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Author Keiper, R. openurl 
  Title Social organization of feral ponies Type Journal Article
  Year 1976 Publication Abbreviated Journal Proc Pennsyl Acad Sci  
  Volume 50 Issue Pages 69-70  
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  Notes from Professor Hans Klingels Equine Reference List Approved no  
  Call Number (up) Serial 1250  
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Author Keiper, R.R.; Moss, M.; Zervanos, S. openurl 
  Title Daily and seasonal patterns of feral ponies on Assateague Island. Type Conference Article
  Year 1980 Publication 2nd Conference on Scientific Research in the National Parks Abbreviated Journal  
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  Area Expedition Conference  
  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2310  
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Author BASHORE, T. L.; KEIPER, R.; TURNER ,J. W. JR; KIRKPATRICK J. F. url  openurl
  Title The accuracy of fixed-wing aerial surveys of feral horses on a coastal barrier island Type Journal Article
  Year 1990 Publication Journal of coastal research Abbreviated Journal J. coast. res  
  Volume 6 Issue Pages 53-56  
  Keywords Accuracy; Airborne methods; Vegetation; Barrier islands; Maryland; Ground methods; United States; North America; America  
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  Series Editor Series Title Abbreviated Series Title  
  Series Volume Series Issue Edition  
  ISSN 0749-0208 ISBN Medium  
  Area Expedition Conference  
  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2221  
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Author Keiper, R.R. openurl 
  Title The behaviour, ecology and social organization of the feral ponies of Assateague Island Type Journal Article
  Year 1979 Publication Proc. I. Conf. Sci. Res. Nat. Parks Abbreviated Journal Proc. I. Conf. Sci. Res. Nat. Parks,  
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  Publisher University of Wyoming Place of Publication Laramie Editor  
  Language Summary Language Original Title  
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  Area Expedition Conference  
  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2309  
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Author Keiper, R.R.; Keenan, M.A. openurl 
  Title Nocturnal activity patterns of feral horses Type Journal Article
  Year 1980 Publication J. Mammal Abbreviated Journal J. Mammal  
  Volume 61 Issue Pages 116-118  
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  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 2311  
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Author Boyd, L.; Keiper, R. openurl 
  Title Behavioural ecology of feral horses Type Book Chapter
  Year 2005 Publication The domestic horse: the origins, development, and management of its behaviour Abbreviated Journal  
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  Publisher Cambridge University Press Place of Publication Cambridge Editor Mills, D. S.; McDonnell S. M.  
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  Notes Approved no  
  Call Number (up) Equine Behaviour @ team @ Serial 5439  
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Author Kirkpatrick, J.F.; Liu, I.M.; Turner, J.W.J.; Naugle, R.; Keiper, R. openurl 
  Title Long-term effects of porcine zonae pellucidae immunocontraception on ovarian function in feral horses (Equus caballus) Type Journal Article
  Year 1992 Publication Journal of reproduction and fertility Abbreviated Journal J Reprod Fertil  
  Volume 94 Issue 2 Pages 437-444  
  Keywords Animals; Contraception, Immunologic/*veterinary; *Egg Proteins; Estrogens, Conjugated (USP)/urine; Female; Glycoproteins/*pharmacology; Horses/immunology/*physiology; *Membrane Glycoproteins; Ovary/drug effects/*physiology; Progesterone/metabolism; *Receptors, Cell Surface; Swine/immunology; Time Factors; Zona Pellucida/*immunology  
  Abstract Ten feral mares free-roaming in Maryland, USA, were inoculated with porcine zonae pellucidae (PZP) protein before the breeding season for three consecutive years (1988-90). Ovarian function was monitored for 51 days during the peak of the breeding season after the third annual PZP inoculation, in seven of these mares and in four untreated control mares, by means of urinary oestrone conjugates and nonspecific progesterone metabolites. None of the ten inoculated mares became pregnant in 1990, compared with 55% of 20 control mares, which included two of the four monitored for ovarian function. Three of the untreated mares demonstrated apparent normal ovarian activity, characterized by preovulatory oestrogen peaks, concurrent progesterone nadirs at ovulation, breeding activity, and luteal-phase progesterone increases after ovulation. Two of the seven monitored PZP-treated mares demonstrated ovulatory cycles that did not result in conception. One was pregnant as a result of conception in 1989 and demonstrated a normal, late-gestation, endocrine profile. The remaining four PZP-treated mares revealed no evidence of ovulation, and urinary oestrogen concentrations were significantly depressed. The experiments indicated that (i) a third consecutive annual PZP booster inoculation is greater than 90% effective in preventing pregnancies in mares and (ii) three consecutive years of PZP treatment may interfere with normal ovarian function as shown by markedly depressed oestrogen secretion.  
  Address Deaconess Research Institute, Billings, MT 59102  
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  ISSN 0022-4251 ISBN Medium  
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  Notes PMID:1317449 Approved no  
  Call Number (up) refbase @ user @ Serial 145  
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Author Rutberg, A.T.; Keiper, R.R. url  doi
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  Title Proximate causes of natal dispersal in feral ponies: some sex differences Type Journal Article
  Year 1993 Publication Animal Behaviour. Abbreviated Journal Anim. Behav.  
  Volume 46 Issue 5 Pages 969-975  
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  Abstract Abstract. Fifteen years of data on natal dispersal age and the context of dispersal for the feral ponies of Assateague Island, Maryland are presented. Ninety-seven per cent of males and 81% of females dispersed from their natal groups by 5 years of age. For animals that left their natal group, average age of dispersal was 20[middle dot]8 months for males and 24[middle dot]6 months for females. Male dispersal age was strongly and significantly correlated with number of peers in the natal group, and males dispersing with peers were significantly older than males dispersing without peers, suggesting that males delayed dispersal when peers were available for interaction. Female dispersal age was not influenced by number of peers, but was correlated with age of first reproduction. Factors not influencing dispersal age in either sex were presence of a younger sibling, maternal band transfers, and maternal age and dominance rank. The relatively high frequency of females failing to disperse from their natal groups is puzzling in light of data showing diminished fecundity in non-dispersing pony mares.  
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  Notes Approved no  
  Call Number (up) refbase @ user @ Serial 518  
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Author Keiper, R.; Houpt, K. openurl 
  Title Reproduction in feral horses: an eight-year study Type Journal Article
  Year 1984 Publication American journal of veterinary research Abbreviated Journal Am J Vet Res  
  Volume 45 Issue 5 Pages 991-995  
  Keywords Abortion, Veterinary/epidemiology; Anestrus; Animal Population Groups/*physiology; Animals; Animals, Wild/*physiology; Birth Rate; Female; Fertility; Horse Diseases/epidemiology; Horses/*physiology; Lactation; Male; Maryland; Pregnancy; *Reproduction; Seasons; Sex Ratio; Virginia  
  Abstract The reproductive rate and foal survival of the free-ranging ponies on Assateague Island National Seashore were studied for 8 years, 1975 to 1982. Most (52%) of the 86 foals were born in May, 13% were born in April, 22.6% in June, 10.4% in July, and less than 1% in August and September. The mean foaling rate was 57.1 +/- 3.9% and the survival rate was 88.3 +/- 3.6%. Forty-eight colts and 55 fillies were born (sex ratio 53% female). Mares less than 3 years old did not foal and the foaling rate of 3-year-old mares was only 23%, that of 4-year-old mares was 46%, that of 5-year-old mares was 53%, and 6-year-old mares was 69%. The relatively poor reproduction rate was believed to be a consequence of the stress of lactating while carrying a foal when forage quality on the island was low. The hypothesis was supported by the higher reproductive rate (74.4 +/- 2.4%) of the ponies in the Chincoteague National Wildlife Refuge on the southern part of the island. Their foals are weaned and sold in July each year. Despite the low reproductive rate on Assateague Island National Seashore , the number of ponies increased from 43 to 80, a 90% increase in the 8-year period or greater than 10%/yr. There were 24 deaths and 8 dispersals from the study area.  
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  Series Volume Series Issue Edition  
  ISSN 0002-9645 ISBN Medium  
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  Notes PMID:6732036 Approved no  
  Call Number (up) refbase @ user @ Serial 665  
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Author Houpt, K.A.; Keiper, R. openurl 
  Title The position of the stallion in the equine dominance hierarchy of feral and domestic ponies. Type Journal Article
  Year 1982 Publication Journal of Animal Science Abbreviated Journal J. Anim Sci  
  Volume 54 Issue 5 Pages 945-950  
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  Notes Approved no  
  Call Number (up) refbase @ user @ Serial 668  
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