixed Demo the horses' actions were more diverse. The results indicate that only a few horses copy behaviours when learning socially from humans. A few may learn through observational conditioning, as some appeared to adapt to demonstrated actions in the course of reaching the learning criterion. Most horses learn socially through enhancement, using humans to learn where, and which aspect of a mechanism has to be manipulated, and by applying individual trial and error learning to reach their goal.
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Corporate Author Thesis
Publisher (up) Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 1435-9456 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Bernauer2019 Serial 6590
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Author Edwards, K.L.; Bansiddhi, P.; Paris, S.; Galloway, M.; Brown, J.L.
Title The development of an immunoassay to measure immunoglobulin A in Asian elephant feces, saliva, urine and serum as a potential biomarker of well-being Type Journal Article
Year 2019 Publication Conservation Physiology Abbreviated Journal conphys
Volume 7 Issue 1 Pages
Keywords
Abstract Additional measures of well-being would be beneficial to the management of a variety of species in human care, including elephants. Immunoglobulin A (IgA) is an immune protein associated with pathogen defense, which has been demonstrated to decrease during times of stress, and increase in response to positive stimuli. This paper describes the development and validation of an enzyme immunoassay (EIA) for the quantification of Asian elephant (Elephas maximus) IgA in feces, saliva, urine, and serum. Samples were collected weekly from four females for 6 months to assess IgA and glucocorticoid (GC) concentrations, establish relationships between these two biomarkers, and determine variability in IgA within and between individuals, and across sample types. IgA was quantified in all four sample types, although urinary concentrations were low and sometimes undetectable in individual samples. Concentrations were highly variable within and between individuals, with fecal, salivary and serum IgA, and fecal, salivary and urinary GCs all differing significantly across individuals. Contrary to previous findings, IgA and GC were generally not correlated. Serum IgA was less variable within individuals, with the exception of one female that experienced a brief illness during the study. However, marked inter-individual differences were still apparent. When data from all individuals were combined, fecal IgA was significantly predicted by salivary and urinary IgA; however, this relationship did not hold when individuals were analyzed separately. Analysis of a fifth female that exhibited a more severe systemic illness demonstrated clear increases in fecal IgA and GC, suggesting these may also be useful health biomarkers. Further investigation is needed to determine what sample type is most reflective of biological state in elephants, and how IgA concentrations are associated with health and positive and negative welfare states. Based on observed variability, a longitudinal approach likely will be necessary to use IgA as a measure of well-being.
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Corporate Author Thesis
Publisher (up) Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 2051-1434 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6591
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Author Cozzi, B.; Povinelli, M.; Ballarin, C.; Granato, A.
Title The Brain of the Horse: Weight and Cephalization Quotients Type Journal Article
Year 2014 Publication Brain, Behavior and Evolution Abbreviated Journal Brain Behav Evol
Volume 83 Issue 1 Pages 9-16
Keywords
Abstract The horse is a common domestic animal whose anatomy has been studied since the XVI century. However, a modern neuroanatomy of this species does not exist and most of the data utilized in textbooks and reviews derive from single specimens or relatively old literature. Here, we report information on the brain of Equus caballus obtained by sampling 131 horses, including brain weight (as a whole and subdivided into its constituents), encephalization quotient (EQ), and cerebellar quotient (CQ), and comparisons with what is known about other relevant species. The mean weight of the fresh brains in our experimental series was 598.63 g (SEM ± 7.65), with a mean body weight of 514.12 kg (SEM ± 15.42). The EQ was 0.78 and the CQ was 0.841. The data we obtained indicate that the horse possesses a large, convoluted brain, with a weight similar to that of other hoofed species of like mass. However, the shape of the brain, the noteworthy folding of the neocortex, and the peculiar longitudinal distribution of the gyri suggest an evolutionary specificity at least partially separate from that of the Cetartiodactyla (even-toed mammals and cetaceans) with whom Perissodactyla (odd-toed mammals) are often grouped.
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Publisher (up) Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0006-8977 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6592
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Author de Oliveira, K.; Soutello, R.V.G.; da Fonseca, R.; Costa, C.; de L. Meirelles, P.R.; Fachiolli, D.F.; Clayton, H.M.
Title Gymnastic Training and Dynamic Mobilization Exercises Improve Stride Quality and Increase Epaxial Muscle Size in Therapy Horses Type Journal Article
Year 2015 Publication Journal of Equine Veterinary Science Abbreviated Journal
Volume 35 Issue 11 Pages 888-893
Keywords Equine; Hippotherapy; Therapeutic exercise; Dynamic mobilization exercise; Physical training; Three-dimensional movement
Abstract The objective was to evaluate the efficacy of gymnastic training (GYM) and dynamic mobilization exercises (DMEs) on stride length (SL) and epaxial muscle size in therapy horses. Nine cross-bred hippotherapy horses that performed three, 25-minute therapeutic riding sessions per week throughout the study period were randomly assigned to three experimental groups: a control group in which the horses were sedentary with no additional physical activity; a group that performed DMEs; and a group that performed both DMEs and additional GYM including pelvic tilting, backing, turning in small circles, and walking over a raised rail to strengthen the abdominal and pelvic stabilizer muscles. The exercises were performed 3 days per week for 3 months, with evaluations at the start and end of the study. Stride quality was assessed by measuring SL and tracking distance (TD). Epaxial muscle size was monitored by ultrasonographic measurement of m. longissimus dorsi (LD) thickness and m. multifidi (MM) cross-sectional area. Paired t tests were used to compare within groups across time, and between groups were detected using analysis of variance with Tukey post hoc test. When walking at 1.3 m/s, SL and TD at walk increased significantly (P < .05) in horses subjected to GYM. Thickness of LD did not change in any group, but cross-sectional area of MM increased significantly by 3.55 cm2 (DME) and 3.78 cm2 (GYM). It was concluded that GYM training improved stride quality and DME-stimulated MM hypertrophy which has been shown to improve intervertebral joint stability in other species.
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Corporate Author Thesis
Publisher (up) Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0737-0806 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6593
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Author Krueger, K.; Marr, I.; Dobler, A.; Palme, R.
Title Preservation of fecal glucocorticoid metabolites and immunoglobulin A through silica gel drying for field studies in horses Type Journal Article
Year 2019 Publication Conservation Physiology Abbreviated Journal conphys
Volume 7 Issue 1 Pages
Keywords
Abstract Non-invasive methods enable stress evaluation through measuring fecal glucocorticoid metabolites (FGMs), and immunoglobulin A (IgA) in the feces avoiding stressful blood drawing or stressful restraining of animals in the field. However, FGMs and IgA are mostly analysed in freshly frozen samples, which is difficult when fresh samples cannot be frozen immediately or frozen samples cannot be stored or transported. Good results were also derived from air-dried fecal samples, which are hampered by unstable air humidity in the field. These difficulties may be overcome, when drying of samples could be induced with colorless silica gel (SiO2) granules in a secure set-up, such as an air tight tube. We determined the speed of drying 1.5 g of a fresh fecal sample from six horses on air and on silica gel. Furthermore, FGMs and IgA were analysed in differently stored subsamples from 12 horses: in frozen fecal samples, in air- or silica gel-dried samples stored for 1 day and for 7 days, and in wet fecal samples kept in a tube at room temperature for 7 days. FGM levels remained stable in feces dried on air or on silica gel for 7 days, whereas IgA quantities showed a significant loss. Under field conditions, when freezing or transporting the frozen samples is not possible and humidity hampers air drying, drying samples on silica gel in air tight tubes appears to be very helpful and reliable for analysing FGMs.
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Corporate Author Thesis
Publisher (up) Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 2051-1434 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6594
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Author Osman, F.; Zeitler-Feicht, M.; Fink, G. W.; Arnhard, S.; Krüger, K.
Title Überprüfung der lichten Weiten von Gitterstäben in der Pferdehaltung [Analysing demands for the clear widths of lattice bars in horse husbandry] Type Journal Article
Year 2020 Publication Abbreviated Journal Landtechnik [Agricultural Engineering]
Volume 75 Issue 1 Pages 24-33
Keywords
Abstract An Gitterstäben, die in der Pferdehaltung an verschiedenen Stellen verwendet werden, können sich Pferde verletzen, wenn sie ihre Köpfe oder ihre Hufe hindurchstecken und nicht zurückziehen können. Um das Verletzungsrisiko zu reduzieren sind lichte Weiten und Materialstärken von Gitterstäben so zu wählen, dass Pferdeköpfe und -hufe entweder nicht zwischen den Freiräumen hindurchpassen oder aber gefahrlos wieder zurückgezogen werden können. Die bisherige Forschung liefert jedoch keine belastbaren Aussagen zu Stababständen (senkrecht und waagerecht), die für Pferde ungefährlich sind. Die in der Praxis verwendeten und in der Literatur empfohlenen Gitterstababstände beruhen auf Erfahrungswerten und technischen Materialeigenschaften. In der vorliegenden Untersuchung wurden Pferdeköpfe und -hufe von insgesamt 480 Pferden (233 Stuten, 204 Wallache und 43 Hengste) von 23 verschiedenen Rassen vermessen, um auf Grundlage der Anatomie der Pferde Aussagen über die Eignung von marktüblichen Stababständen in der Praxis treffen zu können. Es stellte sich heraus, dass bei senkrechten Gitterstäben eine lichte Weite von nicht mehr als 5 cm für alle Pferde ab einem Stockmaß von 110 cm und einem Alter von zwei Jahren als sicher bezeichnet werden kann. Bei waagerechten Gitterstäben erwies sich eine lichte Weite von genau 17 cm als sicher. Dies gilt für alle Pferde ab einem Alter von zwei Jahren oder ab einem Stockmaß von 148 cm.

Kritisch sind die lichten Weiten von Panels zu beurteilen. Hier zeigte sich, dass die handelsüblichen Abstände der Gitterstäbe für die meisten Pferde eine erhebliche Gefahr darstellen. Wenn die Pferde beispielsweise versuchen außerhalb der Panels zu fressen und dabei ihren Kopf durch die Gitterstäbe stecken, kann es leicht passieren, dass sie sich mit dem Kopf zwischen den Gitterstäben verklemmen.

[At bars, used in various places in horse husbandry, horses can hurt themselves when retracting their heads or hooves after pushing them through the interspaces. In order to reduce the risk of injury, the clear widths and material thicknesses of bars should be chosen so that horse heads and hooves either cannot pass between the spaces or can be retracted safely. However, research to date has not provided any reliable information on bar width (vertical and horizontal) that is safe for horses. Grid bar width used in practice and recommended in the literature is based on empirical values and technical material properties. In this study, heads and hooves of 480 horses (233 mares, 204 geldings and 43 stallions) of 23 breeds were measured for making statements about the suitability of standard bar width, when considering the anatomy of the horse. It turned out that for vertical bars, an interspace of no more than five centimetres can be considered to be safe for all horses of a height of 110 centimetres and an age of two years and more. With horizontal lattice bars, a clear width of exactly 17 centimetres proved to be safe. This applies to all horses of a height of 148 centimetres and an age of two years or more. The clear widths of panels must be considered critical for horse welfare. When horses, for example, try to eat outside the panels and put their head through the bars, they may get stuck.]
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Publisher (up) Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6595
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Author Young, R.J.
Title Environmental Enrichment for Captive Animals Type Book Whole
Year 2003 Publication Wiley Interdiscip Rev Cogn Sci Abbreviated Journal
Volume Issue Pages
Keywords
Abstract Environmental enrichment is a simple and effective means of improving animal welfare in any species – companion, farm, laboratory and zoo. For many years, it has been a popular area of research, and has attracted the attention and concerns of animal keepers and carers, animal industry professionals, academics, students and pet owners all over the world.
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Publisher (up) Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6596
Permanent link to this record
 
 
Author McLean, A.N.; Christensen, J.W.
Title The application of learning theory in horse training Type Journal Article
Year 2017 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 190 Issue Pages 18-27
Keywords Behaviour; Conflict theory; Desensitisation; Habituation; Reinforcement; Stress
Abstract The millennia-old practices of horse training markedly predate and thus were isolated from the mid-twentieth century revelation of animal learning processes. From this standpoint, the progress made in the application and understanding of learning theory in horse training is reviewed including a discussion of how learning processes are employed or otherwise under-utilised in training. This review describes the process of habituation and the most commonly applied desensitisation techniques (systematic desensitisation, counter-conditioning, overshadowing, response prevention) and propose two additional techniques (approach conditioning and stimulus blending). The salience of different types of cues, the interaction of operant and classical conditioning and the impact of stress are also discussed. This paper also exposes the inflexibility and occasional inadequacy of the terminology of learning theory when translated from the research laboratory situation to the practical setting in horse training. While learning theory provides a rich toolbox for riders and trainers, the training process is subject to the simultaneous use of multiple learning processes. In addition, learning/behavioural outcomes and trained responses are not just the result of simple stimulus-response based interactions but are further shaped by arousal, affective and attachment states. More research is needed in these areas. For the field of equitation science to progress and to improve clarity and use of learning processes, changes in nomenclature are required. In particular, the use of the terms 'positive' and 'negative' as descriptive labels in both reinforcement and punishment modalities are unacceptably misleading for everyday use. These labels inhibit the understanding and recognition of the learning processes that these terms supposedly represent, yet the learning processes they describe are vital for horse riders, handlers and trainers to understand. We therefore propose that these labels should be re-labelled more appropriately as 'addition' or 'subtraction' reinforcement/punishment. This would enlighten trainers on the correct application of learning theory, and safety and welfare benefits for people and horses would follow. Finally it is also proposed that the term 'conflict theory' be taken up in equitation science to facilitate diagnosis of training-related behaviour disorders and thus enable the emergence of improved training practices. The optimal use of learning theory should be established as a fundamental principle in equestrian education.
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Corporate Author Thesis
Publisher (up) Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0168-1591 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6597
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Author Hendriksen, P.; Elmgreen, K.; Ladewig, J.
Title Trailer-loading of horses: Is there a difference between positive and negative reinforcement concerning effectiveness and stress-related signs? Type Journal Article
Year 2011 Publication Journal of Veterinary Behavior Abbreviated Journal J. Vet. Behav.
Volume 6 Issue 5 Pages 261-266
Keywords negative reinforcement; positive reinforcement; horse; behavior; heart rate; trailer-loading
Abstract The traditional way to train horses is by the application of negative reinforcement (NR). In the past few years, however, the use of positive reinforcement (PR) has become more common. To evaluate the effectiveness and the possible stressor effect of the 2 training methods, 12 horses showing severe trailer-loading problems were selected and exposed to trailer-loading. They were randomly assigned to one of the 2 methods. NR consisted of various degrees of pressure (lead rope pulling, whip tapping). Pressure was removed as soon as the horse complied. PR horses were exposed to clicker training and taught to follow a target into the trailer. Heart rate (HR) was recorded every 5 seconds and behavior denoting discomfort was observed using one-zero sampling with 10 seconds sampling intervals. Training was completed when the horse could enter the trailer upon a signal, or was terminated after a maximum of 15 sessions. Of the 12 horses, 10 reached the criterion within the 15 sessions. One horse was eliminated from the study because of illness and 1 PR horse failed to enter the trailer. A Mann-Whitney U-test indicated that the horses trained with NR displayed significantly more discomfort behavior per training session than horses trained with PR (NR: 13.26 ± 3.25; PR: 3.17 ± 8.93, P < 0.0001) and that horses in the PR group spent less time (second) per session to complete the training criterion (NR: 672.9 ± 247.12; PR: 539.81 ± 166.37, P < 0.01). A Mann-Whitney U-test showed that no difference existed in mean HR (bpm) between the 2 groups (NR: 53.06 ± 11.73 bpm; PR: 55.54 ± 6.7 bpm, P > 0.05), but a Wilcoxon test showed a difference in the PR group between the baseline of HR and mean HR obtained during training sessions (baseline PR: 43 ± 8.83 bpm; PR: 55.54 ± 6.7 bpm, P < 0.05). In conclusion, the PR group provided the fastest training solution and expressed less stress response. Thus, the PR procedure could provide a preferable training solution when training horses in potentially stressing situations.
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Corporate Author Thesis
Publisher (up) Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 1558-7878 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6598
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Author McGreevy, P.D.; McLean, A.N.
Title Punishment in horse-training and the concept of ethical equitation Type Journal Article
Year 2009 Publication Journal of Veterinary Behavior Abbreviated Journal J. Vet. Behav.
Volume 4 Issue 5 Pages 193-197
Keywords conditioning; welfare; ethical equitation; horse; punishment; training
Abstract By definition, punishment makes a response less likely in the future. Because horses are largely trained by negative reinforcement, they are susceptible to inadvertent punishment. Delays in the release of pressure can make desirable responses less likely and thus punish them. This study examines the correct use of negative reinforcement and identifies a continuum between poorly timed negative reinforcement and punishment. It explores some of the problems of non-contingent punishment and the prospect of learned helplessness and experimental neurosis. It concludes by introducing the concept of ethical equitation.
Address
Corporate Author Thesis
Publisher (up) Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 1558-7878 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6599
Permanent link to this record