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Author Baer, K.L.; Potter, G.D.; Friend, T.H.; Beaver, B.V. url  doi
openurl 
  Title Observation effects on learning in horses Type Journal Article
  Year 1983 Publication Applied Animal Ethology Abbreviated Journal Appl. Animal. Ethol.  
  Volume 11 Issue 2 Pages 123-129  
  Keywords  
  Abstract Sixteen horses, divided into 2 groups of 8, were used to study observational learning in horses. One group served as controls while the other group served as the treated group (observers). Observers were allowed to watch a correctly performed discrimination task for 5 days prior to testing their learning response using the same task. Discrimination testing was conducted on all horses daily for 14 days, with criterion set at 7 out of 8 responses correct with the last 5 consecutively correct. The maximum number of trials performed without reaching criterion was limited to 20 per day. Mean trials to criteria (MT) by group were: control, 11.25; observer, 10.70. Mean error (ME) scores were: control, 2.37; observer, 2.02. Average initial discrimination error scores were 11.13 for control and 10.38 for observers (P < 0.10). Asymptote was reached by Day 8 for both control and observer groups. Analysis of variance with repeated measures showed an extreme-day effect indicative of learning (P < 0.01), with non-significant differences in learning rate between experimental groups. Whether the initial ability of the horses to perform a discrimination learning task was enhanced by observation of other horses' performance of that task was not obvious from these data.  
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  Notes Approved no  
  Call Number refbase @ user @ Serial 726  
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Author Horner, V.; Whiten, A. doi  openurl
  Title Learning from others' mistakes limits on understanding a trap-tube task by young chimpanzees (Pan troglodytes) and children (Homo sapiens) Type Journal Article
  Year 2007 Publication Journal of comparative psychology Abbreviated Journal J Comp Psychol  
  Volume 121 Issue 1 Pages 12-21  
  Keywords  
  Abstract A trap-tube task was used to determine whether chimpanzees (Pan troglodytes) and children (Homo sapiens) who observed a model's errors and successes could master the task in fewer trials than those who saw only successes. Two- to 7-year-old chimpanzees and 3- to 4-year-old children did not benefit from observing errors and found the task difficult. Two of the 6 chimpanzees developed a successful anticipatory strategy but showed no evidence of representing the core causal relations involved in trapping. Three- to 4-year-old children showed a similar limitation and tended to copy the actions of the demonstrator, irrespective of their causal relevance. Five- to 6-year-old children were able to master the task but did not appear to be influenced by social learning or benefit from observing errors.  
  Address Centre for Social Learning and Cognitive Evolution, School of Psychology, University of St Andrews, Fife, Scotland, UK. vhorner@rmy.emory.edu  
  Corporate Author Thesis  
  Publisher Place of Publication Washington, D.C. : 1983 Editor  
  Language English Summary Language Original Title  
  Series Editor (down) Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0735-7036 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:17324071 Approved yes  
  Call Number refbase @ user @ Serial 728  
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Author Whiten, A.; Horner, V.; Litchfield, C.A.; Marshall-Pescini, S. url  doi
openurl 
  Title How do apes ape? Type Journal Article
  Year 2004 Publication Learning & Behavior Abbreviated Journal Learn. Behav.  
  Volume 32 Issue 1 Pages 36-52  
  Keywords Adaptation, Psychological; Animals; Behavior, Animal; Hominidae/*psychology; *Imitative Behavior; Imprinting (Psychology); *Learning; Psychological Theory; *Social Environment; *Social Facilitation  
  Abstract In the wake of telling critiques of the foundations on which earlier conclusions were based, the last 15 years have witnessed a renaissance in the study of social learning in apes. As a result, we are able to review 31 experimental studies from this period in which social learning in chimpanzees, gorillas, and orangutans has been investigated. The principal question framed at the beginning of this era, Do apes ape? has been answered in the affirmative, at least in certain conditions. The more interesting question now is, thus, How do apes ape? Answering this question has engendered richer taxonomies of the range of social-learning processes at work and new methodologies to uncover them. Together, these studies suggest that apes ape by employing a portfolio of alternative social-learning processes in flexibly adaptive ways, in conjunction with nonsocial learning. We conclude by sketching the kind of decision tree that appears to underlie the deployment of these alternatives.  
  Address Centre for Social Learning and Cognitive Evolution, Scottish Primate Research Group, School of Psychology, University of St. Andrews, St. Andrews, Fife, Scotland. a.whiten@st-and.ac.uk  
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  Language English Summary Language Original Title  
  Series Editor (down) Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1543-4494 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:15161139 Approved no  
  Call Number refbase @ user @ Serial 734  
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Author Caldwell, C.A.; Whiten, A. doi  openurl
  Title Testing for social learning and imitation in common marmosets, Callithrix jacchus, using an artificial fruit Type Journal Article
  Year 2004 Publication Animal cognition Abbreviated Journal Anim. Cogn.  
  Volume 7 Issue 2 Pages 77-85  
  Keywords Animals; *Association Learning; Callithrix/*psychology; Discrimination Learning; *Feeding Behavior; Female; Food Preferences; Fruit; *Imitative Behavior; Male; *Social Behavior; Social Environment  
  Abstract We tested for social learning and imitation in common marmosets using an artificial foraging task and trained conspecific demonstrators. We trained a demonstrator marmoset to open an artificial fruit, providing a full demonstration of the task to be learned. Another marmoset provided a partial demonstration, controlling for stimulus enhancement effects, by eating food from the outside of the apparatus. We thus compared three observer groups, each consisting of four animals: those that received the full demonstration, those that received the partial demonstration, and a control group that saw no demonstration prior to testing. Although none of the observer marmosets succeeded in opening the artificial fruit during the test periods, there were clear effects of demonstration type. Those that saw the full demonstration manipulated the apparatus more overall, whereas those from the control group manipulated it the least of the three groups. Those from the full-demonstration group also contacted the particular parts of the artificial fruit that they had seen touched (localised stimulus enhancement) to a greater extent than the other two groups. There was also an interaction between the number of hand and mouth touches made to the artificial fruit for the full- and partial-demonstration groups. Whether or not these data represent evidence for imitation is discussed. We also propose that the clear differences between the groups suggest that social learning mechanisms provide real benefits to these animals in terms of developing novel food-processing skills analogous to the one presented here.  
  Address Centre for Social Learning and Cognitive Evolution and Scottish Primate Research Group, University of St Andrews, KY16 9JU, St Andrews, Fife, Scotland. C.A.Caldwell@exeter.ac.uk  
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  Language English Summary Language Original Title  
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  Series Volume Series Issue Edition  
  ISSN 1435-9448 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:15069606 Approved no  
  Call Number refbase @ user @ Serial 735  
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Author Horowitz, A.C. doi  openurl
  Title Do humans ape? Or do apes human? Imitation and intention in humans (Homo sapiens) and other animals Type Journal Article
  Year 2003 Publication Journal of comparative psychology Abbreviated Journal J Comp Psychol  
  Volume 117 Issue 3 Pages 325-336  
  Keywords Adolescent; Adult; Animals; *Appetitive Behavior; Attention; Child, Preschool; Concept Formation; Female; Humans; *Imitative Behavior; Male; Motivation; Pan troglodytes/*psychology; *Problem Solving; *Psychomotor Performance; Reaction Time; Species Specificity  
  Abstract A. Whiten, D. M. Custance, J.-C. Gomez, P. Teixidor, and K. A. Bard (1996) tested chimpanzees' (Pan troglodytes) and human children's (Homo sapiens) skills at imitation with a 2-action test on an “artificial fruit.” Chimpanzees imitated to a restricted degree; children were more thoroughly imitative. Such results prompted some to assert that the difference in imitation indicates a difference in the subjects' understanding of the intentions of the demonstrator (M. Tomasello, 1996). In this experiment, 37 adult human subjects were tested with the artificial fruit. Far from being perfect imitators, the adults were less imitative than the children. These results cast doubt on the inference from imitative performance to an ability to understand others' intentions. The results also demonstrate how any test of imitation requires a control group and attention to the level of behavioral analysis.  
  Address Department of Cognitive Science, University of California, San Diego, CA, USA. ahorowitz@crl.ucsd.edu  
  Corporate Author Thesis  
  Publisher Place of Publication Washington, D.C. : 1983 Editor  
  Language English Summary Language Original Title  
  Series Editor (down) Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0735-7036 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:14498809 Approved yes  
  Call Number refbase @ user @ Serial 736  
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Author Castles, D.L.; Whiten, A.; Aureli, F. url  doi
openurl 
  Title Social anxiety, relationships and self-directed behaviour among wild female olive baboons Type Journal Article
  Year 1999 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.  
  Volume 58 Issue 6 Pages 1207-1215  
  Keywords  
  Abstract Self-directed behaviour (SDB) can be used as a behavioural indicator of stress and anxiety in nonhuman primates (Maestripieri et al. 1992, Animal Behaviour, 44, 967-979). We investigated the effect of nearest neighbours' relative dominance status on the SDB of sexually mature female olive baboons, Papio anubis. When the animal nearest to (within 5 m of) a female was a dominant individual, SDB rates (a combined measure of self-scratching, self-grooming, self-touching, body shaking and yawning) increased by ca. 40% over those observed when the nearest neighbour was a subordinate. The results indicate that (1) SDB can be used as a measure of uncertainty during the social interactions of cercopithecine primates and (2) as there was considerable variation in SDB response according to the nature of the dominant individual, SDB can be used to assess relationship security (i.e. the perceived predictability of a relationship for one partner). Finally, in combination with measures of affiliation rate, SDB may provide insight into relationship value.  
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  Call Number refbase @ user @ Serial 745  
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Author Whiten, A.; Custance, D.M.; Gomez, J.C.; Teixidor, P.; Bard, K.A. doi  openurl
  Title Imitative learning of artificial fruit processing in children (Homo sapiens) and chimpanzees (Pan troglodytes) Type Journal Article
  Year 1996 Publication Journal of comparative psychology (Washington, D.C. : 1983) Abbreviated Journal J Comp Psychol  
  Volume 110 Issue 1 Pages 3-14  
  Keywords Animals; Child, Preschool; Discrimination Learning; Female; Food Preferences/*psychology; *Fruit; Humans; *Imitative Behavior; Male; Mental Recall; Pan troglodytes/*psychology; Social Environment  
  Abstract Observational learning in chimpanzees and young children was investigated using an artificial fruit designed as an analog of natural foraging problems faced by primates. Each of 3 principal components could be removed in 2 alternative ways, demonstration of only one of which was watched by each subject. This permitted subsequent imitation by subjects to be distinguished from stimulus enhancement. Children aged 2-4 years evidenced imitation for 2 components, but also achieved demonstrated outcomes through their own techniques. Chimpanzees relied even more on their own techniques, but they did imitate elements of 1 component of the task. To our knowledge, this is the first experimental evidence of chimpanzee imitation in a functional task designed to simulate foraging behavior hypothesized to be transmitted culturally in the wild.  
  Address Scottish Primate Research Group, University of St. Andrews, Fife, Scotland. aw2@st-andrews.ac.uk  
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  Language English Summary Language Original Title  
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  Series Volume Series Issue Edition  
  ISSN 0735-7036 ISBN Medium  
  Area Expedition Conference  
  Notes PMID:8851548 Approved no  
  Call Number refbase @ user @ Serial 744  
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Author Byrne, R.W. url  openurl
  Title Imitation of novel complex actions: What does the evidence from animals mean? Type Book Chapter
  Year 2002 Publication Advances in the Study of Behavior Abbreviated Journal Adv Stud Behav  
  Volume 31 Issue Pages 77-105  
  Keywords  
  Abstract Summary Underlying the various behaviors that are classified as imitation, there may be several distinct mechanisms, differing in adaptive function, cognitive basis, and computational power. Experiments reporting “true motor imitation” in animals do not as yet give evidence of production learning by imitation; instead, contextual imitation can explain their data, and this can be explained by a simple mechanism (response facilitation) which matches known neural findings. When imitation serves a function in social mimicry, which applies to a wide range of phenomena from neonatal imitation in humans and great apes to pair-bonding in some bird species, the fidelity of the behavioral match is crucial. Learning of novel behavior can potentially be achieved by matching the outcome of a model's action, and it is argued that vocal imitation by birds is a clear example of this method (which is sometimes called emulation). Alternatively, the behavior itself may be perceived in terms of actions that the observer can perform, and thus it may be copied. If the imitation is linear and stringlike (action level), following the surface form rather than the underlying plan, then its utility for learning new instrumental methods is limited. However, the underlying plan of hierarchically organized behavior is visible in output behavior, in subtle but detectable ways, and imitation could instead be based on this organization (program level), extracted automatically by string parsing. Currently, the most likely candidates for such capacities are all great apes. It is argued that this ability to perceive the underlying plan of action, in addition to allowing highly flexible imitation of novel instrumental methods, may have resulted in the competence to understand the intentions (theory of mind) of others.  
  Address  
  Corporate Author Thesis  
  Publisher Academic Press Place of Publication San Diego Editor Snowdon, C. T.; Roper, T. J.;Rosenblatt,J. S.  
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  Notes Approved no  
  Call Number refbase @ user @ Serial 746  
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Author Zentall, T.R url  doi
openurl 
  Title Imitation In Animals: Evidence, Function, And Mechanisms Type Journal Article
  Year 2001 Publication Cybernetics and Systems Abbreviated Journal Cybern Syst  
  Volume 32 Issue Pages 53-96  
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  Abstract The terms sociallearning and social influence have been used descriptively and theoretically to characterize a broad range of animal behavior from physical antipredatory adaptations such as eye spots, which are totally under genetic control, to the human capacity for the exaggeration of individual characteristics, known as caricature, which are largely under cognitive control. In the present review, the various forms of social influence and social learning are identified and distinghished from imitation, a term that generally has been reserved for behavioral matching that cannot be accounted for using simpler specifically predisposed, motivational, or learning mechanisms. It is suggested that much of the ambiguity in the literature concerning the various forms of social learning can be attributed to the distinction between the function of a behavior and the mechanisms responsible for its occurrence. Finally, the various mechanisms that have been proposed to account for imitative learning are presented and an attempt is made to evaluate them.  
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  Notes Approved no  
  Call Number refbase @ user @ Serial 747  
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Author Berger, J. doi  openurl
  Title Organizational systems and dominance in feral horses in the Grand Canyon Type Journal Article
  Year 1977 Publication Behavioral Ecology and Sociobiology Abbreviated Journal Behav. Ecol. Sociobiol.  
  Volume 2 Issue 2 Pages 131-146  
  Keywords  
  Abstract 1. Several aspects of the behavioral ecology of feral horses (Equus caballus) were studied in Grand Canyon, Arizona, USA. Most bands contained three to five horses that included one stallion and his harem. Males that did not obtain a harem remained solitary. Throughout the study bands remained stable in composition.

2. Home ranges for all bands decreased in size in successive warm months, probably due to increased ambient temperature and drought. This resulted in greater utilization of spring areas that led to increased interband confrontation and agonistic display.

3. Territoriality was not observed in individual horses or bands, but bands hierarchial in both inter- and intraband structures. Interband stallion dominance was reinforced through posturing and fighting. Intraband hierarchies, as determined by dominance coefficients, were independent of individual size in three of four bands.

4. Indexes of nervousness (NER), calculated while horses were drinking, showed that stallions were less nervous than mares. A low NER was correlated with individuals leading toward drinking areas, whereas a high NER existed in individuals initiating flight although no single horse acted consistently as a leader.

5. Diurnal activity patterns were correlated with ambient temperatures.
 
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  Notes Approved no  
  Call Number refbase @ user @ Serial 748  
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