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Tavares M.C.H.; Tomaz C. |
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Title |
Working memory in capuchin monkeys (Cebus apella) |
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Journal Article |
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2002 |
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Behavioural Brain Research |
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Behav. Brain. Res. |
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131 |
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131-137 |
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refbase @ user @ |
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3486 |
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Author |
Krzak, W.E.; Gonyou, H.W.; Lawrence, L.M. |
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Title |
Wood chewing by stabled horses: diurnal pattern and effects of exercise |
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Journal Article |
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Year |
1991 |
Publication |
Journal of Animal Science |
Abbreviated Journal |
J. Anim Sci. |
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Volume |
69 |
Issue |
3 |
Pages |
1053-1058 |
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Keywords |
Animal Feed; Animals; *Behavior, Animal; *Circadian Rhythm; Female; Horses/*physiology; Male; *Mastication; *Physical Conditioning, Animal; Sodium Chloride/administration & dosage; Videotape Recording; Wood |
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Abstract |
Nine yearling horses, stabled in individual stalls, were used in a trial to determine the diurnal pattern of wood chewing and the effects of exercise on this behavior. The trial was a Latin square design conducted over three 2-wk periods during which each horse was exposed to each of the three following treatments: 1) no exercise (NE), 2) exercise after the morning feeding (AM), and 3) exercise in the afternoon (PM). Horses were fed a complete pelleted feed in the morning and both pelleted feed and long-stemmed hay in the afternoon. Exercise consisted of 45 min on a mechanical walker followed by 45 min in a paddock with bare soil. Each stall was equipped with two untreated spruce boards during each period for wood chewing. Wood chewing was evaluated by videotaping each horse for 22 h during each period, determining the weight and volume of the boards before and after each period, and by visual appraisal of the boards. Intake of trace mineralized salt was also measured. Wood chewing occurred primarily between 2200 and 1200. All measures of wood chewing were correlated when totals for the entire 6 wk were analyzed. When analysis was performed on 2-wk values, videotape results were not correlated with volume or weight loss of boards. Horses chewed more when on the NE treatment (511 s/d) than when on AM or PM (57 and 136 s/d, respectively; P less than .05). Salt intake tended to be greater for NE than for the other treatments (P less than .10).(ABSTRACT TRUNCATED AT 250 WORDS) |
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Dept. of Anim. Sci., University of Illinois, Urbana 61801 |
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English |
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0021-8812 |
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PMID:2061237 |
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refbase @ user @ |
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1949 |
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Ruid, D.B.; Paul, W.J.; Roell, B.J.; Wydeven, A.P.; Willging, R.C.; Jurewicz, R.L.; Lonsway, D.H. |
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Title |
Wolf-Human Conflicts and Management in Minnesota, Wisconsin, and Michigan |
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Book Chapter |
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2009 |
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Recovery of Gray Wolves in the Great Lakes Region of the United States: An Endangered Species Success Story |
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279-295 |
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Springer New York |
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New York, NY |
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Wydeven, A.P.; Van Deelen, T.R.; Heske, E.J. |
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978-0-387-85952-1 |
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Equine Behaviour @ team @ Ruid2009 |
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6577 |
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Author |
van Duyne, C.; Ras, E.; de Vos, A.E.W.; de Boer, W.F.; Henkens, R.J.H.G.; Usukhjargal, D. |
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Title |
Wolf Predation Among Reintroduced Przewalski Horses in Hustai National Park, Mongolia |
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Journal Article |
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2009 |
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Journal of Wildlife Management |
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Journal of Wildlife Management |
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73 |
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6 |
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836-843 |
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The Wildlife Society |
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0022-541x |
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doi: 10.2193/2008-027 |
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Equine Behaviour @ team @ |
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5776 |
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Harrington, F.H.; Mech, L.D. |
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Wolf howling and its role in territory maintenance |
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1979 |
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Behaviour |
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68 |
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Equine Behaviour @ team @ Harrington1979 |
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6455 |
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Author |
Capitani, C.; Chynoweth, M.; Kusak, J.; Çoban, E.; Sekercioglu, Ç.H. |
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Title |
Wolf diet in an agricultural landscape of north-eastern Turkey |
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Journal Article |
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2016 |
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Mammalia |
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Mammalia |
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80 |
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3 |
Pages |
329-334 |
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Mammalia |
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80 |
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3 |
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Equine Behaviour @ team @ |
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6687 |
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Author |
LANG EM, L.E.H.M.A.N.N.E. von |
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Wildesel in Vergangenheit und Gegenwart |
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Journal Article |
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1972 |
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Zoologische Garten |
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Zool. Garten. |
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41 |
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157-167 |
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from Professor Hans Klingels Equine Reference List |
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no |
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1338 |
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Author |
Gonyou, H.W. |
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Title |
Why the study of animal behavior is associated with the animal welfare issue |
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Journal Article |
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Year |
1994 |
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Journal of Animal Science |
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J. Anim Sci. |
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72 |
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8 |
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2171-2177 |
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Equine Behaviour @ team @ |
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2931 |
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Author |
Hunt, G.R.; Gray R.D.; Taylor, A.H. |
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Title |
Why is tool use rare in animals? |
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Book Whole |
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2013 |
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Tool Use in Animals: Cognition and Ecology |
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Cambridge University Press |
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Cambridge, MA. |
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anz C, Call J, Boesch C |
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Equine Behaviour @ team @ |
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6658 |
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Author |
Hintze, S.; Smith, S.; Patt, A.; Bachmann, I.; Würbel, H. |
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Title |
What eye wrinkles in horses tell us about their emotional state |
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Conference Article |
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2015 |
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Proceedings of the 3. International Equine Science Meeting |
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Proc. 3. Int. Equine. Sci. Mtg |
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Wrinkles above the eye ball are common in domestic horses but may differ in number and shape both between and within individuals. They are caused by contraction of the inner eye brow raiser, and some people working with horses call them “worry wrinkles”, considering them to reflect emotional states. However, as yet no study has formally investigated the relationship between eye wrinkles and emotional state in horses.
The aim of the present study was to induce states of different emotional valence and to assess whether positive emotional states would reduce the expression of eye wrinkles while negative emotional states would increase it. Sixteen horses were confronted in a balanced order with two presumably positively and two negatively valenced situations each. Positive situations included anticipation of a food reward (FA) and petting (P), negative situations included food competition (FC) and waving a plastic bag (PB). Each situation lasted for 60s (TRT) and was preceded by a 60s control phase (CON). Throughout CON and TRT pictures of the eyes were taken, and for each horse four pictures per situation (FA, P, FC, PB) and phase (CON and TRT) were randomly selected (n = 512) and scored in random order and blind to treatment for six outcome variables: overall impression (qualitative), number, angle and markedness of eye wrinkles, presence of eye white, and shape of eye lid.
Data were analysed separately for the right and left eye using linear mixed effects models (angle, number), generalised linear mixed models (eye white, markedness), and ordered logistic regression (qualitative, shape of eye lid), with “situation” (FA, P, FC, PB), “phase” (CON, TRT) and their two-way interaction as fixed effects.
Expression of eye wrinkles did not vary consistently across “situation” and “phase”. Independent of phase, eye white appeared less frequently during P than during FA (z=-3.15, p=0.009), FC (z=-2.94, p=0.02), and PB (z=4.17, p<0.001) in the left eye and during PB (z=4.10, p 0.001) in the right eye. Similarly, wrinkles were less marked during P compared to the other situations in the left eye (FA: z=3.15, p=0.009; FC: z=-2.94, p=0.017; PB: z=4.17, p<0.001) and compared to PB in the right eye (z=4.10, p=0.001), while no differences between situations occurred in number of wrinkles, overall impression and shape of eye lid for both eyes. Consistent with our hypothesis, P induced relaxation of the underlying muscle in the right eye resulting in a wider angle compared to its control phase (interaction situation*phase: F3,10=3.71, p=0.055; post-hoc comparison: z=-3.57, p=0.009), while FC induced muscle contraction, resulting in a sharper angle in the left eye (interaction situation*phase: F3,11=6.57, p=0.011; z=3.73, p=0.005).
We conclude that emotional valence may affect characteristics of eye winkle expression in horses which might therefore be a promising indicator of horses’ emotional states, but further research is needed to validate the relevant outcome variables. |
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eye wrinkles, emotional valence, positive and negative emotions, welfare assessment |
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Corporate Author |
Hintze, S. |
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Equine Behaviour @ team @ |
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5875 |
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