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Author Broom, M.; Cannings, C.
Title Modelling Dominance Hierarchy formation as a Multi-player game Type Journal Article
Year 2002 Publication Journal of Theoretical Biology Abbreviated Journal J. Theor. Biol.
Volume 219 Issue 3 Pages 397-413
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Abstract Animals who live in groups need to divide available resources amongst themselves. This is often achieved by means of a dominance hierarchy, where dominant individuals obtain a larger share of the resources than subordinate individuals. This paper introduces a model of dominance hierarchy formation using a multi-player extension of the classical Hawk-Dove game. Animals play non-independent pairwise games in a Swiss tournament which pairs opponents against those which have performed equally well in the conflict so far, for a fixed number of rounds. Resources are divided according to the number of contests won. The model, and its emergent properties, are discussed in the context of experimental observations.
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Call Number refbase @ user @ Serial 450
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Author Jameson, K.A.; Appleby, M.C.; Freeman, L.C.
Title Finding an appropriate order for a hierarchy based on probabilistic dominance Type Journal Article
Year 1999 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume 57 Issue 5 Pages 991-998
Keywords
Abstract Methods of ranking individuals in a dominance hierarchy that use transitivity of relationships may obscure irregularities. Furthermore, these methods use only a small proportion of the information available from dominance encounters. This paper presents an intuitively appealing and easily implemented alternative to existing methods for ordering dominance data, developed from the work of Batchelder et al. (1992Journal of Mathematical Psychology36, 185-212). The procedure presented here is based on a mathematical model of paired comparisons and it involves only simple estimation procedures. We illustrate its use with data on dominance among red deerCervus elaphus, stags. The results indicate that dominance relationships are well characterized by the scale values that the model provides, and, because the method provides predictions for all pairings of animals, dominance predictions also exist for pairs of animals that have yet to be observed. Moreover, the dominance outcomes predicted by the model using the order scale are highly correlated with actual dominance observations at all levels. Overall, the procedure described provides a solution to the problem of identifying an appropriate order for a near-linear dominance hierarchy.
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Call Number refbase @ user @ Serial 452
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Author Gammell, M.P.; de Vries, H.; Jennings, D.J.; Carlin, C.M.; Hayden, T.J.
Title David's score: a more appropriate dominance ranking method than Clutton-Brock et al.'s index Type Journal Article
Year 2003 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume 66 Issue 3 Pages 601-605
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Call Number refbase @ user @ Serial 453
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Author RHO, J.R.; SRYGLEY, R.B.; CHOE, J.C.
Title Behavioral ecology of the Jeju pony (Equus caballus): Effects of maternal age, maternal dominance hierarchy and foal age on mare aggression Type Journal Article
Year 2004 Publication Ecological Research Abbreviated Journal Ecol. Res.
Volume 19 Issue 1 Pages 55-63
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Abstract On Jeju Island, Korea, dominance hierarchy and maternal care according to maternal age were studied in a herd of Jeju ponies (Equus caballus), consisting of 73 mares, their foals and one stallion. Dominance ranks were nearly linear and increased significantly with the age of mares. Most aggressive encounters involved mares under 5 years old. Mares under the age of 5 years have apparently not established their rank. The mean frequency of aggressive actions of mares per hour increased significantly as the day of parturition approached. Aggressive actions of mares with foals decreased significantly as their foals aged. The overall frequency of aggression of mares with foals also decreased significantly with the age of the mares. Our results suggest that the cost of maternal care is lower for older, more dominant mares than for subordinate ones.
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Call Number refbase @ user @ Serial 459
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Author Feh, C.
Title Alliances and reproductive success in Camargue stallions Type Journal Article
Year 1999 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume 57 Issue 3 Pages 705-713
Keywords
Abstract A study of a herd of Camargue horsesEquus caballus, showed that while the majority of high-ranking stallions held single-male harems, some sons of low-ranking mares, being low ranking themselves, formed alliances that could last a lifetime. The two stallions were each other's closest associate and preferential grooming partner. Alliances were based on coalitions in which either both partners confronted an intruder synchronously or the dominant of the pair tended the female(s) while the subordinate simultaneously displayed towards the rival. Alliance partners were of similar age but were not more closely related to each other than to other stallions in the herd. Long-term paternity data revealed that subordinates sired close to a quarter of the foals born into the alliance group, and significantly more foals than low-ranking stallions in the herd adopting a `sneak'-mating strategy. The dominant appeared to benefit from the presence of his subordinate partner. Fights occurred all year round, and the subordinate stallion of each alliance pair fought outside competitors more than twice as often as the dominant. Forming short-term alliances before defending mares on their own may enhance long-term reproductive success for both partners. Other benefits to both partners include higher survivorship of their foals and increased access to proven reproductive mares. These results suggest that the relationship between alliance partners is based on mutualism, but several conditions for reciprocity seem to be fulfilled: the benefit to the dominant (assistance in fights), and the benefit to the subordinate (access to reproduction), are both costly to the other partner and delayed in time.
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Call Number refbase @ user @ Serial 469
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Author Moons, C.; Heleski, C.R.; Leece, C.M.; Zanella, A.J.
Title Conflicting Results in the Association Between Plasma and Salivary Cortisol Levels in Foals Type Manuscript
Year 2002 Publication Havemeier Workshop Abbreviated Journal
Volume Issue Pages
Keywords
Abstract Introduction

Glucocorticoids are present in many biological fluids as a free fraction or bound to Corticoid

Binding Globulins (CBG) (Matteri et al, 2000). There are conflicting claims regarding the validity of

saliva as a biological fluid to measure cortisol in horses (Lebelt et al, 1996; McGreevy and Pell, 1998;

van der Kolk et al, 2001). Measuring changes in salivary cortisol levels in normal horses and horses

with Cushing`s disease van der Kolk and collaborators (2001) demonstrated the validity of saliva to

assess adrenal function. Puzzling results were reported by McGreevy and Pell (1998) who suggested

that plasma and salivary cortisol concentrations in horses showing oral stereotypies were correlated

but this association was non-existent in control animals. Investigating the responses of foals to

branding and foot-trimming Zanella et al (unpublished results) were unable to identify a relationship

between plasma and salivary cortisol levels in foals. In several species, levels of cortisol in plasma and

saliva are tightly correlated (Fenske, 1996). Cortisol found in blood consists of a fraction bound to

corticoid binding globulin (CBG) and a free, unbound fraction. Free cortisol represents the

biologically active fraction of this steroid hormone. Salivary cortisol reflects the unbound fraction

found in plasma or serum and it passes readily through the parotid membrane (Riad-Fahmy, 1983;

Horning Walker et al,1977). Unbound steroids transfer rapidly between plasma and saliva

(Walker,1989; Scott et al 1990). Saliva flow-rate does not appear to influence saliva cortisol levels in

different species (Hubert and de Jong-Meyer, 1989; Walker 1989, Scott et a, 1990). In horses, Lebelt

et al (1996) reported that salivary and plasma total cortisol in stallions were correlated. We

hypothesized that changes in salivary cortisol in foals would show a pattern that is correlated to that of

plasma free and plasma total cortisol concentrations in foals. In addition, we anticipated that the lack

of good sampling techniques provides an explanation for the failure in determining the association

between salivary and plasma cortisol in foals.
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Call Number refbase @ user @ Serial 470
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Author Feh, C.
Title Relationships and Communication in Socially Natural Horse Herds Type Book Chapter
Year 2005 Publication The domestic horse : the origins, development, and management of its behaviour Abbreviated Journal The domestic horse : the origins, development, and management of its behaviour
Volume Issue Pages
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Abstract Horses are quite unique. In most mammals, sexes segregate and maintain bonds only during the breeding season (Clutton-Brock, 1989). Some canids, a few rodents and primate species such as gorillas, hamadryas baboons and red howler monkeys are the exception, where the same males stay with the same females all year round and over many breeding seasons. Typically, both sexes disperse at puberty in these species. In horses, it was clearly shown that the causes for female dispersal were incest avoidance and not intra-specific competition (Monard, 1996). As a rule, this is confirmed for mammal species where tenure length by males exceeds the age at first reproduction in females (Clutton-Brock, 1989). When horses are allowed to choose their mating partner freely, the inbreeding coefficient of the offspring is lower than expected should they mate randomly (Duncan et al, 1984).
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Publisher Cambridge University Press 2005 Place of Publication Cambridge Editor Mills, D. S. ; McDonnell, , S. M.
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Series Volume Series Issue Edition
ISSN ISBN 13 978-0-521-81414-6 Medium
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Call Number refbase @ user @; Equine Behaviour @ team @ room B 3.092 Serial 472
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Author Nicol, C.J
Title Equine Stereotypies. In: Houpt K.A. (Ed.), Type Book Chapter
Year 2000 Publication Recent Advances in Companion Animal Behavior Problems Abbreviated Journal
Volume Issue Pages
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Publisher International Veterinary Information Service Place of Publication Editor
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Call Number refbase @ user @ Serial 477
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Author Feh, C.
Title Alliances between stallions are more than just multimale groups: reply to Linklater & Cameron (2000) Type Journal Article
Year 2001 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume 61 Issue Pages F27-F30
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Call Number refbase @ user @ Serial 513
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Author Seaman, S.C.; Davidson, H.P.B.; Waran, N.K.
Title How reliable is temperament assessment in the domestic horse (Equus caballus)? Type Journal Article
Year 2002 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 78 Issue 2-4 Pages 175-191
Keywords Temperament assessment; Behavioural tests; Horses; Active and passive copers; Factor analysis
Abstract Differences in behavioural characteristics between individuals of the same species are often described as being due to the temperament of the individuals. These differences can have enormous implications for welfare with some individuals apparently being able to adapt to environmental challenge more easily than others. Such differences have resulted in animals often being described as either `active' copers, which try to escape from or remove an aversive stimulus, or `passive' copers, which show no outward signs of a situation being aversive, thus, appearing to be unaffected. Tests previously developed to assess the temperament of animals have been criticised for several reasons. Behaviour is often recorded and categorised using methods that are not objective and tests are generally carried out once with no consideration of whether or not behavioural responses are consistent over time. This study takes these factors into account. The behaviour of 33 horses was recorded in three types of test--an arena test, response to a person and response to an object. In order to test whether or not responses were consistent over time, the tests were repeated three times with an average of 9 days between trials. Test results were validated using responses from questionnaires completed by the farm team leader. The data were analysed using an initial principal component analysis (PCA) and factor analysis. The horses were found to behave consistently over the three trials in their responses in the arena test. The responses to the person test and the object test were similar to each other; however, these responses were not consistent over trials. The behaviour in the arena test was unable to be used to make a prediction of behaviour in the person and object tests and vice versa. The responses shown by the horses did not enable them to be categorised as either active or passive copers. Behavioural responses in the tests were not predictive of the response to a startle test (water spray), nor could they be used to predict status or response to being reintroduced to the group after testing. There was no relationship between the responses in the tests and the ratings given by the farm team leader. It was concluded that horses vary widely in their responses to artificial behavioural tests, with only the responses to an open-field arena test being consistent over time, and therefore, the only type of test which can indicate some core factor of temperament.
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Call Number refbase @ user @ Serial 520
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