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	Author	Sigurjónsdóttir, H.; Haraldsson, H.
	Title	Significance of Group Composition for the Welfare of Pastured Horses			Type	Journal Article
	Year	2019	Publication	Animals	Abbreviated Journal	Animals
	Volume	9	Issue	14	Pages
	Keywords	horse welfare; aggression; allogrooming; pastured horses; Icelandic horse
	Abstract	We explore how herd composition and management factors correlate with frequencies of social interactions in horse groups. Since the welfare of horses correlates with low aggression levels and social contact opportunities, information of this kind is important. The data are a collection of records of social interactions of 426 Icelandic horses in 20 groups of at least eight horses. The complexities and limitations of the data prohibit useful statistical modelling so the results are presented descriptively. Interesting and informative patterns emerge which can be of use both in management and in future studies. Of special interest are the low levels of agonistic behaviours in breeding groups where one stallion was present. The horses were less agonistic when in groups with young foals and where group membership was stable. Unfamiliar yearlings in peer groups were especially aggressive. Allogrooming was most frequent in groups with relatively more young horses and in unstable and small groups. Interestingly, the horses allogroomed more if they had few preferred allogrooming partners. The findings show that composition (age/sex) and stability of groups are of great importance with respect to aggression levels and opportunities for establishing bonds.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title	Animals
	Series Volume	9	Series Issue	1	Edition
	ISSN	2076-2615	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6510
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	Author	Minero, M.; Canali, E.; Ferrante, V.; Verga, M.; Odberg, F.O.
	Title	Heart rate and behavioural responses of crib-biting horses to two acute stressors			Type	Journal Article
	Year	1999	Publication	The Veterinary Record	Abbreviated Journal	Vet. Rec.
	Volume	145	Issue	15	Pages	430-433
	Keywords	Acute Disease; Animals; Behavior, Animal/physiology; Female; Heart Rate/physiology; Horse Diseases/physiopathology; Horses/physiology/psychology; Male; Stereotyped Behavior/physiology; Stress/physiopathology/psychology/veterinary
	Abstract	The heart rate and behaviour of 14 adult saddle horses, eight crib-biters and six normal controls, were investigated. Initially, the relationship between crib-biting and heart rate was investigated while the horses were undisturbed. The horses were tested when restrained with a lip twitch, and assessed when they were exposed suddenly to the rapid inflation of a balloon. The heart rate of the crib-biters during crib-biting was lower than during other behaviours. The crib-biters had a higher overall mean heart rate (P<0.05) suggesting that they may have had a higher basal sympathetic activity. After the application of the twitch, all the horses had a transient increase in heart rate which returned to basal values more rapidly in the crib-biters. The crib-biters were less reactive to the lip twitch, five of the six investigated remaining calm, and after the release of the twitch, they spent more time nibbling (P<0.05) than the control horses. The crib-biters reacted more strongly to the inflation of the balloon (three of the six reacted), and after it had been inflated they spent more time walking in the box.
	Address	Instituto di Zootecnica Veterinaria, Universita degli Studi di Milano, Italy
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0042-4900	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:10755589			Approved	no
	Call Number	refbase @ user @			Serial	1921
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	Author	Jansen, T.; Forster, P.; Levine, M.A.; Oelke, H.; Hurles, M.; Renfrew, C.; Weber, J.; Olek, K.
	Title	Mitochondrial DNA and the origins of the domestic horse			Type	Journal Article
	Year	2002	Publication	Proceedings of the National Academy of Sciences of the United States of America	Abbreviated Journal	Proc. Natl. Acad. Sci. U.S.A.
	Volume	99	Issue	16	Pages	10905-10910
	Keywords	Animals; Animals, Domestic/classification/genetics; Base Sequence; DNA, Complementary; DNA, Mitochondrial; Evolution, Molecular; Horses/classification/genetics; Molecular Sequence Data; Phylogeny
	Abstract	The place and date of the domestication of the horse has long been a matter for debate among archaeologists. To determine whether horses were domesticated from one or several ancestral horse populations, we sequenced the mitochondrial D-loop for 318 horses from 25 oriental and European breeds, including American mustangs. Adding these sequences to previously published data, the total comes to 652, the largest currently available database. From these sequences, a phylogenetic network was constructed that showed that most of the 93 different mitochondrial (mt)DNA types grouped into 17 distinct phylogenetic clusters. Several of the clusters correspond to breeds and/or geographic areas, notably cluster A2, which is specific to Przewalski's horses, cluster C1, which is distinctive for northern European ponies, and cluster D1, which is well represented in Iberian and northwest African breeds. A consideration of the horse mtDNA mutation rate together with the archaeological timeframe for domestication requires at least 77 successfully breeding mares recruited from the wild. The extensive genetic diversity of these 77 ancestral mares leads us to conclude that several distinct horse populations were involved in the domestication of the horse.
	Address	Biopsytec Analytik GmbH, Marie-Curie-Strasse 1, 53359 Rheinbach, Germany. jansen@biopsytec.com
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0027-8424	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12130666			Approved	no
	Call Number	refbase @ user @			Serial	772
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	Author	Polverini, E.; Cugini, G.; Annoni, F.; Abbruzzetti, S.; Viappiani, C.; Gensch, T.
	Title	Molten globule formation in apomyoglobin monitored by the fluorescent probe Nile Red			Type	Journal Article
	Year	2006	Publication	Biochemistry	Abbreviated Journal	Biochemistry
	Volume	45	Issue	16	Pages	5111-5121
	Keywords	Animals; Apoproteins/chemistry/metabolism; Binding Sites; Computer Simulation; Fluorescent Dyes/analysis; Horses; Hydrogen-Ion Concentration; Models, Molecular; Myoglobin/chemistry/metabolism; Oxazines/*analysis/chemistry; Protein Binding; Protein Folding; Protein Structure, Tertiary
	Abstract	The interaction of nile red (NR) with apomyoglobin (ApoMb) in the native (pH 7) and molten globule (pH 4) states was investigated using experimental and computational methods. NR binds to hydrophobic locations in ApoMb with higher affinity (K(d) = 25 +/- 5 microM) in the native state than in the molten globule state (K(d) = 52 +/- 5 microM). In the molten globule state, NR is located in a more hydrophobic environment. The dye does not bind to the holoprotein, suggesting that the binding site is located at the heme pocket. In addition to monitoring steady-state properties, the fluorescence emission of NR is capable of tracking submillisecond, time-resolved structural rearrangements of the protein, induced by a nanosecond pH jump. Molecular dynamics simulations were run on ApoMb at neutral pH and at pH 4. The structure obtained for the molten globule state is consistent with the experimentally available structural data. The docking of NR with the crystal structure shows that the ligand binds into the binding pocket of the heme group, with an orientation bringing the planar ring system of NR to overlap with the position of two of the heme porphyrin rings in Mb. The docking of NR with the ApoMb structure at pH 4 shows that the dye binds to the heme pocket with a slightly less favorable binding energy, in keeping with the experimental K(d) value. Under these conditions, NR is positioned in a different orientation, reaching a more hydrophobic environment in agreement with the spectroscopic data.
	Address	Dipartimento di Fisica, Universita degli Studi di Parma, Viale G. P. Usberti 7/A, 43100 Parma, Italy
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0006-2960	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16618100			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3763
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	Author	Gulotta, M.; Gilmanshin, R.; Buscher, T.C.; Callender, R.H.; Dyer, R.B.
	Title	Core formation in apomyoglobin: probing the upper reaches of the folding energy landscape			Type	Journal Article
	Year	2001	Publication	Biochemistry	Abbreviated Journal	Biochemistry
	Volume	40	Issue	17	Pages	5137-5143
	Keywords	Animals; Apoproteins/chemistry; Computer Simulation; Horses; Hydrogen-Ion Concentration; Kinetics; Models, Molecular; Myoglobin/chemistry; *Protein Folding; Protein Structure, Secondary; Protein Structure, Tertiary; Spectrometry, Fluorescence/instrumentation/methods; Thermodynamics; Tryptophan/chemistry
	Abstract	An acid-destabilized form of apomyoglobin, the so-called E state, consists of a set of heterogeneous structures that are all characterized by a stable hydrophobic core composed of 30-40 residues at the intersection of the A, G, and H helices of the protein, with little other secondary structure and no other tertiary structure. Relaxation kinetics studies were carried out to characterize the dynamics of core melting and formation in this protein. The unfolding and/or refolding response is induced by a laser-induced temperature jump between the folded and unfolded forms of E, and structural changes are monitored using the infrared amide I' absorbance at 1648-1651 cm(-1) that reports on the formation of solvent-protected, native-like helix in the core and by fluorescence emission changes from apomyoglobin's Trp14, a measure of burial of the indole group of this residue. The fluorescence kinetics data are monoexponential with a relaxation time of 14 micros. However, infrared kinetics data are best fit to a biexponential function with relaxation times of 14 and 59 micros. These relaxation times are very fast, close to the limits placed on folding reactions by diffusion. The 14 micros relaxation time is weakly temperature dependent and thus represents a pathway that is energetically downhill. The appearance of this relaxation time in both the fluorescence and infrared measurements indicates that this folding event proceeds by a concomitant formation of compact secondary and tertiary structures. The 59 micros relaxation time is much more strongly temperature dependent and has no fluorescence counterpart, indicating an activated process with a large energy barrier wherein nonspecific hydrophobic interactions between helix A and the G and H helices cause some helix burial but Trp14 remains solvent exposed. These results are best fit by a multiple-pathway kinetic model when U collapses to form the various folded core structures of E. Thus, the results suggest very robust dynamics for core formation involving multiple folding pathways and provide significant insight into the primary processes of protein folding.
	Address	Department of Biochemistry, Albert Einstein College of Medicine, Bronx, New York 10461, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0006-2960	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11318635			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3789
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	Author	McGreevy, P.D.; Webster, A.J.; Nicol, C.J.
	Title	Study of the behaviour, digestive efficiency and gut transit times of crib-biting horses			Type	Journal Article
	Year	2001	Publication	The Veterinary record	Abbreviated Journal	Vet. Rec.
	Volume	148	Issue	19	Pages	592-596
	Keywords	Animals; Behavior, Animal/physiology; Case-Control Studies; Digestion; Gastrointestinal Motility/drug effects; Horse Diseases/physiopathology; Horses/physiology/psychology; Male; Stereotyped Behavior/physiology; Sulfapyridine/blood; Sulfasalazine/diagnostic use/pharmacology
	Abstract	The spontaneous behaviour and the apparent digestibility of dry matter and fibre and transit times of digesta were compared in four normal horses and four crib-biters. A technique was developed for measuring total gut transit times (TGTT) by using single-stool analysis of the passage of radio-opaque polyethylene markers. Longer TGTT were recorded in the crib-biters than in the normal horses but the orocaecal transit times did not differ. The crib-biters rested less than the normal horses.
	Address	Department of Clinical Veterinary Science, University of Bristol, Langford
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0042-4900	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11386445			Approved	no
	Call Number	refbase @ user @			Serial	86
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	Author	Pinchbeck, G.L.; Clegg, P.D.; Proudman, C.J.; Morgan, K.L.; French, N.P.
	Title	Case-control study to investigate risk factors for horse falls in hurdle racing in England and Wales			Type	Journal Article
	Year	2003	Publication	The Veterinary Record	Abbreviated Journal	Vet. Rec.
	Volume	152	Issue	19	Pages	583-587
	Keywords	Accidental Falls/statistics & numerical data; Animals; Athletic Injuries/epidemiology/etiology/veterinary; Case-Control Studies; England/epidemiology; Horses/*injuries; Risk Factors; Running/injuries; Wales/epidemiology
	Abstract	Between March 1, 2000 and August 31, 2001, a case-control study was conducted on 12 racecourses in England and Wales to identify and quantify the risk factors associated with horse falls in hurdle races. The cases and controls were defined so that variables relating to the horse, the jockey, the race and racecourse, and the jump could be considered. The cases were defined as a jumping effort at a hurdle flight that resulted in a fall, and the controls were defined as a successful jump over a hurdle at any of the 12 racecourses within 14 days before or after the case fall. Conditional logistic regression was used to examine the univariable and multivariable relationships between the predictor variables and the risk of falling. The risk of falling was significantly associated with the position of the jump in the race, and with the distance and speed of the race. A horse's previous racing experience and history were also significantly associated with the risk of falling and horses participating in their first hurdle race were at almost five times greater risk of falling than horses that had hurdled before.
	Address	Epidemiology Group, Department of Veterinary Clinical Science and Animal Husbandry, University of Liverpool, Leahurst, Neston CH64 7TE
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0042-4900	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12762486			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3782
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	Author	Granquist, S.M.; Thorhallsdottir, A.G.; Sigurjonsdottir, H.
	Title	The effect of stallions on social interactions in domestic and semi feral harems			Type	Journal Article
	Year	2012	Publication	Applied Animal Behaviour Science	Abbreviated Journal
	Volume	141	Issue	1–2	Pages	49-56
	Keywords	Horses; Hierarchy; Icelandic horse; Social behaviour; Social bonds
	Abstract	Earlier research indicates that stallions may supress interactions of their harem members, leading to less stable hierarchies and friendship bonds in harems compared to non-stallion groups. In this paper, the effect of the presense of a stallion on the social behaviour of mares was studied by comparing six harems containing stallions to four mixed sex groups not containing stallions. Both temporary and permanent harems were studied, giving the possibility to investigate the effect of group stability on social interactions. A significant linear hierarchy was found in all non-stallion groups that were used for comparison, while the hierarchies were only found to be linear in three of the six harems containing stallions (Landaus h', p < 0.05). Aggression rate was lower (t-test, p < 0.05) and fewer friendship bonds (G-test, p < 0.0001) were found within the harems, compared to the groups without stallions. Stallions seldom intervene directly in interactions between harem members. Thus, our results give support to the hypothesis that stallions may suppress interactions of harem members, but in a more indirect way than with direct interference. In addition, our results give support for earlier findings that aggression rate may be affected by group stability. We found a higher aggression rate in the temporary harems compared to the permanent harems (Kruskal–Wallis, p < 0.05) and in the temporary non-stallion group compared to the permanent non-stallion group. The results have significance for further research on social structure of mammals, and may be applied in management of domestic animals.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5619
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	Author	Virga, V.; Houpt, K.A.
	Title	Prevalence of placentophagia in horses			Type	Journal Article
	Year	2001	Publication	Equine veterinary journal	Abbreviated Journal	Equine Vet J
	Volume	33	Issue	2	Pages	208-210
	Keywords	Animals; Behavior, Animal; Colic/epidemiology/veterinary; Exploratory Behavior; Feeding Behavior; Female; Horse Diseases/epidemiology; Horses; Incidence; New York/epidemiology; *Placenta; Postpartum Period; Pregnancy; Prevalence; Questionnaires
	Abstract
	Address	Animal Behavior Clinic, Cornell University Hospital for Animals, College of Veterinary Medicine, Cornell University, Ithaca, New York 14853, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0425-1644	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11266073			Approved	no
	Call Number	refbase @ user @			Serial	31
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	Author	Houpt, K.A.
	Title	New perspectives on equine stereotypic behaviour			Type
	Year	1995	Publication	Equine veterinary journal	Abbreviated Journal	Equine Vet J
	Volume	27	Issue	2	Pages	82-83
	Keywords	Animals; Horses/psychology; Stereotyped Behavior/physiology
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0425-1644	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:7607153			Approved	no
	Call Number	refbase @ user @			Serial	34
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