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| Author | Krueger, K. | ||||
| Title | Behaviour of horses in the “round pen technique” | Type | Journal Article | ||
| Year | 2007 | Publication | Applied Animal Behaviour Science | Abbreviated Journal | Appl. Anim. Behav. Sci. |
| Volume | 104 | Issue | 1-2 | Pages | 162-170 |
| Keywords | Learning; Dominance relationship; Horse; Human-animal relationships | ||||
| Abstract | I investigated the behavioural background of the way horses learn to follow humans in the “round pen technique” suggested by “horse whisperers” as a gentle method for initial horse training. Though the practicability of this technique has been adequately demonstrated in the past, the horses' behaviour during such training has not yet been documented in detail. In a riding arena, horses, that did not follow the trainer immediately, were chased away so that they galloped around the trainer. Galloping horses showed specific behaviour such as turning the ear to the trainer, chewing, licking, and stretching head and throat downwards. In subsequent trials horses needed to be chased for less time and finally followed immediately, even when conditions were changed or the trainer was replaced by another person. This suggests that horses learn to follow in this particular situation and also show some generalisation. However, following did not occur on a pasture even after several successful trials in the riding arena. | ||||
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refbase @ user @ | Serial | 280 | ||
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| Author | Krueger, K.; Flauger, B. | ||||
| Title | Social learning in horses from a novel perspective | Type | Journal Article | ||
| Year | 2007 | Publication | Behavioural Processes | Abbreviated Journal | Behav. Process. |
| Volume | 76 | Issue | 1 | Pages | 37-39 |
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| Notes | Approved | no | |||
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refbase @ user @ | Serial | 625 | ||
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| Author | Schuetz, A.; Farmer, K.; Krueger, K. | ||||
| Title | Social learning across species: horses (Equus caballus) learn from humans by observation | Type | Journal Article | ||
| Year | 2017 | Publication | Animal Cognition | Abbreviated Journal | Anim. Cogn. |
| Volume | 20 | Issue | 3 | Pages | 567-573 |
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| Abstract | This study examines whether horses can learn by observing humans, given that they identify individual humans and orientate on the focus of human attention. We tested 24 horses aged between 3 and 12. Twelve horses were tested on whether they would learn to open a feeding apparatus by observing a familiar person. The other 12 were controls and received exactly the same experimental procedure, but without a demonstration of how to operate the apparatus. More horses from the group with demonstration (8/12) reached the learning criterion of opening the feeder twenty times consecutively than horses from the control group (2/12), and younger horses seemed to reach the criterion more quickly. Horses not reaching the learning criteria approached the human experimenters more often than those that did. The results demonstrate that horses learn socially across species, in this case from humans. | ||||
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| ISSN | 1435-9456 | ISBN | Medium | ||
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| Notes | Approved | no | |||
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Equine Behaviour @ team @ Schuetz2016 | Serial | 6028 | ||
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| Author | Krueger, K. | ||||
| Title | Perissodactyla Cognition | Type | Book Chapter | ||
| Year | 2017 | Publication | Encyclopedia of Animal Cognition and Behavior | Abbreviated Journal | |
| Volume | Issue | Pages | 1-10 | ||
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| Publisher | Springer International Publishing | Place of Publication | Cham | Editor | Vonk, J.; Shackelford, T. |
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| ISSN | ISBN | 978-3-319-47829-6 | Medium | ||
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| Notes | Approved | no | |||
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Equine Behaviour @ team @ Krueger2017 | Serial | 6187 | ||
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| Author | Krueger, K.; Marr, I.; Farmer, K. | ||||
| Title | Equine Cognition | Type | Book Chapter | ||
| Year | 2017 | Publication | Encyclopedia of Animal Cognition and Behavior | Abbreviated Journal | |
| Volume | Issue | Pages | 1-11 | ||
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| Publisher | Springer International Publishing | Place of Publication | Cham | Editor | Vonk, J.; Shackelford, T. |
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| ISSN | ISBN | 978-3-319-47829-6 | Medium | ||
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| Notes | Approved | no | |||
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Equine Behaviour @ team @ Krueger2017 | Serial | 6181 | ||
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| Author | Krueger, K.; Flauger, B. | ||||
| Title | Horses (Equus caballus) show respect and trust in their owners | Type | Book Chapter | ||
| Year | 2009 | Publication | Program of the 4. Thementagung der Ethologischen Gesellschaft, Februar 12 -14, 2009. | Abbreviated Journal | |
| Volume | Issue | Pages | 32 | ||
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| Publisher | Dpz. | Place of Publication | Göttingen | Editor | Kappeler, P.M.; Schwibbe, M. |
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| Notes | Approved | no | |||
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Equine Behaviour @ team @ Krueger2009 | Serial | 5717 | ||
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| Author | Esch, L.; Wöhr, C.; Erhard, M.; Krueger, K. | ||||
| Title | Horses� (Equus Caballus) Laterality, Stress Hormones, and Task Related Behavior in Innovative Problem-Solving | Type | Journal Article | ||
| Year | 2019 | Publication | Animals | Abbreviated Journal | Animals |
| Volume | 9 | Issue | 5 | Pages | 265 |
| Keywords | innovative behavior; brain lateralization; glucocorticoid metabolites; behavioral traits; equine cognition | ||||
| Abstract | Domesticated horses are constantly confronted with novel tasks. A recent study on anecdotal data indicates that some are innovative in dealing with such tasks. However, innovative behavior in horses has not previously been investigated under experimental conditions. In this study, we investigated whether 16 horses found an innovative solution when confronted with a novel feeder. Moreover, we investigated whether innovative behavior in horses may be affected by individual aspects such as: age, sex, size, motor and sensory laterality, fecal stress hormone concentrations (GCMs), and task-related behavior. Our study revealed evidence for 25% of the horses being capable of innovative problem solving for operating a novel feeder. Innovative horses of the present study were active, tenacious, and may be considered to have a higher inhibitory control, which was revealed by their task related behavior. Furthermore, they appeared to be emotional, reflected by high baseline GCM concentrations and a left sensory and motor laterality. These findings may contribute to the understanding of horses� cognitive capacities to deal with their environment and calls for enriched environments in sports and leisure horse management. | ||||
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Equine Behaviour @ team @ Esch2019 | Serial | 6570 | ||
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| Author | Bernauer, K.; Kollross, H.; Schuetz, A.; Farmer, K.; Krueger, K. | ||||
| Title | How do horses (Equus caballus) learn from observing human action? | Type | Journal Article | ||
| Year | 2020 | Publication | Animal Cognition | Abbreviated Journal | Anim. Cogn. |
| Volume | 23 | Issue | Pages | 1-9 | |
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| Abstract | A previous study demonstrated that horses can learn socially from observing humans, but could not draw any conclusions about the social learning mechanisms. Here we develop this by showing horses four different human action sequences as demonstrations of how to press a button to open a feed box. We tested 68 horses aged between 3 and 12 years. 63 horses passed the habituation phase and were assigned either to the group Hand Demo (N = 13) for which a kneeling person used a hand to press the button, Head Demo (N = 13) for which a kneeling person used the head, Mixed Demo (N = 12) for which a squatting person used both head and hand, Foot Demo (N = 12) in which a standing person used a foot, or No Demo (N = 13) in which horses did not receive a demonstration. 44 horses reached the learning criterion of opening the feeder twenty times consecutively, 40 of these were 75% of the Demo group horses and four horses were 31% of the No Demo group horses. Horses not reaching the learning criterion approached the human experimenters more often than those who did. Significantly more horses used their head to press the button no matter which demonstration they received. However, in the Foot Demo group four horses consistently preferred to use a hoof and two switched between hoof and head use. After the Mixed Demo the horses' actions were more diverse. The results indicate that only a few horses copy behaviours when learning socially from humans. A few may learn through observational conditioning, as some appeared to adapt to demonstrated actions in the course of reaching the learning criterion. Most horses learn socially through enhancement, using humans to learn where, and which aspect of a mechanism has to be manipulated, and by applying individual trial and error learning to reach their goal. | ||||
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| ISSN | 1435-9456 | ISBN | Medium | ||
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| Notes | Approved | no | |||
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Equine Behaviour @ team @ Bernauer2019 | Serial | 6590 | ||
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| Author | Marr, I.; Farmer, K.; Krueger, K. | ||||
| Title | Evidence for Right-Sided Horses Being More Optimistic than Left-Sided Horses | Type | Journal Article | ||
| Year | 2018 | Publication | Animals | Abbreviated Journal | Animals |
| Volume | 8 | Issue | 12 | Pages | 219 |
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| Abstract | An individual's positive or negative perspective when judging an ambiguous stimulus (cognitive bias) can be helpful when assessing animal welfare. Emotionality, as expressed in approach or withdrawal behaviour, is linked to brain asymmetry. The predisposition to process information in the left or right brain hemisphere is displayed in motor laterality. The quality of the information being processed is indicated by the sensory laterality. Consequently, it would be quicker and more repeatable to use motor or sensory laterality to evaluate cognitive bias than to perform the conventional judgment bias test. Therefore, the relationship between cognitive bias and motor or sensory laterality was tested. The horses (n = 17) were trained in a discrimination task involving a box that was placed in either a “positive” or “negative” location. To test for cognitive bias, the box was then placed in the middle, between the trained positive and negative location, in an ambiguous location, and the latency to approach the box was evaluated. Results indicated that horses that were more likely to use the right forelimb when moving off from a standing position were more likely to approach the ambiguous box with a shorter latency (generalized linear mixed model, p < 0.01), and therefore displayed a positive cognitive bias (optimistic). | ||||
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| ISSN | 2076-2615 | ISBN | Medium | ||
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| Notes | Approved | no | |||
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Equine Behaviour @ team @ ani8120219 | Serial | 6439 | ||
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| Author | Krueger, K. | ||||
| Title | Social learning and innovative behaviour in horses | Type | Conference Article | ||
| Year | 2015 | Publication | Proceedings of the 3. International Equine Science Meeting | Abbreviated Journal | Proc. 3. Int. Equine. Sci. Mtg |
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| Keywords | social learning, innovative behaviour, Equus caballus, cognitive capacities | ||||
| Abstract | The evaluation of important parameters for measuring the horses’ cognitive capacities is one of the central topics of the equine behaviour team at Nürtingen-Geislingen University. Social complexity has been said to be one of the settings in which needs for cognitive capacities arise in animals. A variety of studies throughout the last two decades proved the horses’ social complexity to be far more elaborate than previously assumed. Horses form social bonds for the protection of offspring, intervene in encounters of others, identify group mates individually and easily orientate in a fission fusion society. In such socially complex societies, animals will benefit from learning socially. In many bird and primate species the degree of social complexity correlates nicely with the species abilities for social learning. Social learning was, therefore, argued to be an indicator for elaborate mental capacities in animals. We were delighted to prove that horses actually copy social behaviour and techniques for operating a feeding apparatus from older and higher ranking group members. In a recent study we found young horses, at the age of 3 to 12, to copy the operation of a feeding apparatus from a human demonstrator. Social learning seems to work nicely in horses when the social background of the animals is considered. The degree to which individual animals adapt to changes in their social or physical environment by finding innovative solution appears to be the other side of the coin, of whether animals adjust to challenges by social learning. It is not very astonishing, that along with the animals’ social complexity and their ability to learn socially also the degree to which they show innovative behaviour was claimed to be one of the most important demonstrations of advanced cognitive capacities. In a recent approach, we started to ask horse owners and horse keepers in many countries to tell us about unusual behaviour of their horses via a web site (http://innovative-behaviour.org). To date, we received 204 cases of innovative behaviour descriptions from which six cases were clear examples of tool use or borderline tool use. We categorized the innovative behaviours into the classes, a) innovations to gain food, b) innovations to gain freedom, c) social innovations, d) innovations to increase maintenance, and e) innovations that could not be clearly assigned to a category. About 20% of the innovative horses showed more than one innovation. These animals could be termed “true innovators”. Again, young horses were more innovative than older ones with the age group 5 – 9 showing the highest number of innovative behaviour descriptions. In a nutshell, the horses’ cognitive capacities appear to be underestimated throughout the last decades. The horses’ social complexity is far more elaborate than previously assumed, horses learn socially from conspecific and humans, some of them demonstrate innovative behaviour adaptations to their environment and even simple forms of tool use. |
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| Corporate Author | Krueger, K. | Thesis | |||
| Publisher | Xenophon Publishing | Place of Publication | Wald | Editor | |
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| Series Volume | in prep | Series Issue | Edition | ||
| ISSN | ISBN | 978-3-95625-000-2 | Medium | ||
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| Notes | Approved | no | |||
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Equine Behaviour @ team @ | Serial | 5848 | ||
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