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Author |
Hall, C.; Crowell-Davis, S.L.; Warren, R.J. |
Title |
Maternal and developmental behavior of the feral horses of Cumberland Island, Georgia |
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Journal Article |
Year |
1993 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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37 |
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1 |
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85 |
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Equine Behaviour @ team @ |
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2271 |
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Smith-Funk, E.D.; Crowell-Davis, S.L. |
Title |
Maternal behavior of draft mares (Equus caballus) with mule foals (Equus asinus x Equus caballus) |
Type |
Journal Article |
Year |
1992 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
Volume |
33 |
Issue |
2-3 |
Pages |
93-119 |
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Draft mares and their mule foals were observed from the day of birth to Week 17 of each foal's life. The rate of nursing was recorded and the duration of nursing activity to the nearest second. The rate at which foals engaged in nursing activity varied at each age. The duration of nursing bouts varied slightly as the foals matured. Aggression was recorded during both nursing and non-nursing activity for both the mares and foals. Maternal aggression was highest during nursing activity, especially during the pre-nurse nuzzling period. Maternal aggression increased as the foals matured. Mother-directed foal aggression was primarily in response to maternal aggression. Spatial relationships between each focal dyad were recorded when the foals were upright, not nursing and when they were recumbent. Spatial relationships differed based on the foal's state. The activity in which the mare engaged while her foal was recumbent was recorded. The movements of the mares were also recorded during foal recumbency. Mares approached or maintained their distance from their recumbent foal more than they left their recumbent foal in all weeks of the study, except Week 2. |
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Equine Behaviour @ team @ |
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2272 |
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Crowell-Davis, S.L.; Caudle, A.B. |
Title |
Coprophagy by foals: recognition of maternal feces |
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Journal Article |
Year |
1989 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
Volume |
24 |
Issue |
3 |
Pages |
267-272 |
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Six foals were each observed for 4 h per week during the first 6 weeks of life in an experimental situation in which they had access to feces taken from their mother and from another mare which was not pregnant or lactating. The foals sniffed at the feces equally. Two foals engaged in a total of seven bouts of coprophagy. All bouts of coprophagy involved maternal feces (χ2; P<0.01). |
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Equine Behaviour @ team @ |
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2273 |
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Crowell-Davis, S.L.; Houpt, K.A.; Kane, L. |
Title |
Play development in Welsh pony (Equus caballus) foals |
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Journal Article |
Year |
1987 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
Volume |
18 |
Issue |
2 |
Pages |
119-131 |
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The structure of the play of colts and fillies living on pasture was studied from birth (n = 15) for up to 24 weeks. Foal play was categorized as running and bucking alone, running and bucking in a group, interactive (contact or combat) play, play with an object, and play at an adult. The rate of play decreased with increasing age and ambient temperature. Fillies and colts played with equal frequency, but engaged in some different types of play at different rates. There was no difference between colts and fillies in the proportion of play bouts of running and bucking in a group or playing with an object. Fillies engaged in running and bucking alone more than colts. Colts engaged in interactive play and play at an adult more than fillies. While there was no significant difference between colts and fillies in the duration of either type of running and bucking play, the interactive play bouts of colts were significantly longer than those of fillies. Both mares and stallions were tolerant of foal play which involved use of their body as a play object, including mounting play. Both fillies and colts engaged in mounting play. Foals used various natural objects found in the pasture for repeated bouts of play with inanimate objects, a behaviour which may explain, from a developmental perspective, the occasional use of “tools” in adult equids. The sex differences in type of play were consistent with the social structure of unmanaged adults in which males must compete with each other in order to associate with females. |
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Equine Behaviour @ team @ |
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2274 |
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Author |
Crowell-Davis, S.L. |
Title |
Self-grooming by mares and foals of the Welsh pony (Equus caballus) |
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Journal Article |
Year |
1987 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
Volume |
17 |
Issue |
3-4 |
Pages |
197-208 |
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Self-grooming behaviour of 15 mare-foal pairs was studied weekly for the first 24 weeks of the foal's life. Mares self-groomed at mean rates of 1.2-2.2 times h-1 depending on the 4-week period. Foals self-groomed more often, with a peak rate of 12.3 times h-1 occurring during Week 5-8 and a subsequent decline to 6.0 times h-1 by Weeks 21-24. Self-grooming bouts of mares and foals also differed in the proportion devoted to each type of self-grooming. During a greater proportion of their total self-grooming bouts, foals scratched their head and neck with a hind limb or bit and scratched the trunk or hind limbs with the teeth. The mares spent a greater proportion of their self-grooming bouts rubbing their head or neck on an inanimate object, another pony, or their fore limb, or rolling on the ground. Particular types of individual self-grooming bouts were engaged in for different lengths of time. Foals had longer bouts of scratching the head and neck with a hind limb, scratching or biting the fore limb or hind limb with the teeth, and rubbing the head and neck on an inanimate object, while mares had longer bouts of rolling. Some of the differences in preference for particular types of self-grooming may be the result of differences in relative size of various body parts and, consequently, relative ease of a given type of self-grooming. |
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Equine Behaviour @ team @ |
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2275 |
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Author |
Crowell-Davis, S.L.; Houpt, K.A.; Carini, C.M. |
Title |
Mutual grooming and nearest-neighbor relationships among foals of Equus caballus |
Type |
Journal Article |
Year |
1986 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
Volume |
15 |
Issue |
2 |
Pages |
113-123 |
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A 3-year study was carried out on the developmental behavior of foals from birth to 24 weeks of age and the behavior of mares living with foals. Mutual-grooming partners of foals were primarily other foals. The peak frequency of mutual grooming occurred during Weeks 9-12, when fillies mutual-groomed 1.6 times h-1 and colts mutual-groomed 0.9 times h-1. Fillies mutual-groomed more frequently than colts (P < 0.025). Fillies mutual-groomed randomly with colts and other fillies (P < 0.05), whereas colts mutual-groomed almost exclusively with fillies (P = 0.03). At all ages studied, if a foal's nearest neighbor was not its mother, it was more likely to be another foal than would be expected if the foal was associating randomly with non-mother ponies. Fillies were more likely than expected to have a filly rather than a colt as their nearest neighbor (P = 0.01). Thus, during their first few months of life, the foals studied exhibited patterns of behavior which were consistent with the development of the usual social milieu of unmanaged adults, in which several mares form a cohesive herd with one or more stallions associating with them. |
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Equine Behaviour @ team @ |
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2276 |
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Author |
Kaseda, Y.; K. Nozawa, K. |
Title |
Father-daughter matings and its avoidance in Misaki feral horses |
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Journal Article |
Year |
1996 |
Publication |
Animal Science and Technology |
Abbreviated Journal |
Anim Sci Tech |
Volume |
67 |
Issue |
11 |
Pages |
996-1002 |
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Father-daughter matings and its avoidance mechanism were analysed on the basis of data which gained from behavioural observations and paternity tests in Misaki feral horses from 1979 to 1994. Twelve stallions and their 51 daughters had 176 breeding seasons, but they lived in the different home range in 82 breeding seasons. About half of 1- to 3-year-old mares emigrated from natal area to the other and grew up there. Therefore, emigrations of young mares may result reduction of contacts and avoidance of inbreeding with their fathers. The stallions and their daughters lived in the same area in 94 breeding seasons, but there were no cases that daughters which left their natal harem groups before sexual maturity formed again stable consort relations with their natal harem stallions. It is possible that separation of young mares from their natal groups before sexual maturity may result avoidance of formation of consort relation with their fathers. Two father-daughter matings were observed in 124 paternity tests. These two daughters were born in the other harem groups than their father's and left their natal groups before maturity. After maturity, one of them formed a stable consort relation with her father and the other remained together with her father for 2 months in the breeding season. Both of them had not experience to have lived with their fathers before maturity. The persent result supports the hypothesis in wild and semi-wild horses that inbreedings between fathers and daughters may be avioded by the experience to have lived together before sexual maturity. |
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0918-2365. |
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Equine Behaviour @ team @ |
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2307 |
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Author |
Kaseda, Y.; Khalil, A.M. |
Title |
Harem size and reproductive success of stallions in Misaki feral horses |
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Journal Article |
Year |
1996 |
Publication |
Applied Animal Behaviour Science |
Abbreviated Journal |
Appl. Anim. Behav. Sci. |
Volume |
47 |
Issue |
3-4 |
Pages |
163-173 |
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Harem size; Horse social organization; Reproduction; Blood types; Misaki feral horse |
Abstract |
Over a 16-year period (1979-1994), long-term investigations were carried out on 14 Misaki feral stallions to analyze changes in harem size and the reproductive success. Harem size changed with the age of the stallions. Most stallions formed harem groups with four to five mares at the age of 4-6 and then the number of mares increased rapidly to the maximum at the age of 6-9 years. Thereafter, harem size decreased gradually to a minimum with advancing age. The harem size of 60 stable harem groups ranged from 1 to 9, and the average varied from a minimum mean of 1.8 in 1988 to a maximum mean of 5.3 in 1982. Mean harem size increased as adult sex ratio increased and a significant and positive correlation was found between them. One hundred and ninety-eight sire-foal pairs were determined by a paternity test with blood types and consort relations between stallions and mares during the study period. Out of 99 foals which were born in the stable harem groups, the true sires of 84 foals (85%) were the harem stallions in which the foals were born but the remaining 15 foals (15%) were sired by other harem stallions. Two out of three stallions which were studied throughout their lifetime produced 24 and 25 foals in 10 and 11 years of their reproductive lifespan, respectively. Another one produced only five foals in 6 years. The number of foals sired by the harem stallions was less than two over harem size 7 and some of the foals born in the harem were sired by other harem stallions. These results suggest that if a particular stallion monopolizes too many mares, he could not sire so many offspring because he could not always prevent his rival stallions from mating with his mares in wild or feral circumstances. |
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Equine Behaviour @ team @ |
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2308 |
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Kimura, R. |
Title |
Volatile substances in feces, urine and urine-marked feces of feral horses |
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Journal Article |
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2001 |
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Canadian Journal of Animal Science |
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Can. J. Anim. Sci. |
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81 |
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3 |
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411-420 |
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Odors (volatile), excrement, scent-marking, masking, horse (feral), (releaser) pheromone |
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The identity and amount of volatile substances in the feces, urine and feces scent-marked with urine (i.e., feces mixed with urine) of feral horses was determined by acid/steam distillation and gas chromatography-mass spectrometry. The frequency of excretion and scent marking, as evaluated in the breeding and non-breeding seasons, showed clear evidence of seasonal behavioral differences. The concentration of each substance (fatty acids, alcohols, aldehydes, phenols, amines and alkanes) in the feces differed according to maturity, sex and stage in the reproductive process. They had a characteristic chemical fingerprint. Although the levels of tetradecanoic and hexadecanoic acids in the feces of estrous mares were significantly higher than the respective levels in the feces of non-estrous mares, in the case of scent-marked feces by stallions, the levels of them in the feces from estrous mares had decreased to levels similar to those in non-estrous mares. The concentration of these substances in mares were not significantly different. The presence of a high concentration of cresols in the urine of stallions in the breeding season suggests that one role of scent marking by stallions is masking the odor of the feces produced by mares. |
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0008-3984 |
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Equine Museum of Japan, 1-3 Negishidai, Nakaku, Yokohama 231, Japan (hidousch@alles.or.jp) |
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Equine Behaviour @ team @ |
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2314 |
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Rapaport, L.; King, N.E. |
Title |
The behavioral research program at the Washington Park Zoo |
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Journal Article |
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1987 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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18 |
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1 |
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57-66 |
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For the past decade, the Washington Park Zoo, Portland, Oregon, has had an active behavioral research program. The research department is both a zoo-supported research facility for visiting researchers and staff, and an educational facility that teaches practical behavioral research methods to undergraduates. The research education program utilizes students from any of a dozen local colleges and universities. Students receive academic credit for their participation. Active keeper-participation plays a major role in many research projects. Not only does keeper-cooperation facilitate research, but their knowledge of the individual animals often proves invaluable. In addition to involvement in student projects, keepers have also conducted their own research projects. |
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Equine Behaviour @ team @ |
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2324 |
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