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Author Burns, T.E.; Clayton, H.M. openurl 
  Title Comparison of the temporal kinematics of the canter pirouette and collected canter Type Journal Article
  Year (up) 1997 Publication Equine Veterinary Journal. Supplement Abbreviated Journal Equine Vet J Suppl  
  Volume Issue 23 Pages 58-61  
  Keywords Animals; Forelimb/physiology; Gait/*physiology; Hindlimb/physiology; Horses/*physiology; Locomotion/*physiology; Movement/physiology; Time Factors; Videotape Recording; Walking/physiology  
  Abstract The objectives were to compare the temporal characteristics of canter pirouette strides with collected canter strides in elite dressage horses, and to determine whether the stride kinematics of the canter pirouettes fulfilled the requirements specified in the Federation Equestre Internationale Rules for Dressage Events. Eleven horses were videotaped (60 fields/s) during the individual medal competition at the 1992 Olympic Games. Temporal variables were extracted from the videotapes using standard methods. Two strides were analysed on each of the left and right leads and these were pooled to give mean values for the collected canter and the pirouettes. The pirouettes were completed in 4-9 strides, (mean of 6.4). In the collected canter strides, mean duration of the suspension was 0.013 s. There was no suspension in any of the pirouette strides, instead the stance phases of the leading forelimb and trailing hindlimb overlapped by a mean of 0.163 s. In 9 horses the trailing forelimb contacted the ground before the diagonal leading hindlimb in the collected canter, whereas in the pirouettes the leading hindlimb always made contact before the trailing forelimb (mean dissociation 0.164 s), giving the strides a distinct 4 beat rhythm. Due to increases in advanced placement between the diagonal limb pair and between the 2 forelimbs, the stride duration was longer in the pirouette (0.879 s) than the collected canter (0.629 s). It is concluded that the canter pirouette strides did not maintain the rhythm and timing of the the collected canter strides in any of the 11 horses.  
  Address Department of Veterinary Anatomy, Western College of Veterinary Medicine, University of Saskatchewan, Saskatoon, Canada  
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  Notes PMID:9354291 Approved no  
  Call Number Equine Behaviour @ team @ Serial 3737  
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Author Clayton, H.M. openurl 
  Title Classification of collected trot, passage and piaffe based on temporal variables Type Journal Article
  Year (up) 1997 Publication Equine Veterinary Journal. Supplement Abbreviated Journal Equine Vet J Suppl  
  Volume Issue 23 Pages 54-57  
  Keywords Analysis of Variance; Animals; Discriminant Analysis; Gait/*physiology; Hindlimb/physiology; Horses/*physiology; Locomotion/*physiology; Male; Running/physiology; Time Factors; Videotape Recording; Walking/physiology  
  Abstract The objective was to determine whether collected trot, passage and piaffe could be distinguished as separate gaits on the basis of temporal variables. Sagittal plane, 60 Hz videotapes of 10 finalists in the dressage competitions at the 1992 Olympic Games were analysed to measure the temporal variables in absolute terms and as percentages of stride duration. Classification was based on analysis of variance, a graphical method and discriminant analysis. Stride duration was sufficient to distinguish collected trot from passage and piaffe in all horses. The analysis of variance showed that the mean values of most variables differed significantly between passage and piaffe. When hindlimb stance percentage was plotted against diagonal advanced placement percentage, some overlap was found between all 3 movements indicating that individual horses could not be classified reliably in this manner. Using hindlimb stance percentage and diagonal advanced placement percentage as input in a discriminant analysis, 80% of the cases were classified correctly, but at least one horse was misclassified in each movement. When the absolute, rather than percentage, values of the 2 variables were used as input in the discriminant analysis, 90% of the cases were correctly classified and the only misclassifications were between passage and piaffe. However, the 2 horses in which piaffe was misclassified as passage were the gold and silver medallists. In general, higher placed horses tended toward longer diagonal advanced placements, especially in collected trot and passage, and shorter hindlimb stance percentages in passage and piaffe.  
  Address Western College of Veterinary Medicine, University of Saskatchewan, Saskatoon, Canada  
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  Notes PMID:9354290 Approved no  
  Call Number Equine Behaviour @ team @ Serial 3738  
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Author Barrey, E.; Galloux, P. openurl 
  Title Analysis of the equine jumping technique by accelerometry Type Journal Article
  Year (up) 1997 Publication Equine Veterinary Journal. Supplement Abbreviated Journal Equine Vet J Suppl  
  Volume Issue 23 Pages 45-49  
  Keywords *Acceleration; Analysis of Variance; Animals; Forelimb/physiology; Hindlimb/physiology; Horses/*physiology; Locomotion/*physiology; Movement/physiology; Time Factors  
  Abstract The purpose of this study was to demonstrate the relationships between jumping technique and dorsoventral acceleration measured at the sternum. Eight saddle horses of various jumping abilities competed on a selective experimental show jumping course including 14 obstacles. An accelerometric belt fastened onto the thorax continuously measured the dorsoventral acceleration during the course. At each jump, 11 locomotor parameters (acceleration peaks, durations and stride frequency) were obtained from the dorsoventral acceleration-time curves. The type of obstacle significantly influenced the hindlimb acceleration peak at take-off and the landing acceleration peak (P<0.01). The poor jumpers exhibited a higher mean forelimb acceleration peak at take-off, a higher forelimb/hindlimb ratio between peaks of acceleration (F/H), and a lower approach stride frequency than good jumpers. Knocking over an obstacle was significantly associated with a low hindlimb acceleration peak at take-off and a high F/H ratio (P<0.01). In order to observe the continuous changes in the frequency domain of the dorsoventral acceleration during the approach and take-off phase, a Morlet's wavelet analysis was computed for each horse jumping over a series of 3 vertical obstacles. Different patterns of time-frequency images obtained by wavelet analysis were found when the horse either knocked over a vertical obstacle or cleared it. In the latter case, the image pattern showed an instantaneous increase in stride frequency at the end of the approach phase, and a marked energy content in the middle frequency range at take-off.  
  Address INRA Station de Genetique Quantitative et Appliquee, Groupe cheval, Jouy-en-Josas, France  
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  Notes PMID:9354288 Approved no  
  Call Number Equine Behaviour @ team @ Serial 3796  
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Author Galloux, P.; Barrey, E. openurl 
  Title Components of the total kinetic moment in jumping horses Type Journal Article
  Year (up) 1997 Publication Equine Veterinary Journal. Supplement Abbreviated Journal Equine Vet J Suppl  
  Volume Issue 23 Pages 41-44  
  Keywords Algorithms; Animals; Exertion/*physiology; Female; Gravitation; Horses/*physiology; Kinetics; Locomotion/*physiology; Male; Models, Biological; Movement/*physiology; Video Recording  
  Abstract Thirty horses were filmed with a panning camera operating at 50 frames/s as they jumped over a 1.20 x 1.20 m fence. The markers of 9 joints on the horse and 7 joints on the rider were tracked in 2D with the TrackEye system. The centre of gravity and moment of inertia of each segment were calculated using a geometric algorithm and a cylindric model, respectively. The kinetic moment of each part of the horse was calculated after filtering, and resampling of data. This method showed the relative contribution of each body segment to the body overall rotation during the take-off, jump and landing phases. It was found that the trunk, hindlimbs and head-neck had the greatest influence. The coordination between the motion of the body segments allowed the horse to control its angular speed of rotation over the fence. This remained nearly constant during the airborne phase (120 +/- 5 degrees/s). During the airborne phase, the kinetic moment was constant because its value was equal to the moment of the external forces (722 +/- 125 kg x m2/s).  
  Address Ecole Nationale d'Equitation, Terrefort, Saumur, France  
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  Notes PMID:9354287 Approved no  
  Call Number Equine Behaviour @ team @ Serial 3797  
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Author Moehlman, P.D. url  doi
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  Title Behavioral patterns and communication in feral asses (Equus africanus) Type Journal Article
  Year (up) 1998 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 60 Issue 2-3 Pages 125-169  
  Keywords Equids; Feral asses; Behavior patterns; Facial expressions; Postures; Locomotion  
  Abstract The behavior of feral populations of the African wild ass (Equus africanus) were studied in the Northern Panamint Range of Death Valley National Monument for 20 months from 1970 to 1973 [Moehlman, P.D., 1974. Behavior and ecology of feral asses (Equus asinus). PhD dissertation, University of Wisconsin, Madison, 251 pp.; Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Natl. Geogr. Soc. Res. Reports, 1970: 405-411]. Maintenance behavior is described and behavior sequences that were used in social interactions are quantified by sex and age class. Agonistic, sexual, and greeting behavior patterns are described and analyzed in conjunction with the responses they elicited. Mutual grooming mainly occurred between adult males, and between females and their offspring. Five types of vocalizations were distinguished: brays, grunts, growls, snorts, and whuffles. A second population was studied for 1 month on Ossabaw Island, GA (Moehlman, 1979). This population had more permanent social groups and had a higher rate of mutual grooming and foal social play.  
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  Call Number Equine Behaviour @ team @ Serial 2381  
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Author Hodson, E.F.; Clayton, H.M.; Lanovaz, J.L. url  openurl
  Title Temporal analysis of walk movements in the Grand Prix dressage test at the 1996 Olympic Games Type Journal Article
  Year (up) 1999 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 62 Issue 2-3 Pages 89-97  
  Keywords Dressage; Horse; Kinematics; Locomotion; Gait  
  Abstract Video analysis was used to measure temporal characteristics of the collected walk, extended walk and half pirouette at walk of eleven competitors during the team dressage competition at the 1996 Summer Olympic Games in Atlanta, GA. Forelimb stance durations, hind limb stance durations, lateral step intervals and diagonal step intervals were symmetrical for the right and left sides in the collected and extended walk strides, but there were left-right asymmetries in the forelimb stance duration and in the lateral step interval in the half pirouette strides. For both collected and extended walk strides, hind limb stance duration was significantly longer than forelimb stance duration. The mean values for the group of eleven horses showed that the collected and extended walks had a regular rhythm. The half pirouette strides showed an irregularity in which there was a short interval between footfalls of the outside forelimb and inside hind limb, and along interval between footfalls of the inside hind limb and inside forelimb. This irregularity reflected an early placement of the inside hind limb. The stance times of both hind limbs were prolonged and this finding, in combination with the early placement of the inside hind limb, led to an increase in the period of tripedal support in each stride of the half pirouette. This was interpreted as a means of maintaining the horses' balance in the absence of forward movement.  
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  Call Number refbase @ user @ Serial 3960  
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Author Krause Hoare; Hemelrijk; Rubenstein url  doi
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  Title Leadership in fish shoals Type Journal Article
  Year (up) 2000 Publication Fish and Fisheries Abbreviated Journal Fish Fish  
  Volume 1 Issue Pages 82-89  
  Keywords directional locomotion; fish schools; front fish; nutritional state; schooling; shoal leadership; swimming direction  
  Abstract Leadership is not an inherent quality of animal groups that show directional locomotion. However, there are other factors that may be responsible for the occurrence of leadership in fish shoals, such as individual differences in nutritional state between group members. It appears that front fish have a strong influence on directional shoal movements and that individuals that occupy such positions are often characterised by larger body lengths and lower nutritional state. Potential interactions between the two factors and their importance for positioning within shoals need further attention. Initiation of directional movement in stationary shoals and position preferences in mobile shoals need to be addressed separately because they are potentially subject to different constraints. Individuals that initiate a swimming direction may not necessarily be capable of the sustained high swimming performance required to keep the front position or have the motivation to do so, for that matter. More empirical and theoretical work is necessary to look at the factors controlling positioning behaviour within shoals, as well as overall shoal shape and structure. Tracking of marked individuals whose positioning behaviour is monitored over extended time periods of hours or days would be useful. There is an indication that shoal positions are rotated by individuals according to their nutritional needs, with hungry fish occupying front positions only for as long as necessary to regain their nutritional balance. This suggests that shoal members effectively take turns at being leaders. There is a need for three-dimensional recordings of shoaling behaviour using high-speed video systems that allow a detailed analysis of information transfer in shoals of different size. The relationship between leadership and shoal size might provide an interesting field for future research. Most studies to date have been restricted to shoals of small and medium size and more information on larger shoals would be useful.  
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  Call Number Serial 2067  
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Author Buchner, H.H.F.; Obermuller, S.; Scheidl, M. url  openurl
  Title Body Centre of Mass Movement in the Sound Horse Type Journal Article
  Year (up) 2000 Publication The Veterinary Journal Abbreviated Journal  
  Volume 160 Issue 3 Pages 225-234  
  Keywords Horse; centre of mass; kinematics; segment model; locomotion.  
  Abstract The body centre of mass (BCM) is a key factor in the analysis of equine locomotion, as its position and movement determines the distribution and magnitude of loads on the limbs. In this study, the three-dimensional (3D) movement of the BCM in walking and trotting horses was assessed using a kinematic, segmental method. Thirty markers representing 20 body segments were recorded in 12 sound horses while standing, walking and trotting on a treadmill using a high-speed video system. Based on segmental inertial data, 3D positions of the segmental centres of mass as well as the total BCM were calculated. The position within the trunk during square standing and the movements of the BCM were determined for the three planes. The position of the BCM in the standing horse is presented relative to external reference points. At the trot, vertical displacement amplitude of the BCM amounted to 53 (6) mm as mean (sd), which was 27% smaller than external trunk movement. Medio-lateral displacement amplitude of the BCM was 19 (4) mm, 34% less than trunk amplitude. Sagittal forward-backward oscillations of the BCM independent from general forward movement were 13 (3) mm, being 24% less than trunk movements. At the walk, vertical, medio-lateral and sagittal BCM movements were smaller than trunk movements by 43, 65 and 65% respectively. The results show reduced and efficient BCM movements compared to the trunk and form a basis for the assessment of various clinical conditions such as lameness, the influence of a rider and various dressage performances.  
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  Call Number refbase @ user @ Serial 3959  
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Author Wilson, A.M.; McGuigan, M.P.; Su, A.; van Den Bogert, A.J. doi  openurl
  Title Horses damp the spring in their step Type Journal Article
  Year (up) 2001 Publication Nature Abbreviated Journal Nature  
  Volume 414 Issue 6866 Pages 895-899  
  Keywords Animals; Biomechanics; Elasticity; Forelimb; Gait; Horses/anatomy & histology/*physiology; Leg Bones/*physiology; Locomotion; Models, Biological; Muscle Fibers/physiology; Muscle, Skeletal/anatomy & histology/*physiology; Tendons/anatomy & histology/*physiology; Vibration  
  Abstract The muscular work of galloping in horses is halved by storing and returning elastic strain energy in spring-like muscle-tendon units.These make the legs act like a child's pogo stick that is tuned to stretch and recoil at 2.5 strides per second. This mechanism is optimized by unique musculoskeletal adaptations: the digital flexor muscles have extremely short fibres and significant passive properties, whereas the tendons are very long and span several joints. Length change occurs by a stretching of the spring-like digital flexor tendons rather than through energetically expensive length changes in the muscle. Despite being apparently redundant for such a mechanism, the muscle fibres in the digital flexors are well developed. Here we show that the mechanical arrangement of the elastic leg permits it to vibrate at a higher frequency of 30-40 Hz that could cause fatigue damage to tendon and bone. Furthermore, we show that the digital flexor muscles have minimal ability to contribute to or regulate significantly the 2.5-Hz cycle of movement, but are ideally arranged to damp these high-frequency oscillations in the limb.  
  Address Department of Veterinary Basic Sciences, The Royal Veterinary College, Hatfield, Herts AL9 7TA, UK. awilson@rvc.ac.uk  
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  ISSN 0028-0836 ISBN Medium  
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  Notes PMID:11780059 Approved no  
  Call Number Equine Behaviour @ team @ Serial 2300  
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Author Meershoek, L.S.; Schamhardt, H.C.; Roepstorff, L.; Johnston, C. openurl 
  Title Forelimb tendon loading during jump landings and the influence of fence height Type Journal Article
  Year (up) 2001 Publication Equine Veterinary Journal. Supplement Abbreviated Journal Equine Vet J Suppl  
  Volume Issue 33 Pages 6-10  
  Keywords Animals; Biomechanics; Forelimb/injuries/physiology; Horses/injuries/*physiology; Lameness, Animal/etiology; Ligaments, Articular/*physiology; Locomotion/*physiology; Physical Conditioning, Animal; Tendon Injuries/complications/physiopathology/veterinary; Tendons/*physiology; Weight-Bearing/physiology  
  Abstract Lameness in athletic horses is often caused by forelimb tendon injuries, especially in the interosseus tendon (TI) and superficial digital flexor tendon (SDF), but also in the accessory ligament (AL) of the deep digital flexor tendon (DDF). In an attempt to explain the aetiology of these injuries, the present study investigated the loading of the tendons during landing after a jump. In jumping horses, the highest forces can be expected in the trailing limb during landing. Therefore, landing kinematics and ground reaction forces of the trailing forelimb were measured from 6 horses jumping single fences with low to medium heights of 0.80, 1.00 and 1.20 m. The tendon forces were calculated using inverse dynamics and an in vitro model of the lower forelimb. Calculated peak forces in the TI, SDF and DDF + AL during landing were 15.8, 13.9 and 11.7 kN respectively. The relative loading of the tendons (landing forces compared with failure forces determined in a separate study) increased from DDF to TI to SDF and was very high in SDF. This explains the low injury incidence of the DDF and the high injury incidence of the SDF. Fence height substantially influenced SDF forces, whereas it hardly influenced TI forces and did not influence AL strain. Reduction of fence height might therefore limit the risks for SDF injuries, but not for TI and AL injuries.  
  Address Department of Veterinary Anatomy and Physiology, Institute for Fundamental and Clinical Human Movement Sciences, Faculty of Veterinary Medicine, Utrecht University, Utrecht, The Netherlands  
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  Notes PMID:11721571 Approved no  
  Call Number Equine Behaviour @ team @ Serial 3786  
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