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	Author	Shettleworth, S.J.; Krebs, J.R.
	Title	How marsh tits find their hoards: the roles of site preference and spatial memory			Type	Journal Article
	Year	1982	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	8	Issue	4	Pages	354-375
	Keywords	Animals; Appetitive Behavior; Birds; Cues; Discrimination Learning; Memory; Mental Recall; Orientation; *Space Perception
	Abstract	Marsh tits (Parus palustris) store single food items in scattered locations and recover them hours or days later. Some properties of the spatial memory involved were analyzed in two laboratory experiments. In the first, marsh tits were offered 97 sites for storing 12 seeds. They recovered a median of 65% of them 2-3 hr later, making only two errors per seed while doing so. Over trials, they used some sites more often than others, but during recovery they were more likely to visit a site of any preference value if they had stored a seed there that day than if they had not. Recovery performance was much worse if the experimenters moved the seeds between storage and recovery. A fixed search strategy that had some of the same average properties as the tits' search behavior also did worse than the real birds. In Experiment 2, any tendency to visit the same sites on successive daily tests in the aviary was placed in opposition to memory for storage sites by allowing the tits to store more seeds 2 hr after storing a first batch. They tended to avoid individual storage sites holding seeds from the first batch. When the tits searched for all the seeds 2 hr later, they tended to recover more seeds from the second batch than from the first, i.e., there was a recency effect.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:7175447			Approved	no
	Call Number	refbase @ user @			Serial	385
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	Author	Shettleworth, S.J.; Juergensen, M.R.
	Title	Reinforcement and the organization of behavior in golden hamsters: brain stimulation reinforcement for seven action patterns			Type	Journal Article
	Year	1980	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	6	Issue	4	Pages	352-375
	Keywords	Animals; Behavior, Animal/physiology; Cricetinae; Electric Stimulation; Female; Hypothalamus/physiology; Male; Medial Forebrain Bundle/physiology; Mesocricetus; *Reinforcement (Psychology)
	Abstract	Golden hamsters were reinforced with intracranial electrical stimulation of the lateral hypothalamus (ICS) for spending time engaging in one of seven topographically defined action patterns (APs). The stimulation used as reinforcer elicited hoarding and/or feeding and supported high rates of bar pressing. In Experiment 1, hamsters were reinforced successively for digging, open rearing, and face washing. Digging increased most in time spent, and face washing increased least. Experiments 2-5 examined these effects further and also showed that “scrabbling,” like digging, was performed a large proportion of the time, almost without interruption, for contingent ICS but that scratching the body with a hindleg and scent-marking showed relatively little effect of contingent ICS, the latter even in an environment that facilitated marking. In Experiment 6, naive hamsters received ICS not contingent on behavior every 30 sec (fixed-time 30-sec schedule). Terminal behaviors that developed on this schedule were APs that were easy to reinforce in the other experiments, but a facultative behavior, face washing, was one not so readily reinforced. Experiment 7 confirmed a novel prediction from Experiment 6--that wall rearing, a terminal AP, would be performed at a high level for contingent ICS. All together, the results point to both motivational factors and associative factors being involved in the considerable differences in performance among different reinforced activities.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:6968817			Approved	no
	Call Number	refbase @ user @			Serial	386
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	Author	Shettleworth, S.J.
	Title	Reinforcement and the organization of behavior in golden hamsters: Pavlovian conditioning with food and shock unconditioned stimuli			Type	Journal Article
	Year	1978	Publication	Journal of experimental psychology. Animal behavior processes	Abbreviated Journal	J Exp Psychol Anim Behav Process
	Volume	4	Issue	2	Pages	152-169
	Keywords	Acoustic Stimulation; Animals; Behavior, Animal; Conditioning, Classical; Conditioning, Operant; Cricetinae; Electroshock; Female; Food; Male; Punishment; *Reinforcement (Psychology); Reinforcement Schedule
	Abstract	The effects of Pavlovian conditioned stimuli (CSs) for food or shock on a variety of behaviors of golden hamsters were observed in three experiments. The aim was to see whether previously reported differences among the behaviors produced by food reinforcement and punishment procedures could be accounted for by differential effects of Pavlovian conditioning on the behaviors. There was some correspondence between the behaviors observed to the CSs and the previously reported effects of instrumental training. However, the Pavlovian conditioned responses (CRs) alone would not have predicted the effects of instrumental training. Moreover, CRs depended to some extent on the context in which training and testing occurred. These findings, together with others in the literature, suggest that the results of Pavlovian conditioning procedures may not unambiguously predict what system of behaviors will be most readily modified by instrumental training with a given reinforcer.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0097-7403	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:670890			Approved	no
	Call Number	refbase @ user @			Serial	387
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	Author	Gibson, B.M.; Shettleworth, S.J.; McDonald, R.J.
	Title	Finding a goal on dry land and in the water: differential effects of disorientation on spatial learning			Type	Journal Article
	Year	2001	Publication	Behavioural brain research	Abbreviated Journal	Behav. Brain. Res.
	Volume	123	Issue	1	Pages	103-111
	Keywords	Animals; Cues; Environment; Male; Maze Learning/physiology; Orientation/physiology; Rats; Rats, Long-Evans; Spatial Behavior/*physiology; Water
	Abstract	Two previous studies, Martin et al. (J. Exp. Psychol. Anim. Behav. Process. 23 (1997) 183) and Dudchenko et al. (J. Exp. Psychol. Anim. Behav. Process. 23 (1997) 194), report that, compared to non-disoriented controls, rats disoriented before testing were disrupted in their ability to learn the location of a goal on a dry radial-arm maze task, but that both groups learned at the same rate in the Morris water maze. However, the radial-arm maze task was much more difficult than the water maze. In the current set of experiments, we examined the performance of control and disoriented rats on more comparable dry land and water maze tasks. Compared to non-disoriented rats, rats that were disoriented before testing were significantly impaired in locating a goal in a circular dry arena, but not a water tank. The results constrain theoretical explanations for the differential effects of disorientation on different spatial tasks.
	Address	Department of Psychology, University of Toronto, 100 St. George Street, Toronto, Ontario, Canada M5S 3G3. gibson@psych.utoronto.ca
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0166-4328	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:11377733			Approved	no
	Call Number	refbase @ user @			Serial	372
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	Author	Shettleworth, S.J.
	Title	Cognitive ecology: field or label?			Type	Journal Article
	Year	2000	Publication	Trends in Ecology & Evolution	Abbreviated Journal	Trends. Ecol. Evol
	Volume	15	Issue	4	Pages	161
	Keywords
	Abstract
	Address	Depts of Psychology and Zoology, University of Toronto, Toronto, Ontario, Canada M5S 3G3
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0169-5347	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:10717686			Approved	no
	Call Number	refbase @ user @			Serial	373
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	Author	Sekuler, A.B.; Lee, J.A.; Shettleworth, S.J.
	Title	Pigeons do not complete partly occluded figures			Type	Journal Article
	Year	1996	Publication	Perception	Abbreviated Journal	Perception
	Volume	25	Issue	9	Pages	1109-1120
	Keywords	Animals; Columbidae; Visual Perception
	Abstract	One of the most common obstacles to object perception is the fact that objects often occlude parts of themselves and parts of other objects. Perceptual completion has been studied extensively in humans, and researchers have shown that humans do complete partly occluded objects. In an effort to understand more about the mechanisms underlying completion, recent research has extended the study of perceptual completion to other mammalian species. Monkeys and mice also seem to complete two-dimensional representations of partly occluded objects. The present study addresses the question of whether this capacity generalizes to a nonmammalian species, the pigeon (Columba livia). The results point to a limit of the generalizability of perceptual completion: pigeons do not complete partly occluded figures.
	Address	Department of Psychology, University of Toronto, ON, Canada
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0301-0066	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:8983050			Approved	no
	Call Number	refbase @ user @			Serial	377
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	Author	Shettleworth, S.J.
	Title	Taking the best for learning			Type	Journal Article
	Year	2005	Publication	Behavioural processes	Abbreviated Journal	Behav. Process.
	Volume	69	Issue	2	Pages	147-9; author reply 159-63
	Keywords	Algorithms; Animals; Behavior, Animal; Decision Making; Evolution; Learning; Models, Theoretical
	Abstract	Examples of how animals learn when multiple, sometimes redundant, cues are present provide further examples not considered by Hutchinson and Gigerenzer that seem to fit the principle of taking the best. “The best” may the most valid cue in the present circumstances; evolution may also produce species-specific biases to use the most functionally relevant cues.
	Address	Department of Psychology, University of Toronto, Toronto, Ont., Canada M5S 3G3. shettle@psych.utoronto.ca
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15845301			Approved	no
	Call Number	refbase @ user @			Serial	361
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	Author	Sutton, J.E.; Shettleworth, S.J.
	Title	Internal sense of direction and landmark use in pigeons (Columba livia)			Type	Journal Article
	Year	2005	Publication	Journal of comparative psychology (Washington, D.C. : 1983)	Abbreviated Journal	J Comp Psychol
	Volume	119	Issue	3	Pages	273-284
	Keywords	Animals; Columbidae; Conflict (Psychology); Cues; Discrimination Learning; Homing Behavior; Intuition; Orientation; Space Perception; Transfer (Psychology); Visual Perception
	Abstract	The relative importance of an internal sense of direction based on inertial cues and landmark piloting for small-scale navigation by White King pigeons (Columba livia) was investigated in an arena search task. Two groups of pigeons differed in whether they had access to visual cues outside the arena. In Experiment 1, pigeons were given experience with 2 different entrances and all pigeons transferred accurate searching to novel entrances. Explicit disorientation before entering did not affect accuracy. In Experiments 2-4, landmarks and inertial cues were put in conflict or tested 1 at a time. Pigeons tended to follow the landmarks in a conflict situation but could use an internal sense of direction to search when landmarks were unavailable.
	Address	Department of Psychology, University of Toronto, ON, Canada. jsutton7@uwo.ca
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16131256			Approved	no
	Call Number	refbase @ user @			Serial	360
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	Author	Jones, J.E.; Antoniadis, E.; Shettleworth, S.J.; Kamil, A.C.
	Title	A comparative study of geometric rule learning by nutcrackers (Nucifraga columbiana), pigeons (Columba livia), and jackdaws (Corvus monedula)			Type	Journal Article
	Year	2002	Publication	Journal of comparative psychology (Washington, D.C. : 1983)	Abbreviated Journal	J Comp Psychol
	Volume	116	Issue	4	Pages	350-356
	Keywords	Animals; Behavior, Animal/physiology; Birds; Feeding Behavior/physiology; Learning/physiology; Mathematics; Random Allocation; Spatial Behavior/*physiology
	Abstract	Three avian species, a seed-caching corvid (Clark's nutcrackers; Nucifraga columbiana), a non-seed-caching corvid (jackdaws; Corvus monedula), and a non-seed-caching columbid (pigeons; Columba livia), were tested for ability to learn to find a goal halfway between 2 landmarks when distance between the landmarks varied during training. All 3 species learned, but jackdaws took much longer than either pigeons or nutcrackers. The nutcrackers searched more accurately than either pigeons or jackdaws. Both nutcrackers and pigeons showed good transfer to novel landmark arrays in which interlandmark distances were novel, but inconclusive results were obtained from jackdaws. Species differences in this spatial task appear quantitative rather than qualitative and are associated with differences in natural history rather than phylogeny.
	Address	School of Biological Sciences, University of Nebraska-Lincoln, 68588-0118, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12539930			Approved	no
	Call Number	refbase @ user @			Serial	369
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	Author	Gibson, B.M.; Shettleworth, S.J.
	Title	Place versus response learning revisited: tests of blocking on the radial maze			Type	Journal Article
	Year	2005	Publication	Behavioral neuroscience	Abbreviated Journal	Behav Neurosci
	Volume	119	Issue	2	Pages	567-586
	Keywords	Animals; Association Learning; Male; Maze Learning; Memory; Rats; Rats, Long-Evans; Reproducibility of Results
	Abstract	Neurobiological and behavioral research indicates that place learning and response learning occur simultaneously, in parallel. Such findings seem to conflict with theories of associative learning in which different cues compete for learning. The authors conducted place+response training on a radial maze and then tested place learning and response learning separately by reconfiguring the maze in various ways. Consistent with the effects of manipulating place and response systems in the brain (M. G. Packard & J. L. McGaugh, 1996), well-trained rats showed strong place learning and strong response learning. Three experiments using associative blocking paradigms indicated that prior response learning interferes with place learning. Blocking and related tests can be used to better understand how memory systems interact during learning.
	Address	Department of Psychology, University of New Hampshire, Durham, NH 03824-3567, USA. bgibson@cisunix.unh.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7044	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15839803			Approved	no
	Call Number	refbase @ user @			Serial	362
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