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Sakura O, & Matsuzawa T. (1991). Flexibility of wild chimpanzees nut-cracking behavior using stone hammers and anvils: an experimental analysis. Ethology, 87, 237.
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Hampton, R. R. (2001). Animal Minds: Beyond Cognition to Consciousness. Ethology, 107, 1055–1056.
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Houpt, K. A. (1980). Review of some research areas of applied and theoretical interest in domestic animal behavior. Appl. Animal. Ethol., 6(2), 111–119.
Abstract: There are numerous areas worthy of study in the field of domestic animal behavior or applied ethology. In this paper a few areas are offerred as particularly worthy of attention. These areas are worthwhile either because they have received little or no study and are of basic interest or because they have application to current problems of livestock production. The study of cat behavior falls in the former category. Neither the food and water sources, the reproductive success rate nor even the social interactions of cats in the large populations found in both rural and urban environments are known. Pigs as a species have already been the subjects of many behavior studies; nevertheless, there are still gaps in our knowledge of the underlying principles of swine behavior. The physiological basis of maternal behavior, for example, has not been studied in swine or in any domestic species. The sensory basis of udder location by the neonatal piglet deserves study also. Some aspects of olfactory and vocal communication of pigs have been studied, but only one of what may be a large number of pheromones of pigs has been chemically identified. The message conveyed by the vocal interactions between adult swine of the same sex is unknown, as is the role of facial and postural expressions in porcine communication. The two major problems of pig behavior under conditions of intensive livestock management are tail biting and reproductive failure. The application of behavioral techniques to these problems might help to attenuate those problems as well as broaden our understanding of normal pig behavior. Horse behavior has also been a relatively neglected field of study. Of particular interest is the significance of the flehmen gesture used by both mares and stallions in a variety of situations. Flehmen may be related to the function of the vomeronasal organ, but both observational and physiological studies should be performed to verify the hypothesis.
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Houpt, K. A., Law, K., & Martinisi, V. (1978). Dominance hierarchies in domestic horses. Appl. Animal. Ethol., 4(3), 273–283.
Abstract: Dominance hierarchies were studied in 11 herds of domestic horses and ponies (Equus caballus). A paired feeding test was utilized to establish the dominance--subordination relationship between each pair of animals in a herd. Aggressive actions, threats, bites, kicks and chases were also recorded. In small herds linear hierarchies were formed, but in large herds triangular relationships were observed. Aggression was correlated with dominance rank. Body weight, but not age, appear to affect rank in the equine hierarchy. Juvenile horses were more likely to share feed with each other than were adult horses and were usually subordinate to adult horses. The daughters of a dominant mare were dominant within their own herds.
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Ellis, L. (1995). Dominance and reproductive success among nonhuman animals: A cross-species comparison. Ethol. a. Sociob., 16(4), 257–333.
Abstract: This paper updates and extends Dewsbury's (1982) review of the literature on dominance and reproductive success (RS). The findings from approximately 700 studies are included, over two thirds of which were unavailable to Dewsbury. In order to give a highly condensed and yet meaningful overview, the main findings are represented in four tables, one for male nonprimates, one for female nonprimates, one for male primates, and one for female primates. In the tables for males, findings are analyzed in terms of six different indicators of RS, and in the tables for females, in terms of eight RS indicators. Outside the primate order, evidence largely supported the hypothesis that high-ranking males enjoy greater RS than do subordinate males. For females, studies are more evenly divided between those supporting the hypothesis that high rank and RS are positively correlated and those indicating no significant rank-RS relationship. This may reflect both the lower saliency of hierarchical relationships among females, as well as the lower variability in RS among females, relative to males. Among primates, a complex picture has emerged, especially in the case of males. Much of the complexity appears due to the importance of age and seniority in affecting dominance rank. Also, in some primate species, female preferences for sex partners seem to have little to do with the male's dominance rank, at least at the time mating takes place. Nevertheless, the majority of studies suggest that high- to middle-ranking males have at least a slight lifetime reproductive advantage over the lowest ranking males.
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Baer, K. L., Potter, G. D., Friend, T. H., & Beaver, B. V. (1983). Observation effects on learning in horses. Appl. Animal. Ethol., 11(2), 123–129.
Abstract: Sixteen horses, divided into 2 groups of 8, were used to study observational learning in horses. One group served as controls while the other group served as the treated group (observers). Observers were allowed to watch a correctly performed discrimination task for 5 days prior to testing their learning response using the same task. Discrimination testing was conducted on all horses daily for 14 days, with criterion set at 7 out of 8 responses correct with the last 5 consecutively correct. The maximum number of trials performed without reaching criterion was limited to 20 per day. Mean trials to criteria (MT) by group were: control, 11.25; observer, 10.70. Mean error (ME) scores were: control, 2.37; observer, 2.02. Average initial discrimination error scores were 11.13 for control and 10.38 for observers (P < 0.10). Asymptote was reached by Day 8 for both control and observer groups. Analysis of variance with repeated measures showed an extreme-day effect indicative of learning (P < 0.01), with non-significant differences in learning rate between experimental groups. Whether the initial ability of the horses to perform a discrimination learning task was enhanced by observation of other horses' performance of that task was not obvious from these data.
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Shi, J., & Dunbar, R. (2006). Feeding competition within a feral goat population on the Isle of Rum, NW Scotland. J. Ethol., 24(2), 117–124.
Abstract: This study investigated feeding competition within and between different age-sex classes of feral goats (Capra hircus) on the Isle of Rum (northwest Scotland) from August to November 2000 (inclusive). Although contests in a feeding context were common, most were relatively passive: little overt agonistic behaviour was observed between opponents and the distance between feeding animals involved did not change significantly after an interaction. Month (but not sex or habitat type) had a significant effect on feeding interaction rates, and the proportion of interactions involving more intense forms of conflict was highest in November when forage availability was beginning to decline. The results show that the initiator won most feeding encounters, with adult males being dominant over females. The ability to win conflicts increased with age for both males and females. However, it decreased sharply for adult males older than 5 years, which may, in part, explain the reduced overwinter survival of these individuals.
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Carson, K., & Wood-Gush, D. G. M. (1983). Equine behaviour: II. A review of the literature on feeding, eliminative and resting behaviour. Appl. Animal. Ethol., 10(3), 179–190.
Abstract: The literature on the feeding, eliminative and resting behaviour of horses has been reviewed to collate the information available on these subjects. The grazing and eliminative behaviour patterns of domestic horses are unlike those of free-ranging Equidae. The reasons for this are not known, but it can cause wasted grazing of up to 90% of a field. Certain conditions, such as provision of supplementary hay and lack of available herbage, can cause these behaviour patterns to change, although it is not known how to manipulate the grazing behaviour of horses to prevent deterioration of the pasture. Grazing behaviour is influenced by many variables and is more complex than the feeding behaviour of a stabled horse. Horses sleep for approximately 12% of the day and show 4 different sleep/wakefulness states -- alert wakefulness, drowsiness, slow-wave sleep and paradoxical sleep. Horses are able to maintain slow-wave sleep while standing, but they need to lie down for paradoxical sleep to occur, rarely spending more than 30 consecutive minutes in lateral recumbency.
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Sato, S. (1982). Leadership during actual grazing in a small herd of cattle. Appl. Animal. Ethol., 8(1-2), 53–65.
Abstract: An understanding of patterns of leadership during grazing movements is important where the management of grazing cattle is concerned. This paper describes the leadership displayed by grazing cattle by recording the spatial relationship (grazing style) among herd members as the group progressed slowly through a field. Grazing style was divided into “A”, “B” and “C”, meaning following, independence and leading, respectively. The results revealed the following features: (1) the frequency distributions of grazing style and grazing formation used by the herd varied with the seasons; (2) the individual animal variation in grazing style did not fundamentally change with the seasons; (3) there was negative linear correlation between Styles A and C and between Styles A and B. The more any cow followed the grazing movement, the less likely it was to lead the grazing movement or to be independent. Styles C and B tended to be positively related; (4) high, medium and low ranking animals in social dominance showed tendencies to behave in Styles C, A and B, respectively; (5) grazing style and weight gain were not clearly related; (6) the cows that tended to lead, be independent or follow less, tended to get out of their paddocks. The observations suggested (1) that the leader-follower-independent relationship, although modified in each season, did not vary fundamentally, (2) that the active movement of high ranking animals and the independent movement of low ranking animals governed the voluntary formation in grazing, and (3) that as grazing cattle that behaved in a single group and did not escape from their paddock were much easier to manage, the grazing style that expressed these characteristics was one of the significant indices for management of grazing cattle.
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Partridge, B. L. (1981). Internal dynamics and the interrelations of fish in schools. J Comp Physiol Sensory Neural Behav Physiol, 144(3), 313–325.
Abstract: The three-dimensional structure of schools of saithe (Pollachius virens) and the interactions between individuals over time were analyzed in 12,240 frames of videotape sampled at 2.7 Hz. Time series analyses of the interactions between identified individuals allowed testing of assumptions of anonymity vs. leadership in schools and investigation of the transfer of information between individuals by which collective decisions are made. Results include the following:1.Saithe match changes in both swimming direction and speed of their neighbors but correlations are greater for swimming speed. Average speed of the school does not greatly affect correlations between neighboring fish although the reaction latencies may be somewhat increased. As shown previously (Partridge et al. 1980) nearest neighbor distance (NND) decreases with increasing school velocity.2.Saithe simultaneously match the headings and swimming speeds of at least their first two nearest neighbors within the school (NN1 and NN2). Partialling out the correlation between a fish's neighbors demonstrates that a fish's correlation to his second nearest neighbor (NN2) is not simply a transitive function of mutual correlation between the NN1 and NN2.3.Several sources of individual variation in schooling performance were examined. In all respects except one, that of preferred positions within the school, saithe showed no individual differences, i.e., some were not “better schoolers” than others. Although fish in the school differed in length by up to a factor of 2.5, no size related effects in NND or nearest neighbor positioning were found.4.Single Linkage Cluster Analysis (SLCA) of the cross-correlations of fishs' swimming speeds and directions demonstrated quantitatively the existence of subgroups within schools if they contain more than 10-11 members. Subgroups acting more-or-less independently in terms of short term variations in speed and direction nonetheless remained within the school as a whole and were not often apparent to observers since members of one group interdigitated with those of another. How individuals know to which subgroup they belong remains unanswered.
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