| Records |
| Author |
Bobbert, M.F.; Santamaria, S. |
| Title |
Contribution of the forelimbs and hindlimbs of the horse to mechanical energy changes in jumping |
Type |
Journal Article |
| Year |
2005 |
Publication |
The Journal of Experimental Biology |
Abbreviated Journal |
J Exp Biol |
| Volume |
208 |
Issue |
2 |
Pages |
249-260 |
| Keywords |
Animals; Biomechanics; Forelimb/*physiology; Hindlimb/*physiology; Horses/*physiology; Locomotion/*physiology; Muscle, Skeletal/*physiology; Time Factors |
| Abstract |
The purpose of the present study was to gain more insight into the contribution of the forelimbs and hindlimbs of the horse to energy changes during the push-off for a jump. For this purpose, we collected kinematic data at 240 Hz from 23 5-year-old Warmbloods (average mass: 595 kg) performing free jumps over a 1.15 m high fence. From these data, we calculated the changes in mechanical energy and the changes in limb length and joint angles. The force carried by the forelimbs and the amount of energy stored was estimated from the distance between elbow and hoof, assuming that this part of the leg behaved as a linear spring. During the forelimb push, the total energy first decreased by 3.2 J kg(-1) and then increased again by 4.2 J kg(-1) to the end of the forelimb push. At the end of the forelimb push, the kinetic energy due to horizontal velocity of the centre of mass was 1.6 J kg(-1) less than at the start, while the effective energy (energy contributing to jump height) was 2.3 J kg(-1) greater. It was investigated to what extent these changes could involve passive spring-like behaviour of the forelimbs. The amount of energy stored and re-utilized in the distal tendons during the forelimb push was estimated to be on average 0.4 J kg(-1) in the trailing forelimb and 0.23 J kg(-1) in the leading forelimb. This means that a considerable amount of energy was first dissipated and subsequently regenerated by muscles, with triceps brachii probably being the most important contributor. During the hindlimb push, the muscles of the leg were primarily producing energy. The total increase in energy was 2.5 J kg(-1) and the peak power output amounted to 71 W kg(-1). |
| Address |
Institute for Fundamental and Clinical Human Movement Sciences, Vrije Universiteit, van der Boechorstraat 9, NL-1081 BT Amsterdam, The Netherlands. MFBobbert@fbw.vu.nl |
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0022-0949 |
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| Notes |
PMID:15634844 |
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no |
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Serial |
1895 |
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| Author |
Etienne, A.S.; Maurer, R.; Seguinot, V. |
| Title |
Path integration in mammals and its interaction with visual landmarks |
Type |
Journal Article |
| Year |
1996 |
Publication |
The Journal of Experimental Biology |
Abbreviated Journal |
J Exp Biol |
| Volume |
199 |
Issue |
Pt 1 |
Pages |
201-209 |
| Keywords |
Animals; Cognition/physiology; Cricetinae; Gerbillinae; Humans; Locomotion/*physiology; Mammals/*physiology; Mesocricetus; Mice; Proprioception/physiology; Rats; Visual Pathways/*physiology; Visual Perception/*physiology |
| Abstract |
During locomotion, mammals update their position with respect to a fixed point of reference, such as their point of departure, by processing inertial cues, proprioceptive feedback and stored motor commands generated during locomotion. This so-called path integration system (dead reckoning) allows the animal to return to its home, or to a familiar feeding place, even when external cues are absent or novel. However, without the use of external cues, the path integration process leads to rapid accumulation of errors involving both the direction and distance of the goal. Therefore, even nocturnal species such as hamsters and mice rely more on previously learned visual references than on the path integration system when the two types of information are in conflict. Recent studies investigate the extent to which path integration and familiar visual cues cooperate to optimize the navigational performance. |
| Address |
Laboratoire d'Ethologie, FPSE, Universite de Geneve, Carouge, Switzerland |
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English |
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0022-0949 |
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| Notes |
PMID:8576691 |
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no |
| Call Number |
Equine Behaviour @ team @ |
Serial |
2758 |
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| Author |
Dutto, D.J.; Hoyt, D.F.; Clayton, H.M.; Cogger, E.A.; Wickler, S.J. |
| Title |
Moments and power generated by the horse (Equus caballus) hind limb during jumping |
Type |
Journal Article |
| Year |
2004 |
Publication |
The Journal of Experimental Biology |
Abbreviated Journal |
J Exp Biol |
| Volume |
207 |
Issue |
Pt 4 |
Pages |
667-674 |
| Keywords |
Animals; Biomechanics; Hindlimb/*physiology; Horses/*physiology; Locomotion/*physiology |
| Abstract |
The ability to jump over an obstacle depends upon the generation of work across the joints of the propelling limb(s). The total work generated by one hind limb of a horse and the contribution to the total work by four joints of the hind limb were determined for a jump. It was hypothesized that the hip and ankle joints would have extensor moments performing positive work, while the knee would have a flexor moment and perform negative work during the jump. Ground reaction forces and sagittal plane kinematics were simultaneously recorded during each jumping trial. Joint moment, power and work were determined for the metatarsophalangeal (MP), tarsal (ankle), tibiofemoral (knee) and coxofemoral (hip) joints. The hip, knee and ankle all flexed and then extended and the MP extended and then flexed during ground contact. Consistent with our hypothesis, large extensor moments were observed at the hip and ankle joints and large flexor moments at the knee and MP joints throughout ground contact of the hind limb. Peak moments tended to occur earlier in stance in the proximal joints but peak power generation of the hind limb joints occurred at similar times except for the MP joint, with the hip and ankle peaking first followed by the MP joint. During the first portion of ground contact (approximately 40%), the net result of the joint powers was the absorption of power. During the remainder of the contact period, the hind limb generated power. This pattern of power absorption followed by power generation paralleled the power profiles of the hip, ankle and MP joints. The total work performed by one hind limb was 0.71 J kg(-1). Surprisingly, the knee produced 85% of the work (0.60 J kg(-1)) done by the hind limb, and the positive work performed by the knee occurred during the first 40% of the take-off. There is little net work generated by the other three joints over the entire take-off. Velocity of the tuber coxae (a landmark on the pelvis of the animal) was negative (downward) during the first 40% of stance, which perhaps reflects the negative work performed to decrease the potential energy during the first 40% of contact. During the final 60% of contact, the hip, ankle and MP joints generate positive work, which is reflected in the increase of the animal's potential energy. |
| Address |
Department of Kinesiology and Health Promotion, California State Polytechnic University, Pomona, CA 91768, USA. ddutto@csupomona.edu |
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English |
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0022-0949 |
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| Notes |
PMID:14718509 |
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no |
| Call Number |
Equine Behaviour @ team @ |
Serial |
3654 |
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| Author |
Witte, T.H.; Knill, K.; Wilson, A.M. |
| Title |
Determination of peak vertical ground reaction force from duty factor in the horse (Equus caballus) |
Type |
Journal Article |
| Year |
2004 |
Publication |
The Journal of Experimental Biology |
Abbreviated Journal |
J Exp Biol |
| Volume |
207 |
Issue |
Pt 21 |
Pages |
3639-3648 |
| Keywords |
*Acceleration; Animals; Biomechanics; Forelimb/physiology; *Gait; Hindlimb/physiology; Horses/*physiology; Locomotion/*physiology; Telemetry; Time Factors |
| Abstract |
Measurement of peak vertical ground reaction force (GRFz) from multiple limbs simultaneously during high-speed, over-ground locomotion would enhance our understanding of the locomotor mechanics of cursorial animals. Here, we evaluate the accuracy of predicting peak GRFz from duty factor (the proportion of the stride for which the limb is in contact with the ground). Foot-mounted uniaxial accelerometers, combined with UHF FM telemetry, are shown to be practical and accurate for the field measurement of stride timing variables, including duty factor. Direct comparison with the force plate produces a mean error of 2.3 ms and 3.5 ms for the timing of foot on and foot off, respectively, across all gaits. Predictions of peak GRFz from duty factor show mean errors (with positive values indicating an overestimate) of 0.8+/-0.04 N kg(-1) (13%; N=42; mean +/- S.E.M.) at walk, -0.3+/-0.06 N kg(-1) (3%; N=75) at trot, -2.3+/-0.27 N kg(-1) (16%; N=18) for the non-lead limb at canter and +2.1+/-0.7 N kg(-1) (19%; N=9) for the lead limb at canter. The substantial over- and underestimate seen at canter, in the lead and non-lead limbs, respectively, is attributed to the different functions performed by the two limbs in the asymmetrical gaits. The difference in load experienced by the lead and non-lead limbs decreased with increasing speed. |
| Address |
Structure and Motion Lab, The Royal Veterinary College, Hawkshead Lane, Hatfield, Hertfordshire, AL9 7TA, UK |
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English |
Summary Language |
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0022-0949 |
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| Notes |
PMID:15371472 |
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no |
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Equine Behaviour @ team @ |
Serial |
3658 |
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| Author |
Wilson, A.M.; McGuigan, M.P.; Su, A.; van Den Bogert, A.J. |
| Title |
Horses damp the spring in their step |
Type |
Journal Article |
| Year |
2001 |
Publication |
Nature |
Abbreviated Journal |
Nature |
| Volume |
414 |
Issue |
6866 |
Pages |
895-899 |
| Keywords |
Animals; Biomechanics; Elasticity; Forelimb; Gait; Horses/anatomy & histology/*physiology; Leg Bones/*physiology; Locomotion; Models, Biological; Muscle Fibers/physiology; Muscle, Skeletal/anatomy & histology/*physiology; Tendons/anatomy & histology/*physiology; Vibration |
| Abstract |
The muscular work of galloping in horses is halved by storing and returning elastic strain energy in spring-like muscle-tendon units.These make the legs act like a child's pogo stick that is tuned to stretch and recoil at 2.5 strides per second. This mechanism is optimized by unique musculoskeletal adaptations: the digital flexor muscles have extremely short fibres and significant passive properties, whereas the tendons are very long and span several joints. Length change occurs by a stretching of the spring-like digital flexor tendons rather than through energetically expensive length changes in the muscle. Despite being apparently redundant for such a mechanism, the muscle fibres in the digital flexors are well developed. Here we show that the mechanical arrangement of the elastic leg permits it to vibrate at a higher frequency of 30-40 Hz that could cause fatigue damage to tendon and bone. Furthermore, we show that the digital flexor muscles have minimal ability to contribute to or regulate significantly the 2.5-Hz cycle of movement, but are ideally arranged to damp these high-frequency oscillations in the limb. |
| Address |
Department of Veterinary Basic Sciences, The Royal Veterinary College, Hatfield, Herts AL9 7TA, UK. awilson@rvc.ac.uk |
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English |
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0028-0836 |
ISBN |
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| Notes |
PMID:11780059 |
Approved |
no |
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Equine Behaviour @ team @ |
Serial |
2300 |
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| Author |
Ruggieri, V. |
| Title |
The running horse stops: the hypothetical role of the eyes in imagery of movement |
Type |
Journal Article |
| Year |
1999 |
Publication |
Perceptual and motor skills |
Abbreviated Journal |
Percept Mot Skills |
| Volume |
89 |
Issue |
3 Pt 2 |
Pages |
1088-1092 |
| Keywords |
Adult; Cerebral Cortex/physiology; Eye Movements/*physiology; Female; Functional Laterality/physiology; Head/physiology; Humans; *Imagination; Models, Neurological; *Motion Perception; Movement/physiology; *Ocular Physiology; *Optical Illusions; Posture/physiology |
| Abstract |
To examine the hypothetical role of the eyes in visual mental imagery of movement 72 undergraduate women students in psychology were asked to imagine a running horse and then to produce the same mental image without moving the eyes and the head. In 59% of the subjects interesting modifications of the imagined movement appeared: 37% observed an inhibition of the movement and 19% an evident slowing up of the moving figure. The interpretation of this result was made by hypothesizing that the eyes are concretely involved in visual imagery processes. |
| Address |
University of Rome La Sapienza |
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English |
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0031-5125 |
ISBN |
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PMID:10710755 |
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no |
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refbase @ user @ |
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94 |
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| Author |
De Boyer Des Roches, A.; Richard-Yris, M.-A.; Henry, S.; Ezzaouia, M.; Hausberger, M. |
| Title |
Laterality and emotions: visual laterality in the domestic horse (Equus caballus) differs with objects' emotional value |
Type |
Journal Article |
| Year |
2008 |
Publication |
Physiology & Behavior |
Abbreviated Journal |
Physiol. Behav. |
| Volume |
94 |
Issue |
3 |
Pages |
487-490 |
| Keywords |
Animals; Animals, Newborn; Behavior, Animal/physiology; Dominance, Ocular/*physiology; *Emotions; Exploratory Behavior/physiology; Female; Horses/*physiology; Olfactory Pathways/physiology; Pattern Recognition, Visual/*physiology; Photic Stimulation; Pregnancy; Statistics, Nonparametric; Visual Fields/physiology |
| Abstract |
Lateralization of emotions has received great attention in the last decades, both in humans and animals, but little interest has been given to side bias in perceptual processing. Here, we investigated the influence of the emotional valence of stimuli on visual and olfactory explorations by horses, a large mammalian species with two large monocular visual fields and almost complete decussation of optic fibres. We confronted 38 Arab mares to three objects with either a positive, negative or neutral emotional valence (novel object). The results revealed a gradient of exploration of the 3 objects according to their emotional value and a clear asymmetry in visual exploration. When exploring the novel object, mares used preferentially their right eyes, while they showed a slight tendency to use their left eyes for the negative object. No asymmetry was evidenced for the object with the positive valence. A trend for an asymmetry in olfactory investigation was also observed. Our data confirm the role of the left hemisphere in assessing novelty in horses like in many vertebrate species and the possible role of the right hemisphere in processing negative emotional responses. Our findings also suggest the importance of both hemispheres in the processing positive emotions. This study is, to our knowledge, the first to demonstrate clearly that the emotional valence of a stimulus induces a specific visual lateralization pattern. |
| Address |
UMR CNRS 6552 Ethologie-Evolution-Ecologie, Universite de Rennes 1, Avenue du General Leclerc, Campus de Beaulieu, F-35042 Rennes Cedex, France. a.de-boyer@wanadoo.fr |
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English |
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0031-9384 |
ISBN |
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| Notes |
PMID:18455205 |
Approved |
no |
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Equine Behaviour @ team @ |
Serial |
4762 |
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| Author |
Sankey, C.; Henry, S.; Clouard, C.; Richard-Yris, M.-A.; Hausberger, M. |
| Title |
Asymmetry of behavioral responses to a human approach in young naive vs. trained horses |
Type |
Journal Article |
| Year |
2011 |
Publication |
Physiology & Behavior |
Abbreviated Journal |
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| Volume |
104 |
Issue |
3 |
Pages |
464-468 |
| Keywords |
Laterality; Emotionality; Human approach; Horse |
| Abstract |
The aim of this study was to investigate the impact of training experience on young horses (Equus caballus)’ lateralized responses to an approaching human. The results show that the one year old untrained horses display asymmetrical responses to an approaching human, with more negative reactions (escapes, threats) when approached from the left side, while approaches towards the right shoulder elicited more positive behaviors. On the contrary, two years old trained horses reacted equally positively to approaches and contact on both sides. Our findings support those of previous studies investigating a link between emotionality and laterality and confirm the role of the left hemisphere in the processing of novel or negative stimuli. Moreover, the data underline the impact work and training can have on this laterality in horses. |
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0031-9384 |
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no |
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Equine Behaviour @ team @ |
Serial |
5688 |
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Leliveld, L.M.C.; Düpjan, S.; Tuchscherer, A.; Puppe, B. |
| Title |
Behavioural and physiological measures indicate subtle variations in the emotional valence of young pigs |
Type |
Journal Article |
| Year |
2016 |
Publication |
Physiology & Behavior |
Abbreviated Journal |
Physiol. Behav. |
| Volume |
157 |
Issue |
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Pages |
116-124 |
| Keywords |
Emotion; Heart rate; Vocalisation; Emotional valence; Animal welfare; Domestic pig |
| Abstract |
Abstract In the study of animal emotions, emotional valence has been found to be difficult to measure. Many studies of farm animals' emotions have therefore focussed on the identification of indicators of strong, mainly negative, emotions. However, subtle variations in emotional valence, such as those caused by rather moderate differences in husbandry conditions, may also affect animals' mood and welfare when such variations occur consistently. In this study, we investigated whether repeated moderate aversive or rewarding events could lead to measurable differences in emotional valence in young, weaned pigs. We conditioned 105 female pigs in a test arena to either a repeated startling procedure (sudden noises or appearances of objects) or a repeated rewarding procedure (applesauce, toy and straw) over 11 sessions. Control pigs were also regularly exposed to the same test arena but without conditioning. Before and after conditioning, we measured heart rate and its variability as well as the behavioural reactions of the subjects in the test arena, with a special focus on detailed acoustic analyses of their vocalisations. The behavioural and heart rate measures were analysed as changes compared to the baseline values before conditioning. A limited number of the putative indicators of emotional valence were affected by the conditioning. We found that the negatively conditioned pigs showed changes that were significantly different from those in control pigs, namely a decrease in locomotion and an increase in standing. The positively conditioned pigs, however, showed a stronger increase in heart rate and a smaller decrease in SDNN (a heart rate variability parameter indicating changes in autonomic regulation) compared to the controls. Compared to the negatively conditioned pigs, the positively conditioned pigs produced fewer vocalisations overall as well as fewer low-frequency grunts but more high-frequency grunts. The low-frequency grunts of the negatively conditioned pigs also showed lower frequency parameters (bandwidth, maximum frequency, 25% and 50% quartiles) compared to those of the positively conditioned pigs. In any of the statistically significant results, the conditioning accounted for 1.5–11.9% of variability in the outcome variable. Hence, we conclude that repeated moderate aversive and rewarding events have weak but measurable effects on some aspects of behaviour and physiology in young pigs, possibly indicating changes in emotional valence, which could ultimately affect their welfare. The combination of ethophysiological indicators, i.e., the concurrent examination of heart rate measures, behavioural responses and especially vocalisation patterns, as used in the current study, might be a useful way of examining subtle effects on emotional valence in further studies. |
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0031-9384 |
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Equine Behaviour @ team @ |
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6017 |
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| Author |
Rilling, M.E.; Neiworth, J.J. |
| Title |
How animals use images |
Type |
Journal Article |
| Year |
1991 |
Publication |
Science Progress |
Abbreviated Journal |
Sci Prog |
| Volume |
75 |
Issue |
298 Pt 3-4 |
Pages |
439-452 |
| Keywords |
Animals; Association Learning; Columbidae; *Concept Formation; *Imagination; *Mental Recall; Motion Perception; Problem Solving; *Thinking; *Visual Perception |
| Abstract |
Animal cognition is a field within experimental psychology in which cognitive processes formerly studied exclusively with people have been demonstrated in animals. Evidence for imagery in the pigeon emerges from the experiments described here. The pigeon's task was to discriminate, by pecking the appropriate choice key, between a clock hand presented on a video screen that rotated clockwise with constant velocity from a clock hand that violated constant velocity. Imagery was defined by trials on which the line rotated from 12.00 o'clock to 3.00 o'clock, then disappeared during a delay, and reappeared at a final stop location beyond 3.00 o'clock. After acquisition of a discrimination with final stop locations at 3.00 o'clock and 6.00 o'clock, the evidence for imagery was the accurate responding of the pigeons to novel locations at 4.00 o'clock and 7.00 o'clock. Pigeons display evidence of imagery by transforming a representation of movement that includes a series of intermediate steps which accurately represent the location of a moving stimulus after it disappears. |
| Address |
Department of Psychology, Michigan State University, East Lansing 48824 |
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English |
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0036-8504 |
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| Notes |
PMID:1842858 |
Approved |
no |
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Equine Behaviour @ team @ |
Serial |
2831 |
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