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Watts, D.J.; Strogatz, S.H. |
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Collective dynamics of /`small-world/' networks |
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1998 |
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Nature |
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393 |
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6684 |
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440-442 |
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Networks of coupled dynamical systems have been used to model biological oscillators Josephson junction arrays excitable media, neural networks spatial games11, genetic control networks12 and many other self-organizing systems. Ordinarily, the connection topology is assumed to be either completely regular or completely random. But many biological, technological and social networks lie somewhere between these two extremes. Here we explore simple models of networks that can be tuned through this middle ground: regular networks 'rewired' to introduce increasing amounts of disorder. We find that these systems can be highly clustered, like regular lattices, yet have small characteristic path lengths, like random graphs. We call them 'small-world' networks, by analogy with the small-world phenomenon (popularly known as six degrees of separation). The neural network of the worm Caenorhabditis elegans, the power grid of the western United States, and the collaboration graph of film actors are shown to be small-world networks. Models of dynamical systems with small-world coupling display enhanced signal-propagation speed, computational power, and synchronizability. In particular, infectious diseases spread more easily in small-world networks than in regular lattices. |
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0028-0836 |
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10.1038/30918 |
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Equine Behaviour @ team @ |
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4989 |
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Lau, A.N.; Peng, L.; Goto, H.; Chemnick, L.; Ryder, O.A.; Makova, K.D. |
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Horse Domestication and Conservation Genetics of Przewalski's Horse Inferred from Sex Chromosomal and Autosomal Sequences |
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2009 |
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Mol Biol Evol |
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Mol Biol Evol |
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26 |
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1 |
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199-208 |
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Despite their ability to interbreed and produce fertile offspring, there is continued disagreement about the genetic relationship of the domestic horse (Equus caballus) to its endangered wild relative, Przewalski's horse (Equus przewalskii). Analyses have differed as to whether or not Przewalski's horse is placed phylogenetically as a separate sister group to domestic horses. Because Przewalski's horse and domestic horse are so closely related, genetic data can also be used to infer domestication-specific differences between the two. To investigate the genetic relationship of Przewalski's horse to the domestic horse and to address whether evolution of the domestic horse is driven by males or females, five homologous introns (a total of [~]3 kb) were sequenced on the X and Y chromosomes in two Przewalski's horses and three breeds of domestic horses: Arabian horse, Mongolian domestic horse, and Dartmoor pony. Five autosomal introns (a total of [~]6 kb) were sequenced for these horses as well. The sequences of sex chromosomal and autosomal introns were used to determine nucleotide diversity and the forces driving evolution in these species. As a result, X chromosomal and autosomal data do not place Przewalski's horses in a separate clade within phylogenetic trees for horses, suggesting a close relationship between domestic and Przewalski's horses. It was also found that there was a lack of nucleotide diversity on the Y chromosome and higher nucleotide diversity than expected on the X chromosome in domestic horses as compared with the Y chromosome and autosomes. This supports the hypothesis that very few male horses along with numerous female horses founded the various domestic horse breeds. Patterns of nucleotide diversity among different types of chromosomes were distinct for Przewalski's in contrast to domestic horses, supporting unique evolutionary histories of the two species. |
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Equine Behaviour @ team @ |
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4997 |
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Author |
Davis, M. H. |
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Title |
Empathy: A Social Psychological Approach |
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1996 |
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272 |
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Product Description
Empathy has long been a topic of interest to psychologists, but it has been studied in a sometimes bewildering number of ways. In this volume, Mark Davis offers a thorough, evenhanded review of contemporary empathy research, especially work that has been carried out by social and personality psychologists.Davis’ approach is explicitly multidimensional. He draws careful distinctions between situational and dispositional “antecedents” of empathy, cognitive and noncognitive “internal processes,” affective and nonaffective “intrapersonal outcomes,” and the “interpersonal behaviora
l outcomes” that follow. Davis presents a novel organizational model to help classify and interpret previous findings. This book will be of value in advanced undergraduate and graduate courses on altruism, helping, nad moral development.
About the Author
Mark H. Davis is associate professor of psychology at Eckerd College in St. Petersburg, Florida. |
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Westview Press |
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Boulder, CO |
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978-0813330013 |
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Equine Behaviour @ team @ |
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5017 |
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Oakenfull, E.A.; Lim, H.; Ryder, O. |
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A survey of equid mitochondrial DNA: Implications for the evolution, genetic diversity and conservation of Equus |
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2000 |
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Conservation Genetics |
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Conservat Genet |
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1 |
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4 |
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341-355 |
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The evolution, taxonomy and conservation of the genus Equuswere investigated by examining the mitochondrial DNA sequences of thecontrol region and 12S rRNA gene. The phylogenetic analysis of thesesequences provides further evidence that the deepest node in thephylogeny of the extant species is a divergence between twolineages; one leading to the ancestor of modern horses (E.ferus, domestic and przewalskii) and the other to thezebra and ass ancestor, with the later speciation events of the zebrasand asses occurring either as one or more rapid radiations, or withextensive secondary contact after speciation. Examination of the geneticdiversity within species suggested that two of the E. hemionussubspecies (E. h. onager and E. h. kulan) onlyrecently diverged, and perhaps, are insufficiently different to beclassified as separate subspecies. The genetic divergence betweendomestic and wild forms of E. ferus (horse) and E.africanus (African ass) was no greater than expected within anequid species. In E. burchelli (plains zebra) there was anindication of mtDNA divergence between populations increasing withdistance. The implications of these results for equid conservation arediscussed and recommendations are made for conservation action. |
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Equine Behaviour @ team @ |
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5039 |
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Hama, H.; Yogo, M.; Matsuyama, Y. |
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Title |
Effects of stroking horses on both humans' and horses' heart rate responses |
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1996 |
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Japanese Psychological Research |
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Jpn. Psychol. Res. |
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38 |
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2 |
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66-73 |
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The present study examined both human and horse heart rates (HRs) when humans stroked horses for 90 seconds; the subjective arousal levels of the humans were measured by the Tohoku Activation Deactivation Adjective Check List before and after stroking horses. Six male subjects with a positive attitude toward companion animals and 6 male subjects with a negative attitude were selected by their scores on the Pet Attitude Scale, and these two groups, together with a third group, of 6 subjects who were male members of the Doshisha University horse-riding club, participated in this experiment. The HRs of the human subjects during the first 10 seconds immediately after the stroking began were significantly higher than those obtained after that period, but these higher levels gradually returned to baseline levels. This tendency appears more clearly in the negative attitude group. The HRs of the horses increased during the first 20 seconds immediately after the human subjects of the NA group started stroking them, but gradually reduced as the stroking continued. The results of subjective arousal levels suggest a decrease in tension by stroking horses. These results suggest that a certain affectional interaction may exist between humans and companion animals. |
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Equine Behaviour @ team @ |
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5056 |
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Bates, L.A.; Lee, P.C.; Njiraini, N.; Poole, J.H.; Sayialel, K.; Sayialel, S.; Moss, C.J.; Byrne, R. |
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Title |
Do Elephants Show Empathy? |
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2008 |
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Journal of Consciousness Studies |
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J Conscious Stud |
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15 |
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10-11 |
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204-225 |
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Elephants show a rich social organization and display a number of unusual traits. In this paper, we analyse reports collected over a thirty-five year period, describing behaviour that has the potential to reveal signs of empathic understanding. These include coalition formation, the offering of protection and comfort to others, retrieving and 'babysitting' calves, aiding individuals that would otherwise have difficulty in moving, and removing foreign objects attached to others. These records demonstrate that an elephant is capable of diagnosing animacy and goal directedness, and is able to understand the physical competence, emotional state and intentions of others, when they differ from its own. We argue that an empathic understanding of others is the simplest explanation of these abilities, and discuss reasons why elephants appear to show empathy more than other non-primate species. |
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yes |
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Equine Behaviour @ team @ |
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5057 |
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KAUFMANN, J. H. |
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Title |
ON THE DEFINITIONS AND FUNCTIONS OF DOMINANCE AND TERRITORIALITY |
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1983 |
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Biological Reviews |
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Biol Rev |
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58 |
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1 |
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1-20 |
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1. Dominance/subordinance is a relationship between two individuals in which one defers to the other in contest situations. Each such relationship represents an adaptive compromise for each individual in which the benefits and costs of giving in or not giving in are compared. Familiar associates in groups or neighbours on nearby territories may develop relatively stable dominant-subordinate relationships based on individual recognition. Although the aggressive aspects of dominance are usually emphasized, the less conspicuous actions of the subordinate individual are actually more important in maintaining a stable relationship. 2. In evolutionary terms, dominance essentially equals priority of access to resources in short supply. Usually the subordinate, who would probably lose in combat anyway, is better off to bide its time until better able to compete at another time or another place. Both individuals save time, energy, and the risk of injury by recognizing and abiding by an established dominant-subordinate relationship. 3. Dominance can be either absolute or predictably reversible in different locations or at different times. Of the various forms of dominance behaviour, rank hierarchies and territoriality represent the two extremes of absolute and relative dominance, respectively. A dominance hierarchy is the sum total of the adaptive compromises made between individuals in an aggregation or organized group. Many animals seem to be capable of both absolute and relative dominance, and within species-specific limits the balance may shift toward one or the other. High density, or a decrease in available resources, favours a shift from relative to absolute dominance. Some species may exhibit both simultaneously. Social mammals may have intra-group hierarchies and reciprocal territoriality between groups, while the males of lek species may exhibit 'polarized territoriality' by defending small individual territories, with the most dominant males holding the central territories where most of the mating takes place. 4. Territoriality is a form of space-related dominance. Most biologists agree that its most important function is to provide the territory holder with an assured supply of critical resources. Territoriality is selected for only when the individual's genetic fitness is increased because its increased access to resources outweighs the time, energy, and injury costs of territorial behaviour. 5. Territoriality was first defined narrowly as an area from which conspecifics are excluded by overt defence or advertisement. The definition has been variously expanded to include all more or less exclusive areas without regard to possible defence, and finally to include all areas in which the owner is dominant. I define territory as a fixed portion of an individual's or group's range in which it has priority of access to one or more critical resources over others who have priority elsewhere or at another time. This priority of access must be achieved through social interaction. 6. My definition excludes dominance over individual space and moving resources, and includes areas of exclusive use maintained by mutual avoidance. It differs from most other definitions in its explicit recognition of time as a territorial parameter and its rejection of exclusivity and overt defence as necessary components of territorial behaviour. There is an indivisible continuum of degrees of trespass onto territories, and functionally it is priority of access to resources that is important rather than exclusive occupancy. 7. There is a similarly indivisible continuum in the intensity of behaviour needed to achieve priority of access to resources. Deciding whether or not an exclusive area is defended leads to the pointless exercise of trying to decide which cues indicating the owner's presence are conspicuous enough to merit being called defence. Concentrating on overt defence emphasizes the aggressive aspects of territorial behaviour rather than the equally or more important submissive aspects such as passive avoidance. |
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Department of Zoology, University of Florida, Gainesville, FL 32611, U.S.A. |
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Equine Behaviour @ team @ |
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5101 |
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Prins, H.H. |
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Ecology and Behaviour of the African Buffalo: Social Inequality and Decision Making |
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1995 |
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What are the benefits that animals gain from living in a social group? This question has been the primary focus of the author's ecological interest. After many years of original and innovative research on the African buffalo, particularly at Lake Manyara in northern Tanzania, Herbert Prins has now summarized the results of much of this widely-respected work in this fascinating book. While advantages in reduction of the risks of predation or in increased efficiency of foraging on certain types of resources are now widely recognized, until now there has been less attention paid to the idea of the animals themselves as `information centres' and the extent to which the individual may be able to make use of information gathered by conspecifics, adjusting its own behaviour in response. Such a case-study has wide implications for research on social structure and organization in other species, and these are explored within the book. However, it is not a book aimed simply at the academic researcher, zoologist and behavioural ecologist; since it is written in a readable and accessible style, the book will also be enjoyed by wildlife enthusiasts, interested naturalists, wildlife biologists and wildlife managers. |
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Springer Netherland |
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978-0412725203 |
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Equine Behaviour @ team @ |
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5142 |
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Cameron, E.Z.; Setsaas, T.H.; Linklater, W.L. |
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Social bonds between unrelated females increase reproductive success in feral horses |
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2009 |
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Proceedings of the National Academy of Sciences of the United States of America |
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Proc. Natl. Acad. Sci. U.S.A. |
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106 |
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33 |
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13850-13853 |
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In many mammals, females form close social bonds with members of their group, usually between kin. Studies of social bonds and their fitness benefits have not been investigated outside primates, and are confounded by the relatedness between individuals in primate groups. Bonds may arise from kin selection and inclusive fitness rather than through direct benefits of association. However, female equids live in long-term social groups with unrelated members. We present 4 years of behavioral data, which demonstrate that social integration between unrelated females increases both foal birth rates and survival, independent of maternal habitat quality, social group type, dominance status, and age. Also, we show that such social integration reduces harassment by males. Consequently, social integration has strong direct fitness consequences between nonrelatives, suggesting that social bonds can evolve based on these direct benefits alone. Our results support recent studies highlighting the importance of direct benefits in maintaining cooperative behavior, while controlling for the confounding influence of kinship. |
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10.1073/pnas.0900639106 |
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Equine Behaviour @ team @ |
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5152 |
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Hemelrijk, C.K.; Hildenbrandt, H. |
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Self-Organized Shape and Frontal Density of Fish Schools |
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2008 |
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Ethology |
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Ethology |
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114 |
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3 |
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245-254 |
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Abstract Models of swarming (based on avoidance, alignment and attraction) produce patterns of behaviour also seen in schools of fish. However, the significance of such similarities has been questioned, because some model assumptions are unrealistic [e.g. speed in most models is constant with random error, the perception is global and the size of the schools that have been studied is small (up to 128 individuals)]. This criticism also applies to our former model, in which we demonstrated the emergence of two patterns of spatial organization, i.e. oblong school form and high frontal density, which are supposed to function as protection against predators. In our new model we respond to this criticism by making the following improvements: individuals have a preferred ‘cruise speed’ from which they can deviate in order to avoid others or to catch up with them. Their range of perception is inversely related to density, with which we take into account that high density limits the perception of others that are further away. Swarm sizes range from 10 to 2000 individuals. The model is three-dimensional. Further, we show that the two spatial patterns (oblong shape and high frontal density) emerge by self-organization as a side-effect of coordination at two speeds (of two or four body lengths per second) for schools of sizes above 20. Our analysis of the model leads to the development of a new set of hypotheses. If empirical data confirm these hypotheses, then in a school of real fish these patterns may arise as a side-effect of their coordination in the same way as in the model. |
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Blackwell Publishing Ltd |
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1439-0310 |
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Equine Behaviour @ team @ |
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5202 |
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