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| Author |
Lee, J.; Floyd, T.; Erb, H.; Houpt, K. |
| Title |
Preference and demand for exercise in stabled horses |
Type |
Journal Article |
| Year |
2011 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
130 |
Issue |
3-4 |
Pages  |
91-100 |
| Keywords |
Horse; Welfare; Exercise; Operant conditioning; Two choice preference; Treadmill |
| Abstract |
Operant conditioning and two choice preference tests were used to assess the motivation of horses to be released from straight and from box stalls. The motivations for food, a companion, and release into a paddock were compared when the horses had to work for each commodity at increasing fixed ratios of responses (panel presses) to reward in an equine operant conditioning stall. The motivation for food (mean ± SEM = 258 ± 143) responses was much greater than that for either release (38 ± 32) from a straight stall into a large paddock alone or into a small paddock with another horse (95 ± 41) (P = 0.04). When given a two choice preference test between exercise on a treadmill for 20 min or returning to their box stalls, eight of nine horses chose to return to their stalls. In a two choice preference test six of eight horses in box stalls chose to be released into a paddock alone. Horses were given a series of two choice preference tests to determine how long they preferred to be in a paddock. After 15 min in the paddock the horses were re-tested, but all chose the paddock when released into a paddock with three other horses. They were retested every 15 min until they chose to return to their stalls. They chose to stay out for 35 ± 6 min when other horses were in the paddock but for only 17 ± 2 min when they would be alone. When deprived of stall release for 48 h the horses chose to remain in the paddock with other horses for 54 ± 6 min, but showed no compensatory behavior when they were alone (duration chosen = 16 ± 4 min). These findings indicate that horses are not strongly motivated to exercise alone and will choose not to endure forced exercise on a treadmill. The social context of voluntary exercise is important; horses are willing to stay out of their stalls longer if other horses are present and will show compensatory behavior only if other horses are present. These finding have implications for optimizing turnout time for stalled horses. |
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0168-1591 |
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Equine Behaviour @ team @ |
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5330 |
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| Author |
McLean, A.N. |
| Title |
Short-term spatial memory in the domestic horse |
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Journal Article |
| Year |
2004 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
85 |
Issue |
1-2 |
Pages |
93-105 |
| Keywords |
Animal cognition; Delayed response; Equine memory; Horse; Object permanence; Temporal cognition |
| Abstract |
This study investigates the ability of horses to recall a feeding event in a two-point choice apparatus. Twelve horses were individually tested whereby they were maintained immobile in a test arena and visually and aurally experienced the delivery of food into one of two feed goals. The horses were then released to make their choice in two experimental contexts: immediate release after experiencing the delivery of food, and release 10 s after food delivery. Each horse performed 40 immediate-release (IR) trials, followed by forty 10-s release trials over a 3-day period. In addition, the same horses were tested 3 months later in the spring with the same number and sequence of trials. Results were analysed by log-linear analysis of frequencies. Results showed that while horses were able to achieve the correct feed goal choice in the immediate-release trials, they were unsuccessful with the 10-s release trials. This suggests that there are limitations in recall abilities in horses, in that they may not possess a prospective type of memory. There are welfare and training implications in these findings concerning the effects of overestimating the mental abilities of horses during training and the effects of delays in reinforcements. |
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refbase @ user @ |
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403 |
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Smith-Funk, E.D.; Crowell-Davis, S.L. |
| Title |
Maternal behavior of draft mares (Equus caballus) with mule foals (Equus asinus x Equus caballus) |
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Journal Article |
| Year |
1992 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
33 |
Issue |
2-3 |
Pages |
93-119 |
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Draft mares and their mule foals were observed from the day of birth to Week 17 of each foal's life. The rate of nursing was recorded and the duration of nursing activity to the nearest second. The rate at which foals engaged in nursing activity varied at each age. The duration of nursing bouts varied slightly as the foals matured. Aggression was recorded during both nursing and non-nursing activity for both the mares and foals. Maternal aggression was highest during nursing activity, especially during the pre-nurse nuzzling period. Maternal aggression increased as the foals matured. Mother-directed foal aggression was primarily in response to maternal aggression. Spatial relationships between each focal dyad were recorded when the foals were upright, not nursing and when they were recumbent. Spatial relationships differed based on the foal's state. The activity in which the mare engaged while her foal was recumbent was recorded. The movements of the mares were also recorded during foal recumbency. Mares approached or maintained their distance from their recumbent foal more than they left their recumbent foal in all weeks of the study, except Week 2. |
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Equine Behaviour @ team @ |
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2272 |
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Lamoot, I.; Callebaut, J.; Demeulenaere, E.; Vandenberghe, C.; Hoffmann, M. |
| Title |
Foraging behaviour of donkeys grazing in a coastal dune area in temperate climate conditions |
Type |
Journal Article |
| Year |
2005 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
92 |
Issue |
1-2 |
Pages |
93-112 |
| Keywords |
Grazing behaviour; Habitat use; Donkey; Equid; Diet composition |
| Abstract |
A small herd of donkeys was introduced in a coastal dune reserve `Houtsaegerduinen' (ca. 80 ha) in Belgium, in order to slow down expansion of dominant grass and shrub species. The Houtsaegerduinen is a nutrient poor scrub-dominated dune system with a spatially heterogeneous vegetation pattern. Different aspects of the grazing behaviour (grazing time, bite rate, habitat use, diet composition) of the free-ranging donkeys are described and analysed. Behavioural data (of maximum six adult mares) were collected through continuous focal animal observation in three consecutive years (1998-2001). Temporal variation in grazing time, habitat use and diet composition was determined. During daylight, donkeys spent most of their time on grazing (56%). In all 3 years, grazing time was significantly shorter in summer (45% of their time), longest grazing times were achieved in spring (64%). In spring, the donkeys also achieved the highest bite rate (21.5 bites/min). The grassy habitat was preferred for foraging in all seasons, while the use of scrub and woodland was variable over time. Averaged over the four seasons, the general diet consisted for 80% of graminoids, 10% of forbs and 10% of woody plants. However, diet composition varied not only among seasons and years, but depended also on the foraged habitat type. We discuss the possible role of the donkeys in nature management. |
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Equine Behaviour @ team @ |
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2338 |
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| Author |
Jørgensen, G.H.M.; Borsheim, L.; Mejdell, C.M.; Søndergaard, E.; Bøe, K.E. |
| Title |
Grouping horses according to gender--Effects on aggression, spacing and injuries |
Type |
Journal Article |
| Year |
2009 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
120 |
Issue |
1-2 |
Pages |
94-99 |
| Keywords |
Horse; Group housing; Gender; Injuries; Social behaviour |
| Abstract |
Many horse owners tend to group horses according to gender, in an attempt to reduce aggressive interactions and the risk of injuries. The aim of our experiment was to test the effects of such gender separation on injuries, social interactions and individual distance in domestic horses. A total of 66 horses were recruited from 4 different farms in Norway and Denmark and divided into six batches. Within each batch, horses were allotted into one mare group, one gelding group and one mixed gender group, with most groups consisting of three or four animals. After 4-6 weeks of acclimatisation, a trained observer recorded all social interactions using direct, continuous observation 1 h in the morning and 1 h in the afternoon for three consecutive days. Recordings of the nearest neighbour of each horse were performed using instantaneous sampling every 10 min. The horses were inspected for injuries before grouping, day 1 after grouping and after 4-6 weeks. No significant effect of gender composition was found on social interactions (P > 0.05), spacing (P > 0.07) or injuries (P > 0.23). Eighty percent of all aggressive interactions recorded were threats, not involving physical contact. Horses with the smallest space allowance showed the highest mean number of aggressive interactions (28.6 ± 6.1 interactions per 6 h) compared to the mean of all the other batches (8.3 ± 1.0 interactions per 6 h). Very few injuries were found and most were superficial. In conclusion, gender composition does not seem to have any effect on aggression level, spacing or injuries. However, the early social experience of horses, management of feeding and space allowance probably represents more important factors for successful group housing of domestic horses. |
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0168-1591 |
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Equine Behaviour @ team @ |
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5093 |
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van Dierendonck, M.C.; Bandi, N.; Batdorj, D.; Dugerlham, S.; Munkhtsog, B. |
| Title |
Behavioural observations of reintroduced Takhi or Przewalski horses (Equus ferus przewalskii) in Mongolia |
Type |
Journal Article |
| Year |
1996 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
50 |
Issue |
2 |
Pages |
95-114 |
| Keywords |
Horse; Przewalski's horse; Reintroduction; Time budget; Social organisation; Basic reference dataset |
| Abstract |
During 1992 and 1993, 14 reintroduced Przewalski Horses or Takhi (Equus ferus przewalskii) were studied in the Hustain Nuruu Mountain Steppe reserve in Mongolia. Most of the individuals did not know each other before reintroduction. These Takhi were the first of five groups due to be released in the reserve after an acclimatisation period of at least 1 year. During acclimatisation the Takhi, lived visually and acoustically separately, in fenced enclosures of approximately 45 ha each. The observations, mostly scan-sampling, were carried out in each season. The observation bouts were divided over six periods and over two harem herds. Two of the periods were in the same consecutive seasons, so comparison over the years was possible. Social integration within the Takhi herds was very high from the beginning, as described by the spatial relation and synchronisation data. Between 50 and 89% of the observation time, the behaviour of all herd members was synchronised. The amount of time spent grazing by the Takhi (30-68% of the daylight period) was similar to that of feral horses and Takhi in captivity and semi-reserves. The Takhi tended to rest in the morning and have a bimodal period of grazing at dawn and in the afternoon. The Takhi displayed clear habitat preferences for certain activities. They had a strong preference to rest at the highest point in their enclosure. They fed preferably on two or three different vegetation types (with five types available in each enclosure). The amount of time spent grazing during the non-growing seasons (49 +/- 15%) indicates that the feeding value and availability of food were sufficient. Health changes were detected adequately using condition scoring sheets. No supplementary food or water was supplied during the harsh winters. Moreover, low mortality rates and high reproductive success show that the mountain steppe is a habitat which is potentially suitable for establishing a healthy Takhi population. Takhi is the first species to return to its native habitat after living only in zoos for so many generations. |
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762 |
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Parker, G.A.; MacNair, M.R. |
| Title |
Models of parent-offspring conflict. I. Monogamy |
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Journal Article |
| Year |
1978 |
Publication |
Animal Behaviour. |
Abbreviated Journal |
Anim. Behav. |
| Volume |
26 |
Issue |
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97-110 |
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Theoretical models for Trivers (1974) concept of parent-offspring conflict are examined for species in which the effects of the conflict are felt by full sibs. A rare conflictor gene will spread if Image , whereÆ’(m) is the fitness gained by a conflictor relative to a non-conflictor offspring (Æ’(m) >1), and m is the amount of parental investment taken by a conflictor relative to m = 1 for a non-conflictor. The range of m alleles which can spread against the parent optimum decreases as the cost to the parent increases until a point is reached where there is no conflict of evolutionary interests. There would be no polymorphism for conflictor: non-conflictor alleles unless special conditions prevail. The conflictor allele which spreads most rapidly as a rare mutant against the parental optimum is not an evolutionarily stable strategy (ESS). The ESS for parent-offspring conflict in monogamous species has m0 = Æ’(m0)/2[dÆ’(m0)/dm0]. The analytical solutions are confirmed throughout by simulations. |
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10.1016/0003-3472(78)90009-X |
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Equine Behaviour @ team @ |
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4901 |
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Pluhácek, J.; Bartos, L.; Bartosová, J.; Kotrba, R. |
| Title |
Feeding behaviour affects nursing behaviour in captive plains zebra (Equus burchellii) |
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Journal Article |
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2010 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
| Volume |
128 |
Issue |
1-4 |
Pages |
97-102 |
| Keywords |
Equus burchellii; Suckling; Nursing; Equid; Feeding; Zoo |
| Abstract |
Equids spend more than half of the day feeding. Lactation is a very demanding form of maternal investment. In an environment such as a zoo, where no grass but several feeding sites are present, conflict between suckling behaviour of the foal and feeding behaviour of the lactating mother should occur. We observed 20 foals of captive plains zebra, Equus burchellii, at the Dvur Králové Zoo, Czech Republic, and collected data concerning suckling events during 17 months of observation. First, we examined whether feeding by the mother while nursing affected suckling behaviour. We found that when the mother was feeding, the proportion of suckling bouts she terminated decreased with increasing age of the foal, whereas it did not change when she was not feeding. This result supported the trade-off between suckling and feeding behaviour which has been reported in other ungulates. Second, we examined what affected interruptions of feeding behaviour of the mother during the suckling bout. The proportion of interruptions of feeding by the mother during nursing increased with increasing age of her foal. This coincides with declining time spent nursing. In addition, younger mothers interrupted their feeding behaviour during suckling bouts more often than older ones. Mothers interrupted feeding during the suckling bout more often when they nursed a daughter than when they nursed a son. The results of our study show that feeding while suckling could reduce parent-offspring conflict and improve welfare of captive foals and mares. |
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Equine Behaviour @ team @ |
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5280 |
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Klimov, V.V. |
| Title |
Spatial-ethological organization of the herd of Przewalski horses (Equus przewalskii) in Askania-Nova |
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Journal Article |
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1988 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
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21 |
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1-2 |
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99-115 |
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The ethological structure of the herd of Przewalski horses includes hierarchic ranks of horses which determine their social roles in the herd. Besides the age ranks, the wild horses are characterized by the formation of harem groups, a “leading” group of females, a group of bachelor stallions, family groups, etc. The ethological structure determines the spatial one, which is the form of distribution of horses over the territory, and its assimilation and transformation into a system of informative spatial units. Under the influence of “internal” and “external” stimuli, the intragoup regulatory mechanims (social adaptations) manifest themselves, which allow the herd to function in the complicated situation of the reserve and allow humans to control the herd by using these mechanisms. There are grounds to believe that, given the balanced ethological structure of these groups, wild horses could be successfully acclimatized into natural biotopes. |
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Equine Behaviour @ team @ |
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2330 |
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Uher, J.; Asendorpf, J.B.; Call, J. |
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Personality in the behaviour of great apes: temporal stability, cross-situational consistency and coherence in response |
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Journal Article |
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2008 |
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Animal Behaviour. |
Abbreviated Journal |
Anim. Behav. |
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75 |
Issue |
1 |
Pages |
99-112 |
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behaviour prediction; bonobo; bottom-up approach; chimpanzee; gorilla; individual differences; orang-utan; personality; traits |
| Abstract |
Using a multidisciplinary approach, the present study complements ethological behaviour measurements with basic theoretical concepts, methods and approaches of the personality psychological trait paradigm. Its adoptability and usefulness for animal studies are tested exemplarily on a sample of 20 zoo-housed great apes (five of each of the following species): bonobos, Pan paniscus; chimpanzees, Pan troglodytes verus; gorillas, Gorilla gorilla gorilla; and orang-utans, Pongo pygmaeus abelii. Data on 76 single trait-relevant behaviours were recorded in a series of 14 laboratory-based situations and in two different group situations. Data collection was repeated completely after a break of 2 weeks within a 60-day period. All behaviour records were sufficiently reliable. Individual- and variable-oriented analyses showed high/substantial temporal stability on different levels of aggregation. Distinctive and stable individual situational and response profiles clarified the importance of situations and of multiple trait-relevant behaviours. The present study calls for a closer collaboration between behavioural biologists and personality psychologists to tap the full potential of animal personality research. |
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Equine Behaviour @ team @ |
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4278 |
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