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	Author	Langbein, J.; Siebert, K.; Nuernberg, G.; Manteuffel, G.
	Title	The impact of acoustical secondary reinforcement during shape discrimination learning of dwarf goats (Capra hircus)			Type	Journal Article
	Year	2007	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	103	Issue	1-2	Pages	35-44
	Keywords	Dwarf goats; Operant conditioning; Visual discrimination learning; Secondary reinforcement
	Abstract	The use of secondary reinforcement is widely accepted to support operant learning in animals. In farm animals, however, the efficacy of secondary reinforcement has up to now been studied systematically only in horses (“clicker training”), and the results are controversial. We investigated the impact of acoustical secondary reinforcement on voluntary, self-controlled visual discrimination learning of two-dimensional shapes in group-housed dwarf goats (Capra hircus). Learning tests were conducted applying a computer-controlled learning device that was integrated in the animals' home pen. Shapes were presented on a TFT-screen using a four-choice design. Drinking water was used as primary reinforcement. In the control group (Gcontrol, n = 5) animals received only primary reinforcement, whereas in the sound group (Gsound, n = 6) animals got additional acoustical secondary reinforcement. Testing recall of shapes which had been successfully learned by the goats 6 weeks earlier (T1), we found a weak impact of secondary reinforcement on daily learning success (P = 0.07), but not on the number of trials the animals needed to reach the learning criterion (trials to criterion, n.s.). Results in T1 indicated that dwarf goats did not instantly recall previously learned shapes, but, re-learned within 250-450 trials. When learning a set of new shapes (T2), there was a strong influence of secondary reinforcement on daily learning success and on trials to criterion. Animals in Gsound reached the learning criterion earlier (P < 0.05) and needed fewer trials (1320 versus 3700; P < 0.01), compared to animals in Gcontrol. Results suggest that acoustical secondary reinforcement supports visual discrimination learning of dwarf goats, especially when the task is new and the salience of S+ is low.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	ISSN		ISBN		Medium
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3583
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	Author	Henning, J.M.; Zentall, T.R.
	Title	Imitation, social facilitation, and the effects of ACTH 4-10 on rats' bar-pressing behavior			Type	Journal Article
	Year	1981	Publication	The American journal of psychology	Abbreviated Journal	Am J Psychol
	Volume	94	Issue	1	Pages	125-134
	Keywords	Adrenocorticotropic Hormone/pharmacology; Animals; Conditioning, Operant/drug effects; Dose-Response Relationship, Drug; Extinction, Psychological/drug effects; Imitative Behavior/drug effects; Male; Peptide Fragments/pharmacology; Rats; *Social Facilitation
	Abstract	The effects of ACTH 4-10 on rats' imitation learning was examined during the acquisition and extinction of a bar-press response for water reinforcement. Rats were exposed to either a bar-pressing conspecific (OB), an experimentally naive conspecific (ON), or an empty box (OE) during bar-press acquisition. In a factorial design, each rat was then exposed to one of the same three conditions during extinction. An 80 mcg dose of ACTH 4-10 was administered to half of the rats in each group prior to observation. Performance differences during acquisition were generally small, but significant performance differences during extinction were found. Social facilitation was indicated by the finding that rats extinguished in the presence of a conspecific exhibited significantly greater resistance to extinction than rats extinguished in the presence of an empty box. An imitation effect was also found. Rats that observed a bar-pressing conspecific during both acquisition and extinction (group OB-OB) showed significantly greater resistance top extinction than did groups OB-ON, CB-OE, or OE-OE. There were no significant effects of the hormone, however, relative to saline controls.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0002-9556	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:6263117			Approved	no
	Call Number	refbase @ user @			Serial	267
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	Author	Nallan, G.B.; Pace, G.M.; McCoy, D.F.; Zentall, T.R.
	Title	Temporal parameters of the feature positive effect			Type	Journal Article
	Year	1979	Publication	The American journal of psychology	Abbreviated Journal	Am J Psychol
	Volume	92	Issue	4	Pages	703-710
	Keywords	Animals; Columbidae; Conditioning, Operant; Discrimination Learning; Form Perception; Male; Time Perception
	Abstract	Trial duration and intertrial interval duration were parametrically varied between groups of pigeons exposed to a discrimination involving the presence vs. the absence of a dot. Half the groups received the dot as the positive stimulus (feature positive groups) and half the groups received the dot as the negative stimulus (feature negative groups). Faster learning by the feature positive birds (feature positive effect) was found when the trial duration was short (5 sec) regardless of whether the intertrial interval was short (5 sec) or long (30 sec). No evidence for a feature positive effect was found when the trial duration was long (30 sec) regardless of the length of the intertrial interval (30 sec or 180 sec). The results suggest that short trial duration is a necessary condition for the occurrence of the feature positive effect, and neither intertrial interval nor trial duration/intertrial interval ratio are important for its occurrence. The suggestion that mechanisms underlying the feature positive effect and autoshaping might be similar was not supported by the present experiment since the trial duration/intertrial interval ration parameter appears to play an important role in autoshaping but not the feature positive effect.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0002-9556	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:532834			Approved	no
	Call Number	refbase @ user @			Serial	269
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	Author	Williams, J.L.; Friend, T.H.; Nevill, C.H.; Archer, G.
	Title	The efficacy of a secondary reinforcer (clicker) during acquisition and extinction of an operant task in horses			Type	Journal Article
	Year	2004	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	88	Issue	3-4	Pages	331-341
	Keywords	Equine; Horse; Clicker training; Operant conditioning; Secondary reinforcers
	Abstract	“Clicker training” is a popularly promoted training method based on operant conditioning with the use of a secondary reinforcer (the clicker). While this method draws from theories of learning and is used widely, there has been little scientific investigation of its efficacy. We used 60 horses, Equus callabus, and assigned each horse to one of six reinforcement protocols. The reinforcement protocols involved combinations of reinforcers administered (primary versus secondary plus primary), schedule of reinforcement (continuous versus variable ratio), and reinforcers applied during extinction (none or secondary). There were no differences (P>=0.11) between horses which received a secondary reinforcer (click) followed by the primary reinforcer (food) and those which received only the primary reinforcer (food) in the number of trials required to train the horses to touch their noses to a plastic cone (operant response). There also were no differences (P>=0.12) between horses which received the secondary reinforcer plus primary reinforcer and those which received only the primary reinforcer in regards to the number of trials to extinction. We conclude that there is no difference in the amount of training required to learn the operant task or in the task's resistance to extinction between horses that received a secondary reinforcer followed by a primary reinforcer versus horses which received only a primary reinforcer.
	Address
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	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3581
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	Author	Hagen, K.; Broom, D.M.
	Title	Emotional reactions to learning in cattle			Type	Journal Article
	Year	2004	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	85	Issue	3	Pages	203-213
	Keywords	Cattle; Expressive behaviour; Operant learning; Reinforcer
	Abstract	It has been suggested that during instrumental learning, animals are likely to react emotionally to the reinforcer. They may in addition react emotionally to their own achievements. These reactions are of interest with regard to the animals' capacity for self-awareness. Therefore, we devised a yoked control experiment involving the acquisition of an operant task. We aimed to identify the emotional reactions of young cattle to their own learning and to separate these from reactions to a food reward. Twelve Holstein-Friesian heifers aged 7-12 months were divided into two groups. Heifers in the experimental group were conditioned over a 14-day period to press a panel in order to open a gate for access to a food reward. For heifers in the control group, the gate opened after a delay equal to their matched partner's latency to open it. To allow for observation of the heifers' movements during locomotion after the gate had opened, there was a 15m distance in the form of a race from the gate to the food trough. The heart rate of the heifers, and their behaviour when moving along the race towards the food reward were measured. When experimental heifers made clear improvements in learning, they were more likely than on other occasions to have higher heart rates and tended to move more vigorously along the race in comparison with their controls. This experiment found some, albeit inconclusive, indication that cattle may react emotionally to their own learning improvement.
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	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6551
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	Author	McKinley, S.; Young, R.J.
	Title	The efficacy of the model-rival method when compared with operant conditioning for training domestic dogs to perform a retrieval-selection task			Type	Journal Article
	Year	2003	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	81	Issue	4	Pages	357-365
	Keywords	Dogs; Learning; Operant conditioning; Model-rival; Cognition
	Abstract	Traditionally, dogs have been trained by operant conditioning techniques; that is, dogs make a desired behavioural response and this response is reinforced by a reward such as food. This type of training is very effective in training dogs to perform basic obedience behaviours (e.g. `stay'). However, dogs are social animals and should be predisposed to learn from social stimuli. In the present study, we used a modified version of the model-rival technique that has been extensively used in experiments investigating the cognitive ability of parrots. In this technique, social stimuli are used to create in the animal an interest in the object without the use of food or other rewards. Therefore, the animal learns the name of the object (intrinsic reward) and not that the object's name means food. In this experiment we compared the learning ability of nine pet dogs to solve the same retrieval-selection task having been previously trained using operant conditioning or model-rival techniques. The retrieval-selection task was the dogs had to correctly select the commanded object to bring to the experimenter from a group of three similar objects. The results show no difference in the speeds with which the dogs solved the test--demonstrating the efficacy of the model-rival method. This is the first time that the effectiveness of the model-rival technique has been experimentally demonstrated with dogs. Furthermore, we believe that the methodology reported in this paper has applications in dog training and in experiments into dog cognition.
	Address
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	Language		Summary Language		Original Title
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2900
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	Author	Blackmore, T.L.; Foster, T.M.; Sumpter, C.E.; Temple, W.
	Title	An investigation of colour discrimination with horses (Equus caballus)			Type	Journal Article
	Year	2008	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	78	Issue	3	Pages	387-396
	Keywords	Chromatic discrimination; Colour vision; Horse; Operant
	Abstract	The ability of four horses (Equus caballus) to discriminate coloured (three shades of blue, green, red, and yellow) from grey (neutral density) stimuli, produced by back projected lighting filters, was investigated in a two response forced-choice procedure. Pushes of the lever in front of a coloured screen were occasionally reinforced, pushes of the lever in front of a grey screen were never reinforced. Each colour shade was randomly paired with a grey that was brighter, one that was dimmer, and one that approximately matched the colour in terms of brightness. Each horse experienced the colours in a different order, a new colour was started after 85% correct responses over five consecutive sessions or if accuracy showed no trend over sessions. All horses reached the 85% correct with blue versus grey, three horses did so with both yellow and green versus grey. All were above chance with red versus grey but none reached criterion. Further analysis showed the wavelengths of the green stimuli used overlapped with the yellow. The results are consistent with histological and behavioural studies that suggest that horses are dichromatic. They differ from some earlier data in that they indicate horses can discriminate yellow and blue, but that they may have deficiencies in discriminating red and green.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5336
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	Author	Rybarczyk, P.; Koba, Y.; Rushen, J.; Tanida, H.; de Passille, A.M.
	Title	Can cows discriminate people by their faces?			Type	Journal Article
	Year	2001	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	74	Issue	3	Pages	175-189
	Keywords	Dairy cows; Human-animal relationships; Discrimination; Learning; Facial recognition; Operant conditioning
	Abstract	This experiment examines the cues used by cattle to discriminate between people, particularly the role played by facial cues. We trained and tested eight Holstein cows 5 days each week for 2 months. For each cow, we used two people, a rewarder and a non-rewarder, of different size and dressed in overalls of the same colour. The operant chamber was a large box within which stood the two people. The cow could see, smell and touch each person. A lever was placed in front of each person. When the cow pushed the lever in front of the rewarder, it received 75 g of concentrate and nothing when it pushed on the other one. For each test session, the cows made 10 choices. The placement of the people was determined randomly according to the Gellerman series. The success criterion was defined as at least eight correct choices out of 10 trials for two consecutive sessions (binomial law P<0.003). During the shaping, seven cows out of eight learned to press the lever to obtain the food. The cows were then tested in a series of 10 trials with only the rewarder present. Seven out of seven cows succeeded in reaching the success criterion. In experiment 1, both the rewarder and the non-rewarder were present and standing upright at normal height and in full view of the cow. Five out of seven cows achieved the success criterion. In experiment 2, the cows could see only the faces of the two people. None of the cows were able to reach the success criterion. In experiment 3, both people were present standing up and wearing identical masks that completely covered their heads. Five cows out of five achieved the success criterion. In experiment 4, we changed the relative height of the people. Five cows out of five succeeded when the two people stood so they were of equal height but with their faces visible. However, no cows succeeded when the people were both of equal height and had their faces covered. This study suggests that cows seem to use multiple cues to discriminate between people. Cows appear able to use either body height or the face to discriminate between people but use of the face alone is more difficult when the cows cannot see the rest of the body.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	849
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	Author	Zentall, T.R.
	Title	Mental time travel in animals: a challenging question			Type	Journal Article
	Year	2006	Publication	Behavioural processes	Abbreviated Journal	Behav. Process.
	Volume	72	Issue	2	Pages	173-183
	Keywords	Animals; Behavior, Animal; Columbidae; Concept Formation; Conditioning, Operant; Imagination; Memory; Mental Recall; Planning Techniques; Rats; Time Perception; Transfer (Psychology)
	Abstract	Humans have the ability to mentally recreate past events (using episodic memory) and imagine future events (by planning). The best evidence for such mental time travel is personal and thus subjective. For this reason, it is particularly difficult to study such behavior in animals. There is some indirect evidence, however, that animals have both episodic memory and the ability to plan for the future. When unexpectedly asked to do so, animals can report about their recent past experiences (episodic memory) and they also appear to be able to use the anticipation of a future event as the basis for a present action (planning). Thus, the ability to imagine past and future events may not be uniquely human.
	Address	Department of Psychology, University of Kentucky, Lexington, KY 40506-0044, USA. zentall@uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16466863			Approved	no
	Call Number	refbase @ user @			Serial	218
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	Author	Zentall, T.R.
	Title	Support for a theory of memory for event duration must distinguish between test-trial ambiguity and actual memory loss			Type	Journal Article
	Year	1999	Publication	Journal of the experimental analysis of behavior	Abbreviated Journal	J Exp Anal Behav
	Volume	72	Issue	3	Pages	467-472
	Keywords	Animals; Behavior, Animal/physiology; Columbidae; Conditioning, Operant/physiology; Discrimination Learning/physiology; Memory/physiology; Psychological Theory; Time Factors; Time Perception/physiology
	Abstract	Staddon and Higa's (1999) trace-strength theory of timing and memory for event duration can account for pigeons' bias to “choose short” when retention intervals are introduced and to “choose long” when, following training with a fixed retention interval, retention intervals are shortened. However, it does not account for the failure of pigeons to choose short when the intertrial interval is distinct from the retention interval. That finding suggests that stimulus generalization (or ambiguity) between the intertrial interval and the retention interval may result in an effect that has been attributed to memory loss. Such artifacts must be eliminated before a theory of memory for event duration can be adequately tested.
	Address	Department of Psychology, University of Kentucky, Lexington 40506, USA. zentall@pop.uky.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5002	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:10605105			Approved	no
	Call Number	refbase @ user @			Serial	251
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