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Author Boyd, L.E.
Title The behaviour of Przewalski's horses and its importance to their management Type Journal Article
Year 1991 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 29 Issue 1-4 Pages 301-318
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Abstract Przewalski's horses (Equus przewalskii) are believed to be extinct in the wild; the current known population of 797 animals exists wholly in zoos. The Species Survival Commission of the International Union for Conservation of Nature and Natural Resources is proposing to reintroduce this endangered species into its former Mongogian habitat within the next decade. Knowledge of the behavior of harem-forming equids in general and of Przewalski's horses in particular, is of great importance to the captive propagation and eventual reintroduction of this species. Horses are rarely solitary by nature. Solitary captive animals are prone to pacing. Juvenile male feral horses (Equus caballus) form bachelor herds upon dispersal from their natal band. Zoos can set up bachelor herds as a way of managing surplus males. The older, more dominant feral horse bachelors are the first to acquire mares. Bachelors do not generally obtain females until they are 4 or 5 years of age. The first females acquired are usually 1- and 2-year-old fillies dispersing from their natal band. Because of the age differential, the stallions are generally dominant to their mares. Behavioral impotence may result if captive stallions are given a harem at too young an age, especially if the harem contains older, more dominant, females. Typical harem sizes in the wild are 3-5 mares. Captive stallions with too large a harem may become either apathetic or aggressive toward their mares. Wild horses spend 60-70% of their time foraging. Captive animals may quickly consume their limited amounts of food and develop vices out of boredom. Provision of hay ad libitum reduces the amount of pacing seen in captive animals, and virtually eliminates coprophagy.
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Call Number Equine Behaviour @ team @ Serial 2214
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Author McLean, I.G.; Schmitt, N.T.; Jarman, P.J.; Duncan, C.; Wynne, C.D.L.
Title Learning For Life: Training Marsupials To Recognise Introduced Predators Type Journal Article
Year 2000 Publication Behaviour Abbreviated Journal Behaviour
Volume 137 Issue 10 Pages 1361-1376
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Abstract Raising endangered species in captivity for reintroduction necessarily results in animals that lack appropriate skills for coping with problems to be faced in the wild, such as predators. Using classical conditioning techniques involving linking fear of a live dog with the image of a fox, we demonstrate an adjusted fear response for two wallaby species (rufous bettongs Aepyprymnus rufescens, quokkas Setonix brachyurus). No differences in response to the fox were found for wild-caught and captive-born bettongs, even though wild-caught subjects were likely to have encountered canids prior to capture. Attempts to condition a fear response by quokkas to an odour were unsuccessful. An attempt to induce fear of the stuffed fox by linking to fear of humans in quokkas was unsuccessful, but quokkas generalised from fear of the dog to fear of the fox, despite a delay of several weeks. Trained dogs offer a valuable and ethically acceptable mechanism for improving the ability of captive-reared (or sequestered) animals to recognise and cope with predators.
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Call Number Equine Behaviour @ team @ Serial 2282
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Author Feh, C,
Title Long-term paternity data in relation to different aspects of rank for Camargue stallions, Equus caballus Type Journal Article
Year 1990 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume 40 Issue 5 Pages 995-996
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Notes from Professor Hans Klingels Equine Reference List Approved yes
Call Number Serial 1081
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Author HAFEZ, E.S.E.; WILLIAMS, M.; WIERZBOWSKI, S.
Title The Behaviour of Horses. Type Book Whole
Year 1962 Publication In: Hafez. ESE: The Behaviour of Domestic Animals London Abbreviated Journal
Volume Issue Pages 370-396
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Notes from Professor Hans Klingels Equine Reference List Approved no
Call Number Serial 1156
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Author Houpt, K.A.; Fraser, A.F.
Title Przewalski horses Type Journal Article
Year 1988 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 21 Issue 1-2 Pages 1-3
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Call Number Equine Behaviour @ team @ Serial 2302
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Author Klimov, V.V.
Title Spatial-ethological organization of the herd of Przewalski horses (Equus przewalskii) in Askania-Nova Type Journal Article
Year 1988 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 21 Issue 1-2 Pages 99-115
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Abstract The ethological structure of the herd of Przewalski horses includes hierarchic ranks of horses which determine their social roles in the herd. Besides the age ranks, the wild horses are characterized by the formation of harem groups, a “leading” group of females, a group of bachelor stallions, family groups, etc. The ethological structure determines the spatial one, which is the form of distribution of horses over the territory, and its assimilation and transformation into a system of informative spatial units. Under the influence of “internal” and “external” stimuli, the intragoup regulatory mechanims (social adaptations) manifest themselves, which allow the herd to function in the complicated situation of the reserve and allow humans to control the herd by using these mechanisms. There are grounds to believe that, given the balanced ethological structure of these groups, wild horses could be successfully acclimatized into natural biotopes.
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Call Number Equine Behaviour @ team @ Serial 2330
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Author Kolter, L.; Zimmermann, W.
Title Social behaviour of Przewalski horses (Equus p. przewalskii) in the Cologne Zoo and its consequences for management and housing Type Journal Article
Year 1988 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 21 Issue 1-2 Pages 117-145
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Abstract Between 1977 and 1986, two actual rank changes and two unsuccessful attempts occurred among the mares of the Cologne herd. The stallion was at first a low-ranking individual, but attained a dominant position during the last 3 years. At this time he started to split his group and thus to affect attachment relationships among his mares. During his absence of half a year, new bonds resulted and disappeared again some months after his return. Foals were tolerated by the sire for a long time. His behaviour to young, sub-adult mares varied with the individual. Protection of sub-adult mares by adult mares against the stallion's attacks may occur. Young mares protect and guard strange foals from their very first day. Management steps to cope with social and feeding problems consisted of enlargement and adding complexity to the enclosure, the establishment of more feeding sites, building a stable and temporary removal of the stallion.
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Call Number Equine Behaviour @ team @ Serial 2332
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Author Penzhorn,B.L.; Novellie, P.A.
Title Some behavioural traits of Cape mountain zebras and their implications for the management of asmall conservation area Type Journal Article
Year 1991 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 29 Issue 1-4 Pages 293-299
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Abstract The social organisation of mountain zebras (Equus zebra zebra) consists of breeding herds (1 male, 2.4 females (range 1–5) and their offspring) which remain stable over many years, and bachelor groups. Foals leave their maternal herds of their own accord. In a free-ranging population the behaviour of the foals in leaving the herd is probably an adequate mechanism to prevent inbreeding, but inbreeding may occur in confined populations. Individual recognition by means of stripe pattern allows a check to be kept.

Seasonal movement of mountain zebras is associated with a relative change in diet quality (as indicated by crude protein contents of preferred food plants and of faeces) between summer and winter habitats. Any conservation area should be large and varied enough to include both summer and winter habitats. Mountain zebras favour taller grass than most antelope species, harvesting their food at 50–150 mm from the ground. The existence of large populations of antelope could, therefore, be detrimental to zebras.
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Notes from Professor Hans Klingels Equine Reference List Approved no
Call Number Serial 1465
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Author Merkies, K.; McKechnie, M.J.; Zakrajsek, E.
Title Behavioural and physiological responses of therapy horses to mentally traumatized humans Type Journal Article
Year 2018 Publication Applied Animal Behaviour Science Abbreviated Journal
Volume Issue Pages
Keywords Equine-assisted therapy; Ptsd; Horse; Behaviour; Cortisol; Heart rate
Abstract The benefits to humans of equine-assisted therapy (EAT) have been well-researched, however few studies have analyzed the effects on the horse. Understanding how differing mental states of humans affect the behaviour and response of the horse can assist in providing optimal outcomes for both horse and human. Four humans clinically diagnosed and under care of a psychotherapist for Post-Traumatic Stress Disorder (PTSD) were matched physically to four neurotypical control humans and individually subjected to each of 17 therapy horses loose in a round pen. A professional acting coach instructed the control humans in replicating the physical movements of their paired PTSD individual. Both horses and humans were equipped with a heart rate (HR) monitor recording HR every 5secs. Saliva samples were collected from each horse 30 min before and 30 min after each trial to analyze cortisol concentrations. Each trial consisted of 5 min of baseline observation of the horse alone in the round pen after which the human entered the round pen for 2 min, followed by an additional 5 min of the horse alone. Behavioural observations indicative of stress in the horse (gait, head height, ear orientation, body orientation, distance from the human, latency of approach to the human, vocalizations, and chewing) were retrospectively collected from video recordings of each trial and analyzed using a repeated measures GLIMMIX with Tukey's multiple comparisons for differences between treatments and time periods. Horses moved slower (p < 0.0001), carried their head lower (p < 0.0001), vocalized less (p < 0.0001), and chewed less (p < 0.0001) when any human was present with them in the round pen. Horse HR increased in the presence of the PTSD humans, even after the PTSD human left the pen (p < 0.0001). Since two of the PTSD/control human pairs were experienced with horses and two were not, a post-hoc analysis showed that horses approached quicker (p < 0.016) and stood closer (p < 0.0082) to humans who were experienced with horses. Horse HR was lower when with inexperienced humans (p < 0.0001) whereas inexperienced human HR was higher (p < 0.0001). Horse salivary cortisol did not differ between exposure to PTSD and control humans (p > 0.32). Overall, behavioural and physiological responses of horses to humans are more pronounced based on human experience with horses than whether the human is diagnosed with a mental disorder. This may be a reflection of a directness of movement associated with humans who are experienced with horses that makes the horse more attentive. It appears that horses respond more to physical cues from the human rather than emotional cues. This knowledge is important in tailoring therapy programs and justifying horse responses when interacting with a patient in a therapy setting.
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Call Number Equine Behaviour @ team @ Serial 6385
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Author Rutberg, A. T.
Title Inter-group transfer in assateague pony mares Type Journal Article
Year 1990 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.
Volume 40 Issue 5 Pages 945-952
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Abstract Between-previous termgroup transfernext term of adult female previous termponies,next term Equus caballus, was investigated for three consecutive summers on previous Assateaguenext Island, Maryland, U.S.A. Both the previous terminternext term-band movements of individual previous termmares and the marenext term turnover rates of one-male “harem” bands were examined. Long-term previous termtransfersnext term occurred at rates ranging from 0·06 to 0·18 per previous termmarenext term per month. previous termMaresnext term with foals transferred more frequently than previous termmaresnext term without foals, but neither female age, pregnancy, nearest-neighbour distances nor dominance rank affected the likelihood of transferring. Band turnover rates were uncorrelated with the average frequency of previous termmare-marenext term aggression within the band, but new previous termmaresnext term entering a band suffered a transient rise in aggression received. Thus, female aggression did not encourage, and may have discouraged, previous terminternext term-band previous termtransfers.next term Older stallions and stallions who had held bands for 2 years or more had significantly larger and more stable bands. Fewer previous termmarenext term turnovers were seen in bands whose stallions tended to face their previous termmares,next term showed a relatively high proportion of time feeding, and showed a relatively low proportion of time involved in aggression with other stallions, although at marginal levels of significance for all three variables. Thus, variability in stallion attributes, and possibly behaviour, probably plays the strongest role in determining previous termmare transfernext term patterns at previous termAssateague.next term
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Notes from Professor Hans Klingels Equine Reference List Approved yes
Call Number Serial 1535
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