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	Author	Pettifor, R.A.
	Title	The effects of avian mobbing on a potential predator, the European kestrel, Falco tinnunculus			Type	Journal Article
	Year	1990	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	39	Issue	5	Pages	821-827
	Keywords
	Abstract	European kestrels were observed being mobbed by other birds on 63 occasions. Eleven species were involved, and in two instances mobs were composed of more than one species. Both flight-hunting and perch-hunting kestrels flew significantly further between their foraging positions when they were mobbed than when they were not mobbed; on average, mobbing resulted in flight-hunting kestrels moving 6[middle dot]8 times, and perch-hunting kestrels 2[middle dot]7 times, the mean distances moved by non-mobbed birds. The mean strike distance of perch-hunting kestrels attempting to capture birds was significantly less than the distance between perches flown by perch-hunting kestrels when mobbed. These data provide quantitative support for the assumption that mobbing causes a predator to vacate its immediate foraging area. The activity of the kestrels also influenced the frequency that they were mobbed, with kestrels that were flight-hunting being mobbed more than expected compared with ones that were perch-hunting. Kestrels were observed being mobbed throughout the year, and there was no discernible difference in their response to mobbing between seasons. These results are discussed in relation to current ideas on the functions of avian mobbing.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4091
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	Author	Lingle, S.; Rendall, D.; Pellis, S.M.
	Title	Altruism and recognition in the antipredator defence of deer: 1. Species and individual variation in fawn distress calls			Type	Journal Article
	Year	2007	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	73	Issue	5	Pages	897-905
	Keywords	acoustic signals; aggressive defence; altruism; cooperation; mule deer; Odocoileus hemionus; Odocoileus virginianus; olfactory cues; recognition error; white-tailed deer
	Abstract	Mule deer, Odocoileus hemionus, females actively defend fawns against predators, including nonoffspring conspecific fawns and heterospecific white-tailed deer, O. virginianus, fawns. We hypothesized that the defence of nonoffspring fawns was due to a recognition error. During a predator attack, females may have to decide whether to defend a fawn with imperfect information on its identity obtained from hearing only a few distress calls. We examined fawn distress calls to determine whether calls made by the two species and by different individuals within each species were acoustically distinctive. The mean and maximum fundamental frequencies of mule deer fawns were nearly double those of white-tailed deer fawns, with no overlap, enabling us to classify 100% of calls to the correct species using a single trait. A large proportion of calls was also assigned to the correct individual using a multivariate analysis (66% and 70% of mule deer and white-tailed deer fawns, respectively, chance = 6% and 10%); however, there was considerable statistical uncertainty in the probability of correct classification. We observed fawns approach conspecific females in an attempt to nurse; females probed most offspring fawns with their noses before accepting them, and always probed nonoffspring fawns before rejecting them, suggesting that close contact and olfactory information were needed to unequivocally distinguish nonoffspring from offspring fawns. Taken together, these results suggest that acoustic variation alone would probably be sufficient to permit rapid and reliable species discrimination, but it may not be sufficient for mothers to unequivocally distinguish their own fawn from conspecific fawns.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4210
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	Author	Lingle, S.; Rendall, D.; Wilson, W.F.; DeYoung, R.W.; Pellis, S.M.
	Title	Altruism and recognition in the antipredator defence of deer: 2. Why mule deer help nonoffspring fawns			Type	Journal Article
	Year	2007	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	73	Issue	5	Pages	907-916
	Keywords	aggressive defence; altruism; behavioural discrimination; cooperation; motivational constraint; mule deer; Odocoileus hemionus; Odocoileus virginianus; recognition error; white-tailed deer
	Abstract	Both white-tailed deer, Odocoileus virginianus, and mule deer, O. hemionus, females defend fawns against coyotes, Canis latrans, but only mule deer defend nonoffspring conspecific and heterospecific fawns. During a predator attack, females may have to decide whether to defend a fawn while having imperfect information on its identity obtained from hearing a few distress calls. Although imperfect recognition can influence altruistic behaviour, few empirical studies have considered this point when testing functional explanations for altruism. We designed a series of playback experiments with fawn distress calls to test alternative hypotheses (by-product of parental care, kin selection, reciprocal altruism) for the mule deer's defence of nonoffspring, specifically allowing for the possibility that females mistake these fawns for their own. White-tailed deer females approached the speaker only when distress calls of white-tailed deer fawns were played and when their own fawn was hidden, suggesting that fawn defence was strictly a matter of parental care in this species. In contrast, mule deer females responded similarly and strongly, regardless of the caller's identity, the female's reproductive state (mother or nonmother) or the presence of their own offspring. The failure of mule deer females to adjust their responses to these conditions suggests that they do not defend nonoffspring because they mistake them for their own fawns. The lack of behavioural discrimination also suggests that kin selection, reciprocal altruism and defence of the offspring's area are unlikely to explain the mule deer's defence of nonoffspring. We identify causal and functional questions that still need to be addressed to understand why mule deer defend fawns so indiscriminately.
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	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
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	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4211
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	Author	McElreath, R.; Strimling, P.
	Title	How noisy information and individual asymmetries can make `personality' an adaptation: a simple model			Type	Journal Article
	Year	2006	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	72	Issue	5	Pages	1135-1139
	Keywords
	Abstract	Recent attention has been drawn to the existence of individual differences in correlated behaviour across contexts, animal `personality' (Gosling 2001, Psychological Bulletin, 127, 45-86) and behavioural syndromes (Sih et al. 2004b, Quarterly Review of Biology, 79, 241-277). The causes of these patterns of behaviour are subjects of debate. Here, we present a very simple model of how adaptively managing noisy information, combined with differences in individual state, can lead to evolutionarily stable differences in how individuals respond to environmental cues. When information is very noisy, behavioural syndromes are most likely, but as long as there is some error, some types of individuals display the same behaviour in all contexts. In extreme cases, very few individuals display flexible behaviour, and different stable behavioural types dominate the population.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4280
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	Author	de Vries, H.
	Title	An improved test of linearity in dominance hierarchies containing unknown or tied relationships			Type	Journal Article
	Year	1995	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	50	Issue	5	Pages	1375-1389
	Keywords
	Abstract	Appleby (1983, Anim. Behav., 31, 600-608) described a statistical test, based on the work of Kendall (1962, Rank Correlation Methods), for the significance of linearity in dominance hierarchies. He suggested that unknown relationships should be assigned the value 1/2 and that subsequently the same test procedure can be used. In this paper it is shown that incorrect results are obtained by this method whenever there are unknown relationships. Values of the linearity index are systematically too low. P-values can be too high (underestimating the significance) or too low (overestimating), and seem to differ by not much more than a factor two (respectively a half) from the correct P-value. An improved method is developed for testing linearity in a set of dominance relationships containing unknown relationships. Furthermore, it is argued that, if one admits the possibility of tied dominance relationships, which should indeed be assigned the value 1/2, Landau's linearity index is to be preferred to Kendall's index. A randomization test is developed for assessing the significance of linearity or non-linearity in a set of dominance relationships containing unknown or tied relationships. The test statistic employed in this testing procedure is based on Landau's linearity index, but takes the unknown and tied relationships into account.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
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	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4284
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	Author	Cameron, E.Z.; Linklater, W.L.; Stafford, K.J.; Minot, E.O.
	Title	Maternal investment results in better foal condition through increased play behaviour in horses			Type	Journal Article
	Year	2008	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	76	Issue	5	Pages	1511-1518
	Keywords	Equus caballus; feral horse; maternal investment; play
	Abstract	Play behaviour is widespread in mammals, but benefits to play have been difficult to demonstrate. Physical training is one of the many proposed hypotheses, suggesting that males and females should play differently, that increased maternal investment should lead to increases in play, and that increases in play should result in physical advantages. In a population of feral horses, Equus caballus, males and females did not differ in their play behaviour except that males initiated more of their play bouts than females. Maternal condition influenced play behaviour only in males, with sons of mothers in good condition playing more. However, when we controlled for maternal effects by comparing a son and a daughter of the same mother, daughters played more when their mother was in poor condition and sons played more when their mother was in good condition. Mothers of foals that played more lost more condition. Therefore, the difference in play behaviour could not be explained by offspring sex or maternal condition alone, but play behaviour mirrored variation in maternal investment. In addition, those individuals that played more survived better and had better body condition as yearlings despite weaning earlier. Since increased activity has been linked to enhanced musculoskeletal development in domestic horses, we suggest that play provides a link between increased maternal investment, increased body condition and future reproductive success in feral horses, and probably in other species.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0003-3472	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	4709
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	Author	Broom, D.M.; Sena, H.; Moynihan, K.L.
	Title	Pigs learn what a mirror image represents and use it to obtain information			Type	Journal Article
	Year	2009	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	78	Issue	5	Pages	1037-1041
	Keywords	awareness; cognition; learning; mirror; pig; Sus scrofa
	Abstract	Mirror usage has been taken to indicate some degree of awareness in animals. Can pigs, Sus scrofa, obtain information from a mirror? When put in a pen with a mirror in it, young pigs made movements while apparently looking at their image. After 5 h spent with a mirror, the pigs were shown a familiar food bowl, visible in the mirror but hidden behind a solid barrier. Seven out of eight pigs found the food bowl in a mean of 23 s by going away from the mirror and around the barrier. Naïve pigs shown the same looked behind the mirror. The pigs were not locating the food bowl by odour, did not have a preference for the area where the food bowl was and did not go to that area when the food bowl was visible elsewhere. To use information from a mirror and find a food bowl, each pig must have observed features of its surroundings, remembered these and its own actions, deduced relationships among observed and remembered features and acted accordingly. This ability indicates assessment awareness in pigs. The results may have some effects on the design of housing conditions for pigs and may lead to better pig welfare.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0003-3472	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5053
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	Author	Ward, C.; Trisko, R.; Smuts, B.B.
	Title	Third-party interventions in dyadic play between littermates of domestic dogs, Canis lupus familiaris			Type	Journal Article
	Year	2009	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	78	Issue	5	Pages	1153-1160
	Keywords	Canis lupus familiaris; domestic dog; littermates; play; social development; third-party interventions
	Abstract	Interventions occur when animals interfere in competitive interactions between two or more individuals. Interveners can alter the nature of the ongoing interaction by targeting one party (attacking, biting) and supporting the other. Three theories have been proposed to account for intervention behaviour: kin selection, reciprocity and direct benefits. The kin selection hypothesis predicts that interveners will selectively support relatives over nonrelatives; the reciprocity hypothesis predicts that when intervener [`]A' supports individual [`]B', later [`]B' will intervene and support [`]A'; and the direct benefits hypothesis predicts that target/support patterns should serve the immediate interests of the intervener. We tested the reciprocity and direct benefits hypotheses by exploring third-party interventions in play fighting among littermates of domestic dogs. Interveners in dyadic play did not preferentially target or support preferred playmates of the intervener. Interveners targeted the dog in the losing role at the time of the intervention, and they did not show reciprocity in support. Taken together, these last two findings suggest that littermates benefit directly and use interventions opportunistically to practise offence behaviours directed at littermates already behaving subordinately. Opportunities to practise targeting in a playful setting may help structure dominance relationships among littermates. Additionally, the tendency for puppies to do what the other is doing (target the dog in the losing role) may pave the way for synchronizing cooperative behaviours during group hunting and territorial defence. The types of behaviours used to intervene changed over development, but the outcome following an intervention remained stable.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0003-3472	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5090
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	Author	Lusseau, D.; Whitehead, H.; Gero, S.
	Title	Incorporating uncertainty into the study of animal social networks			Type	Journal Article
	Year	2008	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	75	Issue	5	Pages	1809-1815
	Keywords	bootstrap; social behaviour; social network; social structure
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0003-3472	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5173
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	Author	Bonanni, R.; Cafazzo, S.; Valsecchi, P.; Natoli, E.
	Title	Effect of affiliative and agonistic relationships on leadership behaviour in free-ranging dogs			Type	Journal Article
	Year	2010	Publication	Animal Behaviour.	Abbreviated Journal	Anim. Behav.
	Volume	79	Issue	5	Pages	981-991
	Keywords	affiliative relationship; agonistic dominance; Canis lupus familiaris; consensus costs; consensus decisions; domestic dog; formal dominance; individual variation in leadership
	Abstract	Consensus decisions about the nature and timing of group activities allow animals to maintain group cohesiveness, but also entail costs because individuals often differ with respect to their optimal activity budgets. Two mechanisms whereby animals reach a consensus include ‘consistent leadership’, in which a single dominant individual makes the decision, and ‘variable leadership’ in which several group members contribute to the decision outcome. Sharing of consensus decisions is expected to reduce consensus costs to most group members. Both patterns are thought to emerge from the complexity of social relationships of group members. We investigated the distribution of leadership during group departures in two packs of free-ranging dogs, Canis lupus familiaris, and tested how its distribution between individuals was affected by dominance rank-related affiliative and agonistic relationships. Although leadership was not entirely concentrated on a single group member, both packs had a limited number of habitual leaders. In the largest pack, the pattern of leadership changed from ‘variable’ to nearly ‘consistent’ after its size had shrunk. Habitual leaders were usually old and high-ranking individuals. However, high-ranking dogs that received affiliative submissions in greeting ceremonies were more likely to lead than dominant dogs receiving submissions only in agonistic contexts. During resting times, habitual followers associated more closely with habitual leaders than with other followers. These results suggest that in social species collective movements may arise from the effort of subordinates to maintain close proximity with specific valuable social partners.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0003-3472	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5177
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