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Author Wolter, R.; Stefanski, V.; Krueger, K.
Title Parameters for the Analysis of Social Bonds in Horses Type Journal Article
Year 2018 Publication Animals Abbreviated Journal Animals
Volume 8 Issue 11 Pages (down) 191
Keywords feral horses; mutual grooming; social bonds; social bond analysis; spatial proximity
Abstract Social bond analysis is of major importance for the evaluation of social relationships in group housed horses. However, in equine behaviour literature, studies on social bond analysis are inconsistent. Mutual grooming (horses standing side by side and gently nipping, nuzzling, or rubbing each other), affiliative approaches (horses approaching each other and staying within one body length), and measurements of spatial proximity (horses standing with body contact or within two horse-lengths) are commonly used. In the present study, we assessed which of the three parameters is most suitable for social bond analysis in horses, and whether social bonds are affected by individual and group factors. We observed social behaviour and spatial proximity in 145 feral horses, five groups of Przewalskiâ&#65533;&#65533;s horses (N = 36), and six groups of feral horses (N = 109) for 15 h per group, on three days within one week. We found grooming, friendly approaches, and spatial proximity to be robust parameters, as their correlation was affected only by the animalsâ&#65533;&#65533; sex (GLMM: N = 145, SE = 0.001, t = â&#65533;&#65533;2.7, p = 0.008) and the group size (GLMM: N = 145, SE < 0.001, t = 4.255, p < 0.001), but not by the horse breed, the aggression ratio, the social rank, the group, the group composition, and the individuals themselves. Our results show a trend for a correspondence between all three parameters (GLMM: N = 145, SE = 0.004, t = 1.95, p = 0.053), a strong correspondence between mutual grooming and friendly approaches (GLMM: N = 145, SE = 0.021, t = 3.922, p < 0.001), and a weak correspondence between mutual grooming and spatial proximity (GLMM: N = 145, SE = 0.04, t = 1.15, p = 0.25). We therefore suggest either using a combination of the proactive behaviour counts mutual grooming and friendly approaches, or using measurements of close spatial proximity, for the analysis of social bonds in horses within a limited time frame.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 2076-2615 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6428
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Author Drevemo, S.; Fredricson, I.; Hjertén, G.; McMIKEN, D.
Title Early development of gait asymmetries in trotting Standardbred colts Type Journal Article
Year 1987 Publication Equine Veterinary Journal Abbreviated Journal Equine. Vet. J.
Volume 19 Issue 3 Pages (down) 189-191
Keywords
Abstract Summary Ten trotting Standardbred colts were recorded by high-speed cinematography at the ages of eight, 12 and 18 months. The horses were trotting on a treadmill operating at 4.0 m/secs. Five horses were subjected to a programme of intensified training from eight months of age, whereas the others were not trained and acted as controls. The films were analysed on a semi-automatic film-reading equipment and a number of variables used to demonstrate the gait symmetry were calculated and scaled by computer. Certain differences between left and right diagonal and contralateral pair of limbs, respectively, were noted, suggesting that laterality in horses may be inherited. The most pronounced systematic differences were found in 18-month old horses in the trained group. The results show the importance of careful gait examination and comprehensive coordination training at an early age.
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Corporate Author Thesis
Publisher American Medical Association (AMA) Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0425-1644 ISBN Medium
Area Expedition Conference
Notes https://doi.org/10.1111/j.2042-3306.1987.tb01373.x Approved no
Call Number Equine Behaviour @ team @ Serial 6702
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Author McGreevy, P.D.; Harman, A.; McLean, A.; Hawson, L.
Title Over-flexing the horse's neck: A modern equestrian obsession? Type Journal Article
Year 2010 Publication Journal of Veterinary Behavior: Clinical Applications and Research Abbreviated Journal Journal of Veterinary Behavior: Clinical Applications and Research
Volume 5 Issue 4 Pages (down) 180-186
Keywords
Abstract We used an opportunistic review of photographs of different adult and juvenile horses walking, trotting, and cantering (n = 828) to compare the angle of the nasal plane relative to vertical in feral and domestic horses at liberty (n = 450) with ridden horses advertised in a popular Australian horse magazine (n = 378). We assumed that horses in advertisements were shown at, what was perceived by the vendors to be, their best. Of the ridden horses, 68% had their nasal plane behind the vertical. The mean angle of the unridden horses at walk, trot, and canter (30.7 ± 11.5; 27.3 ± 12.0; 25.5 ± 11.0) was significantly greater than those of the ridden horses (1.4 ± 14.1; ?5.1 ± ?11.1; 3.1 ± 15.4, P < 0.001). Surprisingly, unridden domestic horses showed greater angles than feral horses or domestic horses at liberty. We compared adult and juvenile horses in all 3 gaits and found no significant difference. Taken together, these findings demonstrate that the longitudinal neck flexion of the degree desirable by popular opinion in ridden horses is not a common feature of unridden horses moving naturally. Moreover, they suggest that advertised horses in our series are generally being ridden at odds with their natural carriage and contrary to the international rules of dressage (as published by the International Equestrian Federation). These findings are discussed against the backdrop of the established doctrine, which states that carrying a rider necessitates changes in longitudinal flexion, and in the context of the current debate around hyperflexion.
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Corporate Author Thesis
Publisher Elsevier Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 1558-7878 ISBN Medium
Area Expedition Conference
Notes doi: 10.1016/j.jveb.2010.03.004 Approved no
Call Number Equine Behaviour @ team @ Serial 6501
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Author Passilongo, D.; Marchetto, M.; Apollonio, M.
Title Singing in a wolf chorus: structure and complexity of a multicomponent acoustic behaviour Type Journal Article
Year 2017 Publication Hystrix, the Italian Journal of Mammalogy Abbreviated Journal Hysterix
Volume 28 Issue 2 Pages (down) 180-185
Keywords
Abstract Wolf choruses ( Canis lupus ) are complex, multicomponent signals, composed by a series of different vocalizations emitted by a pack. Although howls, the main component, have been highly studied, poor attention has been drawn upon the other vocalizations of the chorus. In this study, we investigate the structure of the chorus by means of the analysis and the quantification of the different components, taking advantage both of the digital sound recording and analysis, and of the modern statistical methodologies. We provide for the first time a detailed, objective description of the types of call emitted during the wolf howlings, combining spectrographic examinations, spectral analyses and automated classifications, with the aim to identify different types of call. Our results show that wolf choruses have a rich, complex structure, that reveals six other types of call, to be added to those howls already described in literature. Wolf choruses are typically composed by other three different types of calls: the bark, i.e. relatively long calls characterized by low frequencies and the presence of harsh components (deterministic chaos); the whimper, characterized by a harmonic structure and a very short duration; and the growl, a call with a noisy structure, low frequencies but relative long duration. Although further investigations are necessary to understand the meaning of the different calls, this research provides a basis for those studies that aim to compare wolves and other canids vocal behaviour.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0394-1914 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Passilongo2017 Serial 6441
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Author de Jong, T.R.; Neumann, I.D.
Title Oxytocin and Aggression Type Book Chapter
Year 2018 Publication Behavioral Pharmacology of Neuropeptides: Oxytocin Abbreviated Journal
Volume Issue Pages (down) 175-192
Keywords
Abstract The neuropeptide oxytocin (OT) has a solid reputation as a facilitator of social interactions such as parental and pair bonding, trust, and empathy. The many results supporting a pro-social role of OT have generated the hypothesis that impairments in the endogenous OT system may lead to antisocial behavior, most notably social withdrawal or pathological aggression. If this is indeed the case, administration of exogenous OT could be the “serenic” treatment that psychiatrists have for decades been searching for.
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Corporate Author Thesis
Publisher Springer International Publishing Place of Publication Cham Editor Hurlemann, R.; Grinevich, V.
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN 978-3-319-63739-6 Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ de Jong2018 Serial 6424
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Author Steinhoff-Wagner, J.
Title Coat Clipping of Horses: A Survey Type Journal Article
Year 2019 Publication Journal of Applied Animal Welfare Science Abbreviated Journal Journal of Applied Animal Welfare Science
Volume 22 Issue 2 Pages (down) 171-187
Keywords Equine, thermoregulation, shaving, winter pelage removal
Abstract Coat clipping is a common practice in sport horses; however, timing, purpose, technique, and clips vary widely, as do the management and feeding of a clipped horse. The aim of this study was to collect data regarding common clipping practices. A questionnaire was published online in Germany and contained 32 questions. Four hundred ninety-eight people answered at least one question, and 373 individuals (7% male, 93% female; ages 14–59 years) completed all the questions. Clipped horses were predominantly used as sport horses (68%), and they were either clipped immediately before or during the winter season (88%) or year-round (7%). The clipping date was scheduled according to hair length (52%), sweat amount (47%), and drying time (47%). Participants primarily used two clips: the hunter clip and the blanket clip, both without clipping the head (23% each). The majority of the clipped horses wore a blanket day and night (> 90%). Future studies with observations in the field are needed to support survey data in an effort to develop welfare recommendations for clipping practices utilized with horses.
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Corporate Author Thesis
Publisher Routledge Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 1088-8705 ISBN Medium
Area Expedition Conference
Notes doi: 10.1080/10888705.2018.1454319 Approved no
Call Number Equine Behaviour @ team @ Serial 6613
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Author Boissy, A.
Title Fear and Fearfulness in Animals Type Journal Article
Year 1995 Publication The Quarterly Review of Biology Abbreviated Journal The Quarterly Review of Biology
Volume 70 Issue 2 Pages (down) 165-191
Keywords
Abstract Persistence of individual differences in animal behavior in reactions to various environmental challenges could reflect basic divergences in temperament, which might be used to predict details of adaptive response. Although studies have been carried out on fear and anxiety in various species, including laboratory, domestic and wild animals, no consistent definition of fearfulness as a basic trait of temperament has emerged. After a classification of the events that may produce a state of fear, this article describes the great variability in behavior and in physiological patterns generally associated with emotional reactivity. The difficulties of proposing fearfulness-the general capacity to react to a variety of potentially threatening situations-as a valid basic internal variable are then discussed. Although there are many studies showing covariation among the psychobiological responses to different environmental challenges, other studies find no such correlations and raise doubts about the interpretation of fearfulness as a basic personality trait. After a critical assessment of methodologies used in fear and anxiety studies, it is suggested that discrepancies among results are mainly due to the modulation of emotional responses in animals, which depend on numerous genetic and epigenetic factors. It is difficult to compare results obtained by different methods from animals reared under various conditions and with different genetic origins. The concept of fearfulness as an inner trait is best supported by two kinds of investigations. First, an experimental approach combining ethology and experimental psychology produces undeniable indicators of emotional reactivity. Second, genetic lines selected for psychobiological traits prove useful in establishing between behavioral and neuroendocrine aspects of emotional reactivity. It is suggested that fearfulness could be considered a basic feature of the temperament of each individual, one that predisposes it to respond similarly to a variety of potentially alarming challenges, but is nevertheless continually modulated during development by the interaction of genetic traits of reactivity with environmental factors, particularly in the juvenile period. Such interaction may explain much of the interindividual variability observed in adaptive responses.
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Corporate Author Thesis
Publisher The University of Chicago Press Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0033-5770 ISBN Medium
Area Expedition Conference
Notes doi: 10.1086/418981 Approved no
Call Number Equine Behaviour @ team @ Serial 6664
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Author Carson, K.; Wood-Gush, D.G.M.
Title Equine behaviour: I. A review of the literature on social and dam--Foal behaviour Type Journal Article
Year 1983 Publication Applied Animal Ethology Abbreviated Journal
Volume 10 Issue 3 Pages (down) 165-178
Keywords
Abstract In most cases, the social organisation of each of the seven species of Equidae existing today outside captivity is either territorial or non-territorial. The striking differences found between these two types of organisation in the social grouping and bonds, mating behaviour, leadership and dominance hierarchies of the animals are examined. It is thought that the non-territorial species show a less primitive type of organisation than the territorial animals. Infant Equidae are precocious animals and are able to follow their dams soon after birth. They stay close by their dams and travel with the herd from an early age and are therefore classified as “followers”, in contrast to the species which have a period of hiding after birth. Dams recognise their foals immediately after birth, whereas it takes 2 or 3 days for a foal to form an attachment to its dam. Being in close proximity to their dams, foals are able to nurse frequently and, unless artificially weaned, a foal will nurse until its dam foals again. Foals start to graze during their first week and as they grow older they spend more time grazing and less time nursing and resting. It is normal for foals to be corprophagic until one month old, and this provides them with bacteria essential for the digestion of fibre. Play behaviour is solitary in very young foals, but after 4 weeks of age, foals play together, with male foals playing more than females and showing more aggressive, fighting movements in play.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0304-3762 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6671
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Author Passilongo, D.; Buccianti, A.; Dessi-Fulgheri, F.; Gazzola, A.; Zaccaronii, M.; Apollonio, M.
Title The Acoustic Structure Of Wolf Howls In Some Eastern Tuscany (Central Italy) Free Ranging Packs Type Journal Article
Year 2010 Publication Bioacoustics Abbreviated Journal Bioacoustics
Volume 19 Issue 3 Pages (down) 159-175
Keywords Canis lupus, acoustic structure, mammal communication, sonogram, fundamental frequency.
Abstract Italian wolf howls are described for the first time from observations between 2003–2008 of a population living in eastern Tuscany, central Italy. A sample of 37 howls selected among single responses and 128 howls included in the choruses of 7 free ranging packs was recorded and analysed. The mean fundamental frequency of the howls ranged between 274–908 Hz. Two main structures recognised by means of multivariate explorative analysis, in particular Principal Component and Cluster Analysis, were ascribed to breaking and flat howls. Discriminant Function Analysis was applied to the recognised groups with the aim to find a general rule for classification. Howls with different features were correctly assigned to the groups obtained by explorative analysis in 95.8% of cases. The analysis of the variables characterising the structure of the howls suggests that maximum frequency and range of fundamental frequency are the most important parameters for classification, while duration does not appear to play any significant role.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6499
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Author Murphy, J.; Arkins, S.
Title Laterality and visuo-spatial ability in the equine: Functional measures of sport horse selection? Type Journal Article
Year 2006 Publication BSAP Occasional Publication Abbreviated Journal BSAP Occasional Publication
Volume 35 Issue Pages (down) 159-170
Keywords
Abstract Laterality in any organism or species can be manifest as morphological, sensory and functional degrees of asymmetry such as hemispheric dominance, handedness or sidedness and other motor functional behaviours and as such is equally important in equitation. The influence of the horses' sex on both the direction and the degree of the laterality was explored within and between 4 experimental procedures in the 1st study. The findings showed that the direction, but not the degree of idiosyncratic motor preference in the horses was strongly sex-related. Male horses exhibited significantly more left lateralized responses and female horses exhibited significantly more right lateralized responses. Visuo-spatial ability is also likely to be important in the performance horse. In many species, moderate to large differences in visuo-spatial ability have been reported between the sexes, with superior visuo-spatial ability being reported in males of all species investigated to date. As no known studies had addressed visuo-spatial ability in the equine, the objective of the 2nd study, was to determine if visuo-spatial ability differed between male and female horses. The results produced the first behavioural demonstration of superior visuo-spatial ability in male horses, similar to that reported in other species. There is evidence to suggest that visuospatial ability and motor laterality are associated with cerebral hemispheric asymmetry and may be intrinsically linked. Brain development and laterality have also been associated with hair patterning, and, in a 3rd study we attempted to identify predictors of lateral bias in motor behaviour in horses. We investigated the relationship between the direction of facial hair whorl rotation and the incidence/direction of laterality in the horse. The findings suggest that direction of facial hair whorl rotation may be a useful indicator of lateralised motor behavioural preferences in the horse. We then attempted to establish if laterality was evident at birth in a 4th study, where we explored if neonatal foals exhibited lateralised patterns during and immediately post the birthing process that were correlated with their facial hair whorl patterns. The results showed a significant association between the sex of the foal and the choice of foreleg presented initially during 2nd stage parturition. Significantly more colt foals led with the left foreleg and significantly more filly foals led with the right foreleg than expected purely by random and the behaviour was correlated with facial hair whorl patterns. The findings also suggest that lateralisation in the horse is determined in utero as has also been shown in humans. Comparisons of wholly intact male and female horses are warranted as they might elucidate additional linkages between motor behaviour, visuo-spatial ability and brain organisation and development in the horse. Further research in this area could lead to more appropriate competition conditions (better fence design/construction on cross-country tracks) and so eliminate unnecessary levels of risk associated with many equestrian sports.
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Corporate Author Thesis
Publisher Cambridge University Press Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition 2018/02/27
ISSN 0263-967x ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6512
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