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	Author	Zharkikh, T.L.; Andersen, L.
	Title	Behaviour of Bachelor Males of the Przewalski Horse (Equus ferus przewalskii) at the Reserve Askania Nova			Type	Journal Article
	Year	2009	Publication	Der Zoologische Garten	Abbreviated Journal	Zoologische Garten
	Volume	78	Issue	5-6	Pages	282-299
	Keywords	Equid; Przewalski horse (Equus ferus przewalskii); Bachelor group; Social behaviour
	Abstract	The aim of this study was to investigate social relationships between Przewalski horses at a high density in a bachelor group housed in a 3.5-ha enclosure. The group consisted of 16 males aged 5 to 16. Behavioural data were collected during 18 days, total 216 h. Fifteen minute focal animal sampling was used; each horse was observed three times a day for a total of 45 min. The occurrence of 25 behaviours was recorded, and group spacing behaviour was studied using nearest neighbour recordings. The group divided into four subgroups; this supports earlier findings of bachelor groups (n>=10) dividing into two or more subgroups if they included several males aged >5 years. The total frequency of social interactions was 14.6±1.1 h-1. Although the density of the group in this study was higher than in other zoos, the males interacted agonistically only 3.6 h-1. The most frequently observed social behaviour categories were friendly interactions. This study shows possibilities to use some investigative behaviours (marking, flehmen, olfactory investigation, etc.) as indicators of social status of animals in a group.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0044-5169	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5098
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	Author	Ryder, O.A.
	Title	Przewalski's horse – putting the wild horse back in the wild			Type	Journal Article
	Year	1988	Publication	Oryx	Abbreviated Journal	Oryx
	Volume	22	Issue		Pages	154-157
	Keywords	Equidae genetics.
	Abstract	The Asian wild horse, or Przewalski's horse (Equus przewalskii), is believed to currently survive only in captivity. There are more than 660 individuals in over 70 zoological collections, and animals are available for reintroduction. The Przewalski's horse had been bred in captivity for 12 generations, and inbreeding has occurred. Genetic variability has been lost, and released animals will require acclimatization on or near release sites. In China, a program is under way for acclimatization, breeding, and release of wild horses, and plans are being made for a similar program in Mongolia. (LCA)
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	Publisher		Place of Publication		Editor
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	Series Editor		Series Title		Abbreviated Series Title
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	Area		Expedition		Conference
	Notes	from Professor Hans Klingels Equine Reference List			Approved	yes
	Call Number				Serial	1537
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	Author	Osthaus, B.; Proops, L.; Hocking, I.; Burden, F.
	Title	Spatial cognition and perseveration by horses, donkeys and mules in a simple A-not-B detour task			Type	Journal Article
	Year	2013	Publication		Abbreviated Journal	Animal Cognition
	Volume	16	Issue	2	Pages	301-305
	Keywords	Equids; Donkeys; Horses; Mules; Dogs; Detour; Spatial reasoning; A-not-B; Perseveration
	Abstract	We investigated perseveration and detour behaviour in 36 equids (Equus caballus, E. asinus, E. caballus × E. asinus) and compared these data to those of a previous study on domestic dogs (Canis familiaris). The animals were required to make a detour through a gap at one end of a straight barrier in order to reach a visible target. After one, two, three or four repeats (A trials), the gap was moved to the opposite end of the barrier (B trials). We recorded initial deviations from the correct solution path and the latency to crossing the barrier. In the A trials, mules crossed the barrier significantly faster than their parental species, the horses and donkeys. In the B trials, following the change of gap location, all species showed a reduction in performance. Both dogs and horses exhibited significant spatial perseveration, going initially to the previous gap location. Donkeys and mules, however, performed at chance level. Our results suggest that hybrid vigour in mules extends to spatial abilities.
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	Corporate Author				Thesis
	Publisher	Springer-Verlag	Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5694
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	Author	Lamoot, I.; Callebaut, J.; Degezelle, T.; Demeulenaere, E.; Laquiere, J.; Vandenberghe, C.; Hoffmann, M.
	Title	Eliminative behaviour of free-ranging horses: do they show latrine behaviour or do they defecate where they graze?			Type	Journal Article
	Year	2004	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	86	Issue	1-2	Pages	105-121
	Keywords	Equids; Faeces avoidance; Grazing behaviour; Spatial differentiation; Urine; Horse marking
	Abstract	In contrast to horses in pastures, it is thought that free-ranging horses do not perform latrine behaviour, i.e. a behavioural pattern whereby the animals graze and defecate in separate areas. However, few studies deal with this particular subject, reporting contrasting conclusions. We hypothesize that horses free-ranging in large heterogeneous areas do not perform latrine behaviour. Thus, we believe that grazing and elimination behaviour are spatially related: where horses graze, they will also defecate. Behavioural data were collected from Konik horses, Haflinger horses, Shetland ponies and donkeys, grazing in different nature reserves (54-80 ha). Data for the different equids were analyzed separately, as well as data for mares and stallions (Konik and donkey stallions only). We investigated the proportion of the number of defecations/urinations while grazing on the total number of defecations/urinations; furthermore, we searched for the sequence of behaviours representing latrine behaviour in the strict sense. Additionally, we analyzed the correlation between grazing behaviour and eliminative behaviour on both vegetation type level and patch level. All the female equids often continued grazing while defecating. During urination, grazing ceases in the majority of instances. Cases where a mare terminated grazing in a certain vegetation type and sward height to eliminate in another vegetation type or in another sward height within the same vegetation type were rarely observed. On the vegetation type level as well as on the patch level, there was a highly significant (P<0.001) positive correlation between grazing time and number of eliminations (or eliminating time). The high values of the correlation coefficients (in case of the defecation variables r ranges between 0.553 and 0.955; in case of the urination variables r ranges between 0.370 and 0.839) illustrate that the spatial distribution of the eliminative behaviour can be explained to a high degree by the spatial distribution of the grazing behaviour. Results in the case of the stallions are preliminary, but indicate the same pattern. Horses, free-ranging in large heterogeneous areas, do not perform latrine behaviour, but defecate where they graze. Possibly, animal density is of major importance to explain this behavioural difference with horses in pastures. We suggest that also spatial vegetation heterogeneity and plant productivity of the grazed area, as well as parasite status of the grazing animals could play a role.
	Address
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	Publisher		Place of Publication		Editor
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2339
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	Author	Moehlman, P.D.
	Title	*Behavioral patterns and communication in feral asses (Equus africanus)*			Type	Journal Article
	Year	1998	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	60	Issue	2-3	Pages	125-169
	Keywords	Equids; Feral asses; Behavior patterns; Facial expressions; Postures; Locomotion
	Abstract	The behavior of feral populations of the African wild ass (Equus africanus) were studied in the Northern Panamint Range of Death Valley National Monument for 20 months from 1970 to 1973 [Moehlman, P.D., 1974. Behavior and ecology of feral asses (Equus asinus). PhD dissertation, University of Wisconsin, Madison, 251 pp.; Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Natl. Geogr. Soc. Res. Reports, 1970: 405-411]. Maintenance behavior is described and behavior sequences that were used in social interactions are quantified by sex and age class. Agonistic, sexual, and greeting behavior patterns are described and analyzed in conjunction with the responses they elicited. Mutual grooming mainly occurred between adult males, and between females and their offspring. Five types of vocalizations were distinguished: brays, grunts, growls, snorts, and whuffles. A second population was studied for 1 month on Ossabaw Island, GA (Moehlman, 1979). This population had more permanent social groups and had a higher rate of mutual grooming and foal social play.
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2381
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	Author	Moehlman, P.D.
	Title	*Feral asses (Equus africanus): intraspecific variation in social organization in arid and mesic habitats*			Type	Journal Article
	Year	1998	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	60	Issue	2-3	Pages	171-195
	Keywords	Equids; Feral asses; Social organization; Mating systems; Intraspecific variation
	Abstract	Feral asses have been studied in the arid habitats of the southwestern United States [Moehlman, P.D., 1974. Behavior and ecology of feral asses (Equus asinus). PhD dissertation, University of Wisconsin, Madison, 251 pp.; Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Nat. Geogr. Soc. Res. Reports 1970, 405-411.; Woodward, S.L., 1979. The social system of feral asses (Equus asinus). Z. Tierpsychol. 49, 304-316] and in the mesic habitat of Ossabaw Island, Georgia [Moehlman, P.D., 1979, ibid; McCort, W.D., 1980. The feral asses (Equus asinus) of Ossabaw Island, Georgia. PhD dissertation, Pennsylvania State University, University Park, 219 pp]. The feral ass populations in these two locales exhibited intraspecific variation in polygynous mating systems and social organization which were consistent with the ecological classification of mating systems of Emlen and Oring (1977) [Emlen, S.T., Oring, S.W., 1977. Ecology, sexual selection, and the evolution of mating systems. Science 197 (4300), 215-223]. Feral asses in the arid environment have a `resource defense' polygynous mating system, and those in the mesic habitat exhibit `female (harem) defense' polygyny. The intraspecific variation observed in feral asses encompasses the interspecific variation observed in the family Equidae.
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	Language		Summary Language		Original Title
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	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2382
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	Author	Moehlman, P.D.; Fowler, L.E.; Roe, J.H.
	Title	*Feral asses (Equus africanus) of Volcano Alcedo, Galapagos: behavioral ecology, spatial distribution, and social organization*			Type	Journal Article
	Year	1998	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	60	Issue	2-3	Pages	197-210
	Keywords	Equids; Feral asses; Social organization; Mating systems; Intraspecific variation; Galapagos
	Abstract	Feral asses were studied on Volcano Alcedo, Galapagos Islands, Ecuador, during the wet season of 1980. On the volcano rim during March/April, two stable groups were observed to have a `female (harem) defense' polygynous mating system [Emlen, S.T., Oring, S.W., 1977. Ecology, sexual selection, and the evolution of mating systems. Science 197 (4300), pp. 215-223] and social behavior patterns and feeding ecology similar to feral asses living in a habitat where forage and climate are similar, e.g., Ossabaw Island, Georgia [Moehlman, P.D., 1979. Behavior and ecology of feral asses (Equus asinus). Nat. Geogr. Soc. Res. Rep., 1970, pp. 405-411; Moehlman, P.D., 1997. Feral asses (Equus africanus): intraspecific variation in social organization in arid and mesic habitats. J. Appl. Anim. Behav. Sci., this issue; McCort, W.D., 1980. The feral asses (Equus asinus) of Ossabaw Island, Georgia., PhD Dissertation, Pennsylvania State University, University Park, 219 pp.].
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2383
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	Author	Boyd, L.; Bandi, N.
	Title	Reintroduction of takhi, Equus ferus przewalskii, to Hustai National Park, Mongolia: time budget and synchrony of activity pre- and post-release			Type	Journal Article
	Year	2002	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	78	Issue	2-4	Pages	87-102
	Keywords	Equids; Przewalski's horses; Takhi; Behaviour patterns; Time budgets; Behavioural synchrony; Reintroduction
	Abstract	A harem of takhi (Equus ferus przewalskii) was observed during introduction to the Hustain Nuruu Steppe Reserve of Mongolia. The goals were to examine whether the harem exhibited significant behavioural synchrony, whether release had an effect on time budget, and what the implications might be regarding acclimatisation to the wild. Behaviours were scan sampled every 10 min between the hours of 06:00 and 22:00, twice before release, twice immediately after release, and twice 2 years after reintroduction. Time budgets were constructed from these data. Considerable behavioural synchrony was evidenced both before and after release. Crepuscular grazing and midday resting were typical, regardless of the date relative to release. Upon release, the amount of time spent moving doubled for all age classes. It is suggested that this increase resulted from exploration. The amount of time spent grazing and standing remained unchanged; the increased amount of time spent moving came at the expense of resting. Two years later, the horses still spent more time moving than when captive. Somewhat less time was spent grazing, although the difference was not significant. More time was spent resting in 1996 than immediately after release. These time budgets provide evidence of successful acclimatisation to the wild. Trekking between favoured sites could account for the persistent increase in time spent moving, with concomitantly less time needed to meet nutritional needs by grazing and more time available for resting. Housing captive takhi in large enclosures is evidently insufficient to permit the amount of movement exhibited by this wild harem. The time budget of the 1- and 2-year olds was more similar to that of adults than foals, indicating approaching adulthood. That 1- and 2-year olds were nursed, without loss of body condition by the dam, provided additional evidence that the takhi achieved excellent nutritional status in the wild.
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	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
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	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3690
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	Author	Pluhácek, J.; Bartos, L.; Bartosová, J.; Kotrba, R.
	Title	Feeding behaviour affects nursing behaviour in captive plains zebra (Equus burchellii)			Type	Journal Article
	Year	2010	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	128	Issue	1-4	Pages	97-102
	Keywords	Equus burchellii; Suckling; Nursing; Equid; Feeding; Zoo
	Abstract	Equids spend more than half of the day feeding. Lactation is a very demanding form of maternal investment. In an environment such as a zoo, where no grass but several feeding sites are present, conflict between suckling behaviour of the foal and feeding behaviour of the lactating mother should occur. We observed 20 foals of captive plains zebra, Equus burchellii, at the Dvur Králové Zoo, Czech Republic, and collected data concerning suckling events during 17 months of observation. First, we examined whether feeding by the mother while nursing affected suckling behaviour. We found that when the mother was feeding, the proportion of suckling bouts she terminated decreased with increasing age of the foal, whereas it did not change when she was not feeding. This result supported the trade-off between suckling and feeding behaviour which has been reported in other ungulates. Second, we examined what affected interruptions of feeding behaviour of the mother during the suckling bout. The proportion of interruptions of feeding by the mother during nursing increased with increasing age of her foal. This coincides with declining time spent nursing. In addition, younger mothers interrupted their feeding behaviour during suckling bouts more often than older ones. Mothers interrupted feeding during the suckling bout more often when they nursed a daughter than when they nursed a son. The results of our study show that feeding while suckling could reduce parent-offspring conflict and improve welfare of captive foals and mares.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5280
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	Author	Lamoot, I.; Callebaut, J.; Demeulenaere, E.; Vandenberghe, C.; Hoffmann, M.
	Title	Foraging behaviour of donkeys grazing in a coastal dune area in temperate climate conditions			Type	Journal Article
	Year	2005	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	92	Issue	1-2	Pages	93-112
	Keywords	Grazing behaviour; Habitat use; Donkey; Equid; Diet composition
	Abstract	A small herd of donkeys was introduced in a coastal dune reserve `Houtsaegerduinen' (ca. 80 ha) in Belgium, in order to slow down expansion of dominant grass and shrub species. The Houtsaegerduinen is a nutrient poor scrub-dominated dune system with a spatially heterogeneous vegetation pattern. Different aspects of the grazing behaviour (grazing time, bite rate, habitat use, diet composition) of the free-ranging donkeys are described and analysed. Behavioural data (of maximum six adult mares) were collected through continuous focal animal observation in three consecutive years (1998-2001). Temporal variation in grazing time, habitat use and diet composition was determined. During daylight, donkeys spent most of their time on grazing (56%). In all 3 years, grazing time was significantly shorter in summer (45% of their time), longest grazing times were achieved in spring (64%). In spring, the donkeys also achieved the highest bite rate (21.5 bites/min). The grassy habitat was preferred for foraging in all seasons, while the use of scrub and woodland was variable over time. Averaged over the four seasons, the general diet consisted for 80% of graminoids, 10% of forbs and 10% of woody plants. However, diet composition varied not only among seasons and years, but depended also on the foraged habitat type. We discuss the possible role of the donkeys in nature management.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2338
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