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Author (up) McLean, A.N.; Christensen, J.W.
Title The application of learning theory in horse training Type Journal Article
Year 2017 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 190 Issue Pages 18-27
Keywords Behaviour; Conflict theory; Desensitisation; Habituation; Reinforcement; Stress
Abstract The millennia-old practices of horse training markedly predate and thus were isolated from the mid-twentieth century revelation of animal learning processes. From this standpoint, the progress made in the application and understanding of learning theory in horse training is reviewed including a discussion of how learning processes are employed or otherwise under-utilised in training. This review describes the process of habituation and the most commonly applied desensitisation techniques (systematic desensitisation, counter-conditioning, overshadowing, response prevention) and propose two additional techniques (approach conditioning and stimulus blending). The salience of different types of cues, the interaction of operant and classical conditioning and the impact of stress are also discussed. This paper also exposes the inflexibility and occasional inadequacy of the terminology of learning theory when translated from the research laboratory situation to the practical setting in horse training. While learning theory provides a rich toolbox for riders and trainers, the training process is subject to the simultaneous use of multiple learning processes. In addition, learning/behavioural outcomes and trained responses are not just the result of simple stimulus-response based interactions but are further shaped by arousal, affective and attachment states. More research is needed in these areas. For the field of equitation science to progress and to improve clarity and use of learning processes, changes in nomenclature are required. In particular, the use of the terms 'positive' and 'negative' as descriptive labels in both reinforcement and punishment modalities are unacceptably misleading for everyday use. These labels inhibit the understanding and recognition of the learning processes that these terms supposedly represent, yet the learning processes they describe are vital for horse riders, handlers and trainers to understand. We therefore propose that these labels should be re-labelled more appropriately as 'addition' or 'subtraction' reinforcement/punishment. This would enlighten trainers on the correct application of learning theory, and safety and welfare benefits for people and horses would follow. Finally it is also proposed that the term 'conflict theory' be taken up in equitation science to facilitate diagnosis of training-related behaviour disorders and thus enable the emergence of improved training practices. The optimal use of learning theory should be established as a fundamental principle in equestrian education.
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Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0168-1591 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6597
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Author (up) McVey, A.; Wilkinson, A.; Mills, D.S.
Title Social learning in horses: the effect of using a group leader demonstrator on the performance of familiar conspecifics in a detour task Type Journal Article
Year 2018 Publication Applied Animal Behaviour Science Abbreviated Journal
Volume Issue Pages
Keywords Equine; Imitation; Leader; Social facilitation; Social learning
Abstract Learning through the observation of others allows the transfer of information without the costs incurred during individual trial and error learning. Horses (Equus caballus) are a highly social species, which might be expected to be capable of learning from others, but experimental findings are inconsistent, and potentially confounded by social facilitation effects not related directly to the learning of the task. We refined the methods used in previous equine social learning studies, to examine and distinguish specific social influences on learning of a task: we used predefined group leaders rather than agonistically dominant individuals to demonstrate a detour task to familiar conspecific observers; in addition we had two control groups: a non-observer (true control) and a group with the demonstrator simply present at the goal (social facilitation control). 44 socially kept horses were allocated to one of the three test conditions and took part in five trials each. Success rate, latency and detour direction were recorded. There was no significant difference between the three groups in the likelihood of them succeeding in the task nor latency to succeed; however there was a significant difference in the route chosen by the groups, with the true control choosing the side with the entrance gate significantly more than either the observer group or social facilitation group. Both of the latter two groups chose to go in the same direction relative to themselves, regardless of which side the gate was. Seven out of nine horses in the observer group chose the same direction as their demonstrator every time. Our results show a significant role of social facilitation on detour behaviour and highlight the importance of including adequate controls for simpler cognitive influences on behaviour before claims can be made about the specific learning of motor actions or goal directed behaviour. Social cues may be important to horses if the task is sufficiently challenging and motivationally important, so future work should consider more demanding, but ecologically relevant situations, in order to maximise the potential revelation of social learning effects which do not depend on simple local or stimulus enhancement effects.
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Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0168-1591 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6395
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Author (up) Mech, L.D.
Title The Wolf: The Ecology and Behaviour of an Endangered Species Type Book Whole
Year 1970 Publication Abbreviated Journal
Volume Issue Pages
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Publisher The Natural History Press, Garden City Place of Publication New York Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Mech1970 Serial 6480
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Author (up) Meddock, T.; Osborn, D.
Title Neophobia in wild and laboratory mice Type Journal Article
Year 1968 Publication Psychol Sci Abbreviated Journal
Volume 12 Issue Pages
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Notes Approved no
Call Number Equine Behaviour @ team @ Meddock1968 Serial 6366
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Author (up) Meek, P.D.; Ballard, G.-A.; Fleming, P.J.S.
Title The pitfalls of wildlife camera trapping as a survey tool in Australia Type Journal Article
Year 2015 Publication Australian Mammalogy Abbreviated Journal Aust. Mammal.
Volume 37 Issue 1 Pages 13-22
Keywords camera trap constraints, critical review, remote cameras.
Abstract Camera trapping is a relatively new addition to the wildlife survey repertoire in Australia. Its rapid adoption has been unparalleled in ecological science, but objective evaluation of camera traps and their application has not kept pace. With the aim of motivating practitioners to think more about selection and deployment of camera trap models in relation to research goals, we reviewed Australian camera trapping studies to determine how camera traps have been used and how their technological constraints may have affected reported results and conclusions. In the 54 camera trapping articles published between 1991 and 2013, mammals (86%) were studied more than birds (10%) and reptiles (3%), with small to medium-sized mammals being most studied. Australian camera trapping studies, like those elsewhere, have changed from more qualitative to more complex quantitative investigations. However, we found that camera trap constraints and limitations were rarely acknowledged, and we identified eight key issues requiring consideration and further research. These are: camera model, camera detection system, camera placement and orientation, triggering and recovery, camera trap settings, temperature differentials, species identification and behavioural responses of the animals to the cameras. In particular, alterations to animal behaviour by camera traps potentially have enormous influence on data quality, reliability and interpretation. The key issues were not considered in most Australian camera trap papers and require further study to better understand the factors that influence the analysis and interpretation of camera trap data and improve experimental design.
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Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6704
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Author (up) Mejdell, C.M.; Buvik, T.; Jørgensen, G.H.M.; Bøe, K.E.
Title Horses can learn to use symbols to communicate their preferences Type Journal Article
Year 2016 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 184 Issue Pages 66-73
Keywords Operant conditioning; Blanket; Rug; Thermoregulation; Cognition; Clicker training
Abstract This paper describes a method in which horses learn to communicate by touching different neutral visual symbols, in order to tell the handler whether they want to have a blanket on or not. Horses were trained for 10-15min per day, following a training program comprising ten steps in a strategic order. Reward based operant conditioning was used to teach horses to approach and touch a board, and to understand the meaning of three different symbols. Heat and cold challenges were performed to help learning and to check level of understanding. At certain stages, a learning criterion of correct responses for 8-14 successive trials had to be achieved before proceeding. After introducing the free choice situation, on average at training day 11, the horse could choose between a “no change” symbol and the symbol for either “blanket on” or “blanket off” depending on whether the horse already wore a blanket or not. A cut off point for performance or non-performance was set to day 14, and 23/23 horses successfully learned the task within this limit. Horses of warm-blood type needed fewer training days to reach criterion than cold-bloods (P<0.05). Horses were then tested under differing weather conditions. Results show that choices made, i.e. the symbol touched, was not random but dependent on weather. Horses chose to stay without a blanket in nice weather, and they chose to have a blanket on when the weather was wet, windy and cold (χ2=36.67, P<0.005). This indicates that horses both had an understanding of the consequence of their choice on own thermal comfort, and that they successfully had learned to communicate their preference by using the symbols. The method represents a novel tool for studying preferences in horses.
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0168-1591 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6617
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Author (up) Mejdell, C.M.; Buvik, T.; Jørgensen, G.H.M.; Bøe, K.E.
Title Horses can learn to use symbols to communicate their preferences Type Journal Article
Year 2016 Publication Applied Animal Behaviour Science Abbreviated Journal Appl. Anim. Behav. Sci.
Volume 184 Issue Pages 66-73
Keywords Operant conditioning; Blanket; Rug; Thermoregulation; Cognition; Clicker training
Abstract This paper describes a method in which horses learn to communicate by touching different neutral visual symbols, in order to tell the handler whether they want to have a blanket on or not. Horses were trained for 10-15min per day, following a training program comprising ten steps in a strategic order. Reward based operant conditioning was used to teach horses to approach and touch a board, and to understand the meaning of three different symbols. Heat and cold challenges were performed to help learning and to check level of understanding. At certain stages, a learning criterion of correct responses for 8-14 successive trials had to be achieved before proceeding. After introducing the free choice situation, on average at training day 11, the horse could choose between a “no change” symbol and the symbol for either “blanket on” or “blanket off” depending on whether the horse already wore a blanket or not. A cut off point for performance or non-performance was set to day 14, and 23/23 horses successfully learned the task within this limit. Horses of warm-blood type needed fewer training days to reach criterion than cold-bloods (P<0.05). Horses were then tested under differing weather conditions. Results show that choices made, i.e. the symbol touched, was not random but dependent on weather. Horses chose to stay without a blanket in nice weather, and they chose to have a blanket on when the weather was wet, windy and cold (χ2=36.67, P<0.005). This indicates that horses both had an understanding of the consequence of their choice on own thermal comfort, and that they successfully had learned to communicate their preference by using the symbols. The method represents a novel tool for studying preferences in horses.
Address
Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0168-1591 ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6651
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Author (up) Mench, J.A.; Morrow-Tesch, J.; Chu, L.-R.
Title Environmental enrichment for farm animals Type Journal Article
Year 1998 Publication Lab Animal Abbreviated Journal Lab Anim.
Volume 27 Issue Pages 32-36
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Corporate Author Thesis
Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN ISBN ISSN : 0093-7355 Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6188
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Author (up) Meriggi, A.; Dagradi, V.; Dondina, O.; Perversi, M.; Milanesi, P.; Lombardini, M.; Raviglione, S.; Repossi, A.
Title Short-term responses of wolf feeding habits to changes of wild and domestic ungulate abundance in Northern Italy Type Journal Article
Year 2014 Publication Ethology Ecology & Evolution Abbreviated Journal Ethology Ecology & Evolution
Volume 27 Issue 4 Pages 389-411
Keywords
Abstract
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Corporate Author Thesis
Publisher Taylor & Francis Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
Series Volume Series Issue Edition
ISSN 0394-9370 ISBN Medium
Area Expedition Conference
Notes doi: 10.1080/03949370.2014.986768 Approved no
Call Number Equine Behaviour @ team @ Serial 6688
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Author (up) Meriggi,A.; Lovari, S.
Title A Review of Wolf Predation in Southern Europe: Does the Wolf Prefer Wild Prey to Livestock? Type Journal Article
Year 1996 Publication Journal of Applled Ecology Abbreviated Journal J. Appl. Ecol
Volume 33 Issue Pages 1561-1571
Keywords Canis lupus, conservation, food habits, prey abundance, prey availability.
Abstract 1. The recent recovery of the wolf in southern Europe has not yet removed the risk

of local extinction. Wolf populations are fragmented and often comprise fewer than

500 individuals. In North America, northern and eastern Europe, wolves feed maiiily

on wild herbivores. In southern Europe, this canid has apparently adapted to feed

also on fruit, rubbish, livestock, small and medium-size mammals.

2. The main conservation problem lies with predation o n domestic ~ingulates,w liich

leads to extensive killing of wolves. The reintroduction of wild large herbivores has

been advocated as a means of reducing attacks on livestock, but predatiori on the

latter may remain high if domestic ungulates are locally abundant.

3. Our synthesis of 15 studies, published in the last 15 years, on food habits of the

wolf in southern Europe, has shown that ungulates have been the main diet component

overall. A significant inverse correlation was found between the occurrence (%) of

wild and domestic ungulates in the diet. The presence of relatively few wild ungulate

species was necessary to reduce predation on livestock.

4. Selection of wild and domestic ungulate prey was influenced mainly by their local

abundance, but also by their accessibility. Feeding dependence on rubbish was local

and rare. In Italy, the consumption of riibbish/fruit and that of ungulates was significantly

negatively correlated. Diet breadth increased as the presence of large prey

in tlie diet decreased.

5. The simultaneous reintroduction of severa1 wild ungulate species is likely to reduce

predation on livestock and may prove to be one of the most effective conservation

measures.
Address
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Publisher Place of Publication Editor
Language Summary Language Original Title
Series Editor Series Title Abbreviated Series Title
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ISSN ISBN Medium
Area Expedition Conference
Notes Approved no
Call Number Equine Behaviour @ team @ Serial 6387
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