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Author Lee, J.; Floyd, T.; Erb, H.; Houpt, K. url  doi
openurl 
  Title Preference and demand for exercise in stabled horses Type Journal Article
  Year 2011 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 130 Issue 3-4 Pages (up) 91-100  
  Keywords Horse; Welfare; Exercise; Operant conditioning; Two choice preference; Treadmill  
  Abstract Operant conditioning and two choice preference tests were used to assess the motivation of horses to be released from straight and from box stalls. The motivations for food, a companion, and release into a paddock were compared when the horses had to work for each commodity at increasing fixed ratios of responses (panel presses) to reward in an equine operant conditioning stall. The motivation for food (mean ± SEM = 258 ± 143) responses was much greater than that for either release (38 ± 32) from a straight stall into a large paddock alone or into a small paddock with another horse (95 ± 41) (P = 0.04). When given a two choice preference test between exercise on a treadmill for 20 min or returning to their box stalls, eight of nine horses chose to return to their stalls. In a two choice preference test six of eight horses in box stalls chose to be released into a paddock alone. Horses were given a series of two choice preference tests to determine how long they preferred to be in a paddock. After 15 min in the paddock the horses were re-tested, but all chose the paddock when released into a paddock with three other horses. They were retested every 15 min until they chose to return to their stalls. They chose to stay out for 35 ± 6 min when other horses were in the paddock but for only 17 ± 2 min when they would be alone. When deprived of stall release for 48 h the horses chose to remain in the paddock with other horses for 54 ± 6 min, but showed no compensatory behavior when they were alone (duration chosen = 16 ± 4 min). These findings indicate that horses are not strongly motivated to exercise alone and will choose not to endure forced exercise on a treadmill. The social context of voluntary exercise is important; horses are willing to stay out of their stalls longer if other horses are present and will show compensatory behavior only if other horses are present. These finding have implications for optimizing turnout time for stalled horses.  
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  ISSN 0168-1591 ISBN Medium  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5330  
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Author McLean, A.N. url  doi
openurl 
  Title Short-term spatial memory in the domestic horse Type Journal Article
  Year 2004 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 85 Issue 1-2 Pages (up) 93-105  
  Keywords Animal cognition; Delayed response; Equine memory; Horse; Object permanence; Temporal cognition  
  Abstract This study investigates the ability of horses to recall a feeding event in a two-point choice apparatus. Twelve horses were individually tested whereby they were maintained immobile in a test arena and visually and aurally experienced the delivery of food into one of two feed goals. The horses were then released to make their choice in two experimental contexts: immediate release after experiencing the delivery of food, and release 10 s after food delivery. Each horse performed 40 immediate-release (IR) trials, followed by forty 10-s release trials over a 3-day period. In addition, the same horses were tested 3 months later in the spring with the same number and sequence of trials. Results were analysed by log-linear analysis of frequencies. Results showed that while horses were able to achieve the correct feed goal choice in the immediate-release trials, they were unsuccessful with the 10-s release trials. This suggests that there are limitations in recall abilities in horses, in that they may not possess a prospective type of memory. There are welfare and training implications in these findings concerning the effects of overestimating the mental abilities of horses during training and the effects of delays in reinforcements.  
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  Notes Approved no  
  Call Number refbase @ user @ Serial 403  
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Author Smith-Funk, E.D.; Crowell-Davis, S.L. url  openurl
  Title Maternal behavior of draft mares (Equus caballus) with mule foals (Equus asinus x Equus caballus) Type Journal Article
  Year 1992 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 33 Issue 2-3 Pages (up) 93-119  
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  Abstract Draft mares and their mule foals were observed from the day of birth to Week 17 of each foal's life. The rate of nursing was recorded and the duration of nursing activity to the nearest second. The rate at which foals engaged in nursing activity varied at each age. The duration of nursing bouts varied slightly as the foals matured. Aggression was recorded during both nursing and non-nursing activity for both the mares and foals. Maternal aggression was highest during nursing activity, especially during the pre-nurse nuzzling period. Maternal aggression increased as the foals matured. Mother-directed foal aggression was primarily in response to maternal aggression. Spatial relationships between each focal dyad were recorded when the foals were upright, not nursing and when they were recumbent. Spatial relationships differed based on the foal's state. The activity in which the mare engaged while her foal was recumbent was recorded. The movements of the mares were also recorded during foal recumbency. Mares approached or maintained their distance from their recumbent foal more than they left their recumbent foal in all weeks of the study, except Week 2.  
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  Call Number Equine Behaviour @ team @ Serial 2272  
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Author Lamoot, I.; Callebaut, J.; Demeulenaere, E.; Vandenberghe, C.; Hoffmann, M. url  doi
openurl 
  Title Foraging behaviour of donkeys grazing in a coastal dune area in temperate climate conditions Type Journal Article
  Year 2005 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 92 Issue 1-2 Pages (up) 93-112  
  Keywords Grazing behaviour; Habitat use; Donkey; Equid; Diet composition  
  Abstract A small herd of donkeys was introduced in a coastal dune reserve `Houtsaegerduinen' (ca. 80 ha) in Belgium, in order to slow down expansion of dominant grass and shrub species. The Houtsaegerduinen is a nutrient poor scrub-dominated dune system with a spatially heterogeneous vegetation pattern. Different aspects of the grazing behaviour (grazing time, bite rate, habitat use, diet composition) of the free-ranging donkeys are described and analysed. Behavioural data (of maximum six adult mares) were collected through continuous focal animal observation in three consecutive years (1998-2001). Temporal variation in grazing time, habitat use and diet composition was determined. During daylight, donkeys spent most of their time on grazing (56%). In all 3 years, grazing time was significantly shorter in summer (45% of their time), longest grazing times were achieved in spring (64%). In spring, the donkeys also achieved the highest bite rate (21.5 bites/min). The grassy habitat was preferred for foraging in all seasons, while the use of scrub and woodland was variable over time. Averaged over the four seasons, the general diet consisted for 80% of graminoids, 10% of forbs and 10% of woody plants. However, diet composition varied not only among seasons and years, but depended also on the foraged habitat type. We discuss the possible role of the donkeys in nature management.  
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  Call Number Equine Behaviour @ team @ Serial 2338  
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Author Heffner, R.S.; Heffner, H.E. url  openurl
  Title Localization of tones by horses: use of binaural cues and the role of the superior olivary complex Type Journal Article
  Year 1986 Publication Behavioral Neuroscience Abbreviated Journal Behav Neurosci  
  Volume 100 Issue 1 Pages (up) 93-103  
  Keywords Animals; Auditory Pathways/physiology; Auditory Perception/*physiology; Avoidance Learning/physiology; Brain Mapping; Electroshock; Female; Horses/*physiology; Male; Olivary Nucleus/anatomy & histology/*physiology; Orientation/physiology; Pitch Perception/physiology; Sound Localization/*physiology  
  Abstract The ability of horses to use binaural time and intensity difference cues to localize sound was assessed in free-field localization tests by using pure tones. The animals were required to discriminate the locus of a single tone pip ranging in frequency from 250 Hz to 25 kHz emitted by loudspeakers located 30 degrees to the left and right of the animals' midline (60 degrees total separation). Three animals were tested with a two-choice procedure; 2 additional animals were tested with a conditioned avoidance procedure. All 5 animals were able to localize 250 Hz, 500 Hz, and 1 kHz but were completely unable to localize 2 kHz and above. Because the frequency of ambiguity for the binaural phase cue delta phi for horses in this test was calculated to be 1.5 kHz, these results indicate that horses can use binaural time differences in the form of delta phi but are unable to use binaural intensity differences. This finding was supported by an unconditioned orientation test involving 4 additional horses, which showed that horses correctly orient to a 500-Hz tone pip but not to an 8-kHz tone pip. Analysis of the superior olivary complex, the brain stem nucleus at which binaural interactions first take place, reveals that the lateral superior olive (LSO) is relatively small in the horse and lacks the laminar arrangement of bipolar cells characteristic of the LSO of most mammals that can use binaural delta I.  
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  ISSN 0735-7044 ISBN Medium  
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  Notes PMID:3954885 Approved no  
  Call Number Equine Behaviour @ team @ Serial 5634  
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Author Plotnik, J.; Nelson, P.A.; de Waal, F.B.M. openurl 
  Title Visual field information in the face perception of chimpanzees (Pan troglodytes) Type Journal Article
  Year 2003 Publication Annals of the New York Academy of Sciences Abbreviated Journal Ann N Y Acad Sci  
  Volume 1000 Issue Pages (up) 94-98  
  Keywords Animals; *Facial Expression; Pan troglodytes; Recognition (Psychology); Visual Fields/*physiology; Visual Perception/*physiology  
  Abstract Evidence for a visual field advantage (VFA) in the face perception of chimpanzees was investigated using a modification of a free-vision task. Four of six chimpanzee subjects previously trained on a computer joystick match-to-sample paradigm were able to distinguish between images of neutral face chimeras consisting of two left sides (LL) or right sides (RR) of the face. While an individual's ability to make this distinction would be unlikely to determine their suitability for the VFA tests, it was important to establish that distinctive information was available in test images. Data were then recorded on their choice of the LL vs. RR chimera as a match to the true, neutral image; a bias for one of these options would indicate an hemispatial visual field advantage. Results suggest that chimpanzees, unlike humans, do not exhibit a left visual field advantage. These results have important implications for studies on laterality and asymmetry in facial signals and their perception in primates.  
  Address Department of Animal Science, Cornell University, Ithaca, New York 14853, USA. jmp63@cornell.edu  
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  ISSN 0077-8923 ISBN Medium  
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  Notes PMID:14766624 Approved no  
  Call Number refbase @ user @ Serial 175  
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Author Jørgensen, G.H.M.; Borsheim, L.; Mejdell, C.M.; Søndergaard, E.; Bøe, K.E. url  doi
openurl 
  Title Grouping horses according to gender--Effects on aggression, spacing and injuries Type Journal Article
  Year 2009 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 120 Issue 1-2 Pages (up) 94-99  
  Keywords Horse; Group housing; Gender; Injuries; Social behaviour  
  Abstract Many horse owners tend to group horses according to gender, in an attempt to reduce aggressive interactions and the risk of injuries. The aim of our experiment was to test the effects of such gender separation on injuries, social interactions and individual distance in domestic horses. A total of 66 horses were recruited from 4 different farms in Norway and Denmark and divided into six batches. Within each batch, horses were allotted into one mare group, one gelding group and one mixed gender group, with most groups consisting of three or four animals. After 4-6 weeks of acclimatisation, a trained observer recorded all social interactions using direct, continuous observation 1 h in the morning and 1 h in the afternoon for three consecutive days. Recordings of the nearest neighbour of each horse were performed using instantaneous sampling every 10 min. The horses were inspected for injuries before grouping, day 1 after grouping and after 4-6 weeks. No significant effect of gender composition was found on social interactions (P > 0.05), spacing (P > 0.07) or injuries (P > 0.23). Eighty percent of all aggressive interactions recorded were threats, not involving physical contact. Horses with the smallest space allowance showed the highest mean number of aggressive interactions (28.6 ± 6.1 interactions per 6 h) compared to the mean of all the other batches (8.3 ± 1.0 interactions per 6 h). Very few injuries were found and most were superficial. In conclusion, gender composition does not seem to have any effect on aggression level, spacing or injuries. However, the early social experience of horses, management of feeding and space allowance probably represents more important factors for successful group housing of domestic horses.  
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  ISSN 0168-1591 ISBN Medium  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 5093  
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Author van Dierendonck, M.C.; Bandi, N.; Batdorj, D.; Dugerlham, S.; Munkhtsog, B. url  openurl
  Title Behavioural observations of reintroduced Takhi or Przewalski horses (Equus ferus przewalskii) in Mongolia Type Journal Article
  Year 1996 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.  
  Volume 50 Issue 2 Pages (up) 95-114  
  Keywords Horse; Przewalski's horse; Reintroduction; Time budget; Social organisation; Basic reference dataset  
  Abstract During 1992 and 1993, 14 reintroduced Przewalski Horses or Takhi (Equus ferus przewalskii) were studied in the Hustain Nuruu Mountain Steppe reserve in Mongolia. Most of the individuals did not know each other before reintroduction. These Takhi were the first of five groups due to be released in the reserve after an acclimatisation period of at least 1 year. During acclimatisation the Takhi, lived visually and acoustically separately, in fenced enclosures of approximately 45 ha each. The observations, mostly scan-sampling, were carried out in each season. The observation bouts were divided over six periods and over two harem herds. Two of the periods were in the same consecutive seasons, so comparison over the years was possible. Social integration within the Takhi herds was very high from the beginning, as described by the spatial relation and synchronisation data. Between 50 and 89% of the observation time, the behaviour of all herd members was synchronised. The amount of time spent grazing by the Takhi (30-68% of the daylight period) was similar to that of feral horses and Takhi in captivity and semi-reserves. The Takhi tended to rest in the morning and have a bimodal period of grazing at dawn and in the afternoon. The Takhi displayed clear habitat preferences for certain activities. They had a strong preference to rest at the highest point in their enclosure. They fed preferably on two or three different vegetation types (with five types available in each enclosure). The amount of time spent grazing during the non-growing seasons (49 +/- 15%) indicates that the feeding value and availability of food were sufficient. Health changes were detected adequately using condition scoring sheets. No supplementary food or water was supplied during the harsh winters. Moreover, low mortality rates and high reproductive success show that the mountain steppe is a habitat which is potentially suitable for establishing a healthy Takhi population. Takhi is the first species to return to its native habitat after living only in zoos for so many generations.  
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  Call Number refbase @ user @ Serial 762  
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Author Thiel, D.; Jenni-Eiermann, S.; Palme, R. doi  openurl
  Title Measuring corticosterone metabolites in droppings of capercaillies (Tetrao urogallus) Type Journal Article
  Year 2005 Publication Annals of the New York Academy of Sciences Abbreviated Journal Ann N Y Acad Sci  
  Volume 1046 Issue Pages (up) 96-108  
  Keywords Adrenocorticotropic Hormone/administration & dosage/analysis/metabolism; Animals; Circadian Rhythm; Corticosterone/administration & dosage/*analysis/*metabolism; Feces/*chemistry; Female; Freezing; Galliformes/*metabolism; Male; Reproducibility of Results; Sex Factors; Temperature; Time Factors; Tritium/diagnostic use  
  Abstract The capercaillie (Tetrao urogallus), the largest grouse species in the world, is decreasing in numbers in major parts of its distribution range. Disturbances by human outdoor activities are discussed as a possible reason for this population decline. An indicator for disturbances is the increase of the glucocorticoid corticosterone, a stress hormone, which helps to cope with life-threatening situations. However, repeated disturbances might result in a long-term increase of the basal corticosterone concentration, which can result in detrimental effects like reduced fitness and survival of an animal. To measure corticosterone metabolites (CMs) noninvasively in the droppings of free-living capercaillies, first an enzyme immunoassay (EIA) in captive birds had to be selected and validated. Therefore, the excretion pattern of intravenously injected radiolabeled corticosterone was determined and 3H metabolites were characterized. High-performance liquid chromatography (HPLC) separations of the samples containing peak concentrations revealed that corticosterone was extensively metabolized. The HPLC fractions were tested in several EIAs for glucocorticoid metabolites. The physiological relevance of this method was proved after pharmacological stimulation of the adrenocortical activity. Only the recently established cortisone assay, measuring CMs with a 3,11-dione structure, detected an expressed increase of concentrations following ACTH stimulation. To set up a sampling protocol suited for the field, we examined the influence of various storage conditions and time of day on concentrations of CMs.  
  Address Swiss Ornithological Institute, 6204 Sempach, Switzerland. dominik.thiel@vogelwarte.ch  
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  Notes PMID:16055846 Approved no  
  Call Number Equine Behaviour @ team @ Serial 4079  
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Author Huff, A.N.; Meacham, T.N.; Wahlberg, M.L. url  doi
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  Title Feeds and feeding: A review Type Journal Article
  Year 1985 Publication Journal of Equine Veterinary Science Abbreviated Journal  
  Volume 5 Issue 2 Pages (up) 96-108  
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  Notes Approved no  
  Call Number Equine Behaviour @ team @ Serial 4667  
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