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Author Bernauer, K.; Kollross, H.; Schuetz, A.; Farmer, K.; Krueger, K. url  doi
openurl 
  Title How do horses (Equus caballus) learn from observing human action? Type Journal Article
  Year 2020 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume 23 Issue Pages 1-9  
  Keywords  
  Abstract A previous study demonstrated that horses can learn socially from observing humans, but could not draw any conclusions about the social learning mechanisms. Here we develop this by showing horses four different human action sequences as demonstrations of how to press a button to open a feed box. We tested 68 horses aged between 3 and 12 years. 63 horses passed the habituation phase and were assigned either to the group Hand Demo (N = 13) for which a kneeling person used a hand to press the button, Head Demo (N = 13) for which a kneeling person used the head, Mixed Demo (N = 12) for which a squatting person used both head and hand, Foot Demo (N = 12) in which a standing person used a foot, or No Demo (N = 13) in which horses did not receive a demonstration. 44 horses reached the learning criterion of opening the feeder twenty times consecutively, 40 of these were 75% of the Demo group horses and four horses were 31% of the No Demo group horses. Horses not reaching the learning criterion approached the human experimenters more often than those who did. Significantly more horses used their head to press the button no matter which demonstration they received. However, in the Foot Demo group four horses consistently preferred to use a hoof and two switched between hoof and head use. After the Mixed Demo the horses' actions were more diverse. The results indicate that only a few horses copy behaviours when learning socially from humans. A few may learn through observational conditioning, as some appeared to adapt to demonstrated actions in the course of reaching the learning criterion. Most horses learn socially through enhancement, using humans to learn where, and which aspect of a mechanism has to be manipulated, and by applying individual trial and error learning to reach their goal.  
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  Series Volume Series Issue Edition  
  ISSN 1435-9456 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Bernauer2019 Serial 6590  
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Author Brinkmann, L.; Gerken, M.; Riek, A. url  doi
openurl 
  Title Effect of long-term feed restriction on the health status and welfare of a robust horse breed, the Shetland pony (Equus ferus caballus) Type Journal Article
  Year 2013 Publication Research in Veterinary Science Abbreviated Journal Res. Vet. Sci.  
  Volume 94 Issue 3 Pages 826-831  
  Keywords Animal welfare; Blood parameter; Extensive housing; Feed restriction; Horse; Winter conditions  
  Abstract Outdoor group housing is increasingly recognized as an appropriate housing system for domesticated horses. The objective of this study was therefore to investigate the effect of potential feed shortage in semi-natural horse keeping systems in winter on animal health and welfare. In 10 female Shetland ponies blood concentrations (NEFA, total protein (TP), total bilirubin (TB), beta-hydroxybutyrate (BHB) and thyroxine (T4)), body mass and the body condition score (BCS) were monitored for 7months including a 4months period of feed restriction in five of the 10 ponies. Restrictively fed animals lost 18.4±2.99% of their body mass and the BCS decreased by 2.2±0.8 points (BCS scale: 0=emaciated, 5=obese). Feed restriction led to a continuous increase in TB (P<0.001) and NEFA (P<0.01) concentrations compared to control ponies. The TP and BHB values only differed at the end of the trial with lower concentrations in restricted fed mares (P<0.05). Feed restriction had no effect on thyroxine concentrations. TB concentrations in the feed restricted group were out of the reference range during the entire feeding trial. The increased NEFA concentrations in feed restricted compared to control ponies suggest that fat was mobilized. The BCS, as well as plasma NEFA and TB concentrations were good indicators for a rapid detection of possible health problems caused by undernourishment in horses when kept under semi-natural conditions. In contrast, blood parameters of the control animals were within the reference ranges, suggesting that a year round outdoor housing with additional feed supply is an adequate housing system for a robust horse breed like the Shetland pony.  
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  Publisher (down) Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0034-5288 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6601  
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Author Baragli, P.; Scopa, C.; Maglieri, V.; Palagi, E. url  doi
openurl 
  Title If horses had toes: demonstrating mirror self recognition at group level in Equus caballus Type Journal Article
  Year 2021 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume Issue Pages  
  Keywords  
  Abstract Mirror self-recognition (MSR), investigated in primates and recently in non-primate species, is considered a measure of self-awareness. Nowadays, the only reliable test for investigating MSR potential skills consists in the untrained response to a visual body mark detected using a reflective surface. Here, we report the first evidence of MSR at group level in horses, by facing the weaknesses of methodology present in a previous pilot study. Fourteen horses were used in a 4-phases mirror test (covered mirror, open mirror, invisible mark, visible colored mark). After engaging in a series of contingency behaviors (looking behind the mirror, peek-a-boo, head and tongue movements), our horses used the mirror surface to guide their movements towards their colored cheeks, thus showing that they can recognize themselves in a mirror. The analysis at the group level, which 'marks' a turning point in the analytical technique of MSR exploration in non-primate species, showed that horses spent a longer time in scratching their faces when marked with the visible mark compared to the non-visible mark. This finding indicates that horses did not see the non-visible mark and that they did not touch their own face guided by the tactile sensation, suggesting the presence of MSR in horses. Although a heated debate on the binary versus gradualist model in the MSR interpretation exists, recent empirical pieces of evidence, including ours, indicate that MSR is not an all-or-nothing phenomenon that appeared once in phylogeny and that a convergent evolution mechanism can be at the basis of its presence in phylogenetically distant taxa.  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1435-9456 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Baragli2021 Serial 6631  
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Author Jarausch, A.; Harms, V.; Kluth, G.; Reinhardt, I.; Nowak, C. url  doi
openurl 
  Title How the west was won: genetic reconstruction of rapid wolf recolonization into Germany's anthropogenic landscapes Type Journal Article
  Year 2021 Publication Heredity Abbreviated Journal Heredity  
  Volume Issue Pages  
  Keywords  
  Abstract Following massive persecution and eradication, strict legal protection facilitated a successful reestablishment of wolf packs in Germany, which has been ongoing since 2000. Here, we describe this recolonization process by mitochondrial DNA control-region sequencing, microsatellite genotyping and sex identification based on 1341 mostly non-invasively collected samples. We reconstructed the genealogy of German wolf packs between 2005 and 2015 to provide information on trends in genetic diversity, dispersal patterns and pack dynamics during the early expansion process. Our results indicate signs of a founder effect at the start of the recolonization. Genetic diversity in German wolves is moderate compared to other European wolf populations. Although dispersal among packs is male-biased in the sense that females are more philopatric, dispersal distances are similar between males and females once only dispersers are accounted for. Breeding with close relatives is regular and none of the six male wolves originating from the Italian/Alpine population reproduced. However, moderate genetic diversity and inbreeding levels of the recolonizing population are preserved by high sociality, dispersal among packs and several immigration events. Our results demonstrate an ongoing, rapid and natural wolf population expansion in an intensively used cultural landscape in Central Europe.  
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  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1365-2540 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Jarausch2021 Serial 6638  
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Author Bruns, A.; Waltert, M.; Khorozyan, I. url  doi
openurl 
  Title The effectiveness of livestock protection measures against wolves (Canis lupus) and implications for their co-existence with humans Type Journal Article
  Year 2020 Publication Global Ecology and Conservation Abbreviated Journal  
  Volume 21 Issue Pages e00868  
  Keywords Carnivore; Depredation; Efficiency; Germany; Intervention; Predator  
  Abstract Wolves (Canis lupus) can kill domestic livestock resulting in intense conflicts with humans. Damage to livestock should be reduced to facilitate human-wolf coexistence and ensure positive outcomes of conservation efforts. Current knowledge on the effectiveness of livestock protection measures from wolves is limited and scattered in the literature. In this study, we compiled a dataset of 30 cases describing the application of 11 measures of protecting cattle and smaller livestock against wolves, estimated their effectiveness as a relative risk of damage, and identified the best measures for damage reduction. We found that: (1) lethal control and translocation were less effective than other measures, (2) deterrents, especially fladry which is a fence with ropes marked by hanging colored flags that sway in the wind and provide a visual warning signal, were more effective than guarding dogs; (3) deterrents, fencing, calving control and herding were very effective, but the last two measures included only one case each; and (4) protection of cattle was more effective than that of small stock (sheep and goats, or sheep only) and mixed cattle and small stock. In all of these cases, the relative risk of damage was reduced by 50-100%. Considering Germany as an example of a country with a recovering wolf population and escalating human-wolf conflicts, we suggest electric fences and electrified fladry as the most promising measures, which under suitable conditions can be accompanied by well-trained livestock guarding dogs, and the temporary use of deterrents during critical periods such as calving and lambing seasons. Further research in this field is of paramount importance to efficiently mitigate human-wolf conflicts.  
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  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 2351-9894 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6641  
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Author Krueger, K.; Esch, L.; Farmer, K.; Marr, I. doi  openurl
  Title Basic Needs in Horses?--A Literature Review Type Magazine Article
  Year 2021 Publication Animals Abbreviated Journal Animals  
  Volume 11 Issue 6 Pages 1798  
  Keywords abnormal behaviour; active responses; horse; movement; passive responses; roughage; stress; social contact  
  Abstract Every animal species has particular environmental requirements that are essential for its welfare, and when these so-called “basic needs” are not fulfilled, the animals suffer. The basic needs of horses have been claimed to be social contact, social companionship, free movement and access to roughage. To assess whether horses suffer when one or more of the four proposed basic needs are restricted, we examined several studies (n = 38) that reported behavioural and physiological reactions to these restrictions. We assigned the studies according to the four types of responses investigated: (a) Stress, (b) Active, (c) Passive, and (d) Abnormal Behaviour. Furthermore, the number of studies indicating that horses reacted to the restrictions were compared with the number of studies reporting no reaction. The limited number of studies available on single management restrictions did not allow conclusions to be drawn on the effect of each restriction separately, especially in the case of social companionship. However, when combinations of social contact, free movement and access to roughage were restricted, many of the horses had developed responses consistent with suffering. Passive Responses, indicating acute suffering, and Abnormal Behaviour, indicating suffering currently or at some time in the past, were especially clearly demonstrated. This provides further evidence of the usefulness of assessing behavioural parameters in combination with physiological measurements when evaluating horse welfare. This meta-analysis of the literature confirms that it is justified to claim that social contact, free movement and access to roughage are basic needs in horses.  
  Address  
  Corporate Author Thesis  
  Publisher (down) Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title Animals  
  Series Volume 11 Series Issue 6 Edition  
  ISSN 2076-2615 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6645  
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Author Bergmüller, R.; Taborsky, M. url  doi
openurl 
  Title Animal personality due to social niche specialisation Type Journal Article
  Year 2010 Publication Trends in Ecology & Evolution Abbreviated Journal  
  Volume 25 Issue 9 Pages 504-511  
  Keywords  
  Abstract The existence of 'animal personality', i.e. consistent individual differences in behaviour across time and contexts, is an evolutionary puzzle that has recently generated considerable research interest. Although social factors are generally considered to be important, it is as yet unclear how they might select for personality. Drawing from ecological niche theory, we explore how social conflict and alternative social options can be key factors in the evolution and development of consistent individual differences in behaviour. We discuss how animal personality research might benefit from insights into the study of alternative tactics and illustrate how selection can favour behavioural diversification and consistency due to fitness benefits resulting from conflict reduction among social partners.  
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  Corporate Author Thesis  
  Publisher (down) Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0169-5347 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6646  
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Author Baumgartner, M.; Boisson, T.; Erhard, M.H.; Zeitler-Feicht, M.H. doi  openurl
  Title Common Feeding Practices Pose A Risk to the Welfare of Horses When Kept on Non-Edible Bedding Type Journal Article
  Year 2020 Publication Animals Abbreviated Journal Animals  
  Volume 10 Issue Pages 441  
  Keywords horse behaviour; feed intake pause; bedding; welfare indicator; feeding practices; roughage; horse welfare; individual housing system  
  Abstract During the evolution of the horse, an extended period of feed intake, spread over the entire 24-h period, determined the horsesâ&#65533;&#65533; behaviour and physiology. Horses will not interrupt their feed intake for more than 4 h, if they have a choice. The aim of the present study was to investigate in what way restrictive feeding practices (non ad libitum) affect the horsesâ&#65533;&#65533; natural feed intake behaviour. We observed the feed intake behaviour of 104 horses on edible (n = 30) and non-edible bedding (n = 74) on ten different farms. We assessed the duration of the forced nocturnal feed intake interruption of horses housed on shavings when no additional roughage was available. Furthermore, we comparatively examined the feed intake behaviour of horses housed on edible versus non-edible bedding. The daily restrictive feeding of roughage (2 times a day: n = 8; 3 times a day: n = 2), as it is common in individual housing systems, resulted in a nocturnal feed intake interruption of more than 4 hours for the majority (74.32%, 55/74) of the horses on shavings (8:50 ± 1:25 h, median: 8:45 h, minimum: 6:45 h, maximum: 13:23 h). In comparison to horses on straw, horses on shavings paused their feed intake less frequently and at a later latency. Furthermore, they spent less time on consuming the evening meal than horses on straw. Our results of the comparison of the feed-intake behaviour of horses on edible and non-edible bedding show that the horsesâ&#65533;&#65533; ethological feeding needs are not satisfied on non-edible bedding. If the horses accelerate their feed intake (also defined as â&#65533;&#65533;rebound effectâ&#65533;&#65533;), this might indicate that the horsesâ&#65533;&#65533; welfare is compromised. We conclude that in addition to the body condition score, the longest duration of feed intake interruption (usually in the night) is an important welfare indicator of horses that have limited access to roughage.  
  Address  
  Corporate Author Thesis  
  Publisher (down) Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title Animals  
  Series Volume 10 Series Issue 3 Edition  
  ISSN 2076-2615 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6647  
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Author Trösch, M.; Pellon, S.; Cuzol, F.; Parias, C.; Nowak, R.; Calandreau, L.; Lansade, L. url  doi
openurl 
  Title Horses feel emotions when they watch positive and negative horse-human interactions in a video and transpose what they saw to real life Type Journal Article
  Year 2020 Publication Animal Cognition Abbreviated Journal Anim. Cogn.  
  Volume 23 Issue 4 Pages 643-653  
  Keywords  
  Abstract Animals can indirectly gather meaningful information about other individuals by eavesdropping on their third-party interactions. In particular, eavesdropping can be used to indirectly attribute a negative or positive valence to an individual and to adjust one's future behavior towards that individual. Few studies have focused on this ability in nonhuman animals, especially in nonprimate species. Here, we investigated this ability for the first time in domestic horses (Equus caballus) by projecting videos of positive and negative interactions between an unknown human experimenter (a “positive” experimenter or a “negative” experimenter) and an actor horse. The horses reacted emotionally while watching the videos, expressing behavioral (facial expressions and contact-seeking behavior) and physiological (heart rate) cues of positive emotions while watching the positive video and of negative emotions while watching the negative video. This result shows that the horses perceived the content of the videos and suggests an emotional contagion between the actor horse and the subjects. After the videos were projected, the horses took a choice test, facing the positive and negative experimenters in real life. The horses successfully used the interactions seen in the videos to discriminate between the experimenters. They touched the negative experimenter significantly more, which seems counterintuitive but can be interpreted as an appeasement attempt, based on the existing literature. This result suggests that horses can indirectly attribute a valence to a human experimenter by eavesdropping on a previous third-party interaction with a conspecific.  
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  Corporate Author Thesis  
  Publisher (down) Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 1435-9456 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Trösch2020 Serial 6649  
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Author Ahrendt, L.P.; Labouriau, R.; Malmkvist, J.; Nicol, C.J.; Christensen, J.W. url  doi
openurl 
  Title Development of a standard test to assess negative reinforcement learning in horses Type Journal Article
  Year 2015 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 169 Issue Pages 38-42  
  Keywords Algometry; Horse behaviour; Learning performance; Operant conditioning; Pressure-release; Horse training  
  Abstract Most horses are trained by negative reinforcement. Currently, however, no standardised test for evaluating horses' negative reinforcement learning ability is available. The aim of this study was to develop an objective test to investigate negative reinforcement learning in horses. Twenty-four Icelandic horses (3 years old) were included in this study. The horses were tested in a pressure-release task on three separate days with 10, 7 and 5 trials on each side, respectively. Each trial consisted of pressure being applied on the hindquarter with an algometer. The force of the pressure was increased until the horse moved laterally away from the point of pressure. There was a significant decrease in required force over trials on the first test day (P<0.001), but not the second and third day. The intercepts on days 2 and 3 differed significantly from day 1 (P<0.001), but not each other. Significantly stronger force was required on the right side compared to the left (P<0.001), but there was no difference between first and second side tested (P=0.56). Individual performance was evaluated by median-force and the change in force over trials on the first test day. These two measures may explain different characteristics of negative reinforcement learning. In conclusion, this study presents a novel, standardised test for evaluating negative reinforcement learning ability in horses.  
  Address  
  Corporate Author Thesis  
  Publisher (down) Place of Publication Editor  
  Language Summary Language Original Title  
  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0168-1591 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 6650  
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