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	Author	Broom, M.
	Title	A unified model of dominance hierarchy formation and maintenance			Type	Journal Article
	Year	2002	Publication	Journal of theoretical biology	Abbreviated Journal	J. Theor. Biol.
	Volume	219	Issue	1	Pages	63-72
	Keywords	Animals; Behavior, Animal; Feeding Behavior; Models, Psychological; *Social Dominance; Social Environment
	Abstract	In many different species it is common for animals to spend large portions of their lives in groups. Such groups need to divide available resources amongst the individuals they contain and this is often achieved by means of a dominance hierarchy. Sometimes hierarchies are stable over a long period of time and new individuals slot into pre-determined positions, but there are many situations where this is not so and a hierarchy is formed out of a group of individuals meeting for the first time. There are several different models both of the formation of such dominance hierarchies and of already existing hierarchies. These models often treat the two phases as entirely separate, whereas in reality, if there is a genuine formation phase to the hierarchy, behaviour in this phase will be governed by the rewards available, which in turn depends upon how the hierarchy operates once it has been formed. This paper describes a method of unifying models of these two distinct phases, assuming that the hierarchy formed is stable. In particular a framework is introduced which allows a variety of different models of each of the two parts to be used in conjunction with each other, thus enabling a wide range of situations to be modelled. Some examples are given to show how this works in practice.
	Address	Centre for Statistics and Stochastic Modelling, School of Mathematical Sciences, University of Sussex, Falmer, Brighton, BN1 9QH, U.K. m.broom@sussex.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0022-5193	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:12392975			Approved	no
	Call Number	refbase @ user @			Serial	439
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	Author	Heitor, F.; do Mar Oom, M.; Vicente, L.
	Title	Social relationships in a herd of Sorraia horses Part I. Correlates of social dominance and contexts of aggression			Type	Journal Article
	Year	2006	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	73	Issue	2	Pages	170-177
	Keywords	Age Factors; Aggression; Animals; Female; Hierarchy, Social; Horses/psychology; Male; Sex Factors; Social Dominance; *Social Environment; Statistics, Nonparametric
	Abstract	Factors related to dominance rank and the functions of aggression were studied in a herd of Sorraia horses, Equus caballus, under extensive management. Subjects were 10 adult mares 5-18 years old and a stallion introduced into the group for breeding. Dominance relationships among mares were clear, irrespective of rank difference, and remained stable after introduction of the stallion. The dominance hierarchy was significantly linear and rank was positively correlated with age and total aggressiveness. Higher-ranking mares received lower frequency and intensity of agonistic interactions. Nevertheless, higher-ranking dominants were not more likely to elicit submission from their subordinates than lower-ranking dominants. Neither close-ranking mares nor mares with less clear dominance relationships were more aggressive towards each other. Agonistic interactions seemed to be used more importantly in regulation of space than to obtain access to food or to reassert dominance relationships. Contexts of aggression were related to mare rank. The results suggest that dominance relationships based on age as a conventional criterion were established to reduce aggressiveness in a herd where the costs of aggression are likely to outweigh the benefits.
	Address	Centro de Biologia Ambiental, Faculdade de Ciencias da Universidade de Lisboa, Campo Grande, Edificio C2, 1749-016 Lisboa, Portugal
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0376-6357	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16815645			Approved	no
	Call Number	refbase @ user @			Serial	292
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	Author	Gajdon, G.K.; Fijn, N.; Huber, L.
	Title	Limited spread of innovation in a wild parrot, the kea (Nestor notabilis)			Type	Journal Article
	Year	2006	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	9	Issue	3	Pages	173-181
	Keywords	Animal Communication; Animals; Diffusion of Innovation; Feeding Behavior; Female; Imitative Behavior; Learning; Male; Motor Skills; Parrots; *Problem Solving; Sex Factors; Social Dominance; Social Environment; Statistics, Nonparametric
	Abstract	In the local population of kea in Mount Cook Village, New Zealand, some keas open the lids of rubbish bins with their bill to obtain food scraps within. We investigated the extent to which this innovation has spread in the local population, and what factors limit the acquisition of bin opening. Only five males of 36 individually recognised birds were observed to have performed successful bin opening. With one exception there were always other keas present, watching successful bin opening. Seventeen additional individuals were seen to have benefitted from lid opening. Their foraging success was less than that of the bin openers. Social status of bin openers did not differ from scrounging males. Among the individuals that were regularly seen at the site of the bins but were not successful in bin opening, social status and the ratio of feeding directly from open bins correlated with the amount of opening attempts. We conclude that scrounging facilitated certain behavioural aspects of bin opening rather than inhibiting them. The fact that only 9% of opening attempts were successful, and the long period of time required to increase efficiency in lid opening shows that mainly individual experience, and to a lesser extent insight and social learning, play key roles in acquisition of the opening technique. The results indicate that the spread of innovative solutions of challenging mechanical problems in animals may be restricted to only a few individuals.
	Address	Department for Behavior, Neurobiology and Cognition, University of Vienna, 1090, Vienna, Austria. gyula.gajdon@univie.ac.at
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16568276			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2472
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	Author	Huber, L.; Gajdon, G.K.
	Title	Technical intelligence in animals: the kea model			Type	Journal Article
	Year	2006	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	9	Issue	4	Pages	295-305
	Keywords	Animals; Comprehension; Intelligence; Models, Psychological; Observation; Parrots; Problem Solving; Psychological Theory; Social Environment; *Tool Use Behavior
	Abstract	The ability to act on information flexibly is one of the cornerstones of intelligent behavior. As particularly informative example, tool-oriented behavior has been investigated to determine to which extent nonhuman animals understand means-end relations, object affordances, and have specific motor skills. Even planning with foresight, goal-directed problem solving and immediate causal inference have been a focus of research. However, these cognitive abilities may not be restricted to tool-using animals but may be found also in animals that show high levels of curiosity, object exploration and manipulation, and extractive foraging behavior. The kea, a New Zealand parrot, is a particularly good example. We here review findings from laboratory experiments and field observations of keas revealing surprising cognitive capacities in the physical domain. In an experiment with captive keas, the success rate of individuals that were allowed to observe a trained conspecific was significantly higher than that of naive control subjects due to their acquisition of some functional understanding of the task through observation. In a further experiment using the string-pulling task, a well-probed test for means-end comprehension, we found the keas finding an immediate solution that could not be improved upon in nine further trials. We interpreted their performance as insightful in the sense of being sensitive of the relevant functional properties of the task and thereby producing a new adaptive response without trial-and-error learning. Together, these findings contribute to the ongoing debate on the distribution of higher cognitive skills in the animal kingdom by showing high levels of sensorimotor intelligence in animals that do not use tools. In conclusion, we suggest that the 'Technical intelligence hypothesis' (Byrne, Machiavellian intelligence II: extensions and evaluations, pp 289-211, 1997), which has been proposed to explain the origin of the ape/monkey grade-shift in intelligence by a selection pressure upon an increased efficiency in foraging behavior, should be extended, that is, applied to some birds as well.
	Address	Department for Behavior, Neurobiology and Cognition, University of Vienna, Althanstrasse 14, 1090, Vienna, Austria. ludwig.huber@univie.ac.at
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16909237			Approved	no
	Call Number				Serial	2085
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	Author	Cheney, D.L.; Seyfarth, R.M.
	Title	The representation of social relations by monkeys			Type	Journal Article
	Year	1990	Publication	Cognition	Abbreviated Journal	Cognition
	Volume	37	Issue	1-2	Pages	167-196
	Keywords	Animals; Cercopithecus aethiops/psychology; Concept Formation; Dominance-Subordination; Female; Macaca fascicularis/psychology; Male; Social Behavior; *Social Environment
	Abstract	Monkeys recognize the social relations that exist among others in their group. They know who associates with whom, for example, and other animals' relative dominance ranks. In addition, monkeys appear to compare types of social relations and make same/different judgments about them. In captivity, longtailed macaques (Macaca fascicularis) trained to recognize the relation between one adult female and her offspring can identify the same relation among other mother-offspring pairs, and distinguish this relation from bonds between individuals who are related in a different way. In the wild, if a vervet monkey (Cercopithecus aethiops) has seen a fight between a member of its own family and a member of Family X, this increases the likelihood that it will act aggressively toward another member of Family X. Vervets act as if they recognize some similarity between their own close associates and the close associates of others. To make such comparisons the monkeys must have some way of representing the properties of social relationships. We discuss the adaptive value of such representations, the information they contain, their structure, and their limitations.
	Address	Department of Anthropology, University of Pennsylvania, Philadelphia 19104
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0010-0277	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:2269006			Approved	no
	Call Number	refbase @ user @			Serial	702
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	Author	Cheney, D.L.; Seyfarth, R.M.; Silk, J.B.
	Title	The responses of female baboons (Papio cynocephalus ursinus) to anomalous social interactions: evidence for causal reasoning?			Type	Journal Article
	Year	1995	Publication	Journal of comparative psychology (Washington, D.C. : 1983)	Abbreviated Journal	J Comp Psychol
	Volume	109	Issue	2	Pages	134-141
	Keywords	Animals; Attention; Auditory Perception; Awareness; Concept Formation; Dominance-Subordination; Fear; Female; Hierarchy, Social; Papio/psychology; *Social Behavior; Social Environment; Vocalization, Animal
	Abstract	Baboons' (Papio cynocephalus ursinus) understanding of cause-effect relations in the context of social interactions was examined through use of a playback experiment. Under natural conditions, dominant female baboons often grunt to more subordinate mothers when interacting with their infants. Mothers occasionally respond to these grunts by uttering submissive fear barks. Subjects were played causally inconsistent call sequences in which a lower ranking female apparently grunted to a higher ranking female, and the higher ranking female apparently responded with fear barks. As a control, subjects heard a sequence made causally consistent by the inclusion of grunts from a 3rd female that was dominant to both of the others. Subjects responded significantly more strongly to the causally inconsistent sequences, suggesting that they recognized the factors that cause 1 individual to give submissive vocalizations to another.
	Address	Department of Biology, University of Pennsylvania, Philadelphia 19104, USA
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:7758289			Approved	no
	Call Number	refbase @ user @			Serial	348
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	Author	Bugnyar, T.; Heinrich, B.
	Title	Pilfering ravens, Corvus corax, adjust their behaviour to social context and identity of competitors			Type	Journal Article
	Year	2006	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	9	Issue	4	Pages	369-376
	Keywords	Adaptation, Psychological; Animals; Behavior, Animal; Competitive Behavior; Crows/physiology; Deception; Feeding Behavior; Female; Male; Social Behavior; *Social Environment
	Abstract	Like other corvids, food-storing ravens protect their caches from being pilfered by conspecifics by means of aggression and by re-caching. In the wild and in captivity, potential pilferers rarely approach caches until the storers have left the cache vicinity. When storers are experimentally prevented from leaving, pilferers first search at places other than the cache sites. These behaviours raise the possibility that ravens are capable of withholding intentions and providing false information to avoid provoking the storers' aggression for cache protection. Alternatively, birds may refrain from pilfering to avoid conflicts with dominants. Here we examined whether ravens adjust their pilfer tactics according to social context and type of competitors. We allowed birds that had witnessed a conspecific making caches to pilfer those caches either in private, together with the storer, or together with a conspecific bystander that had not created the caches (non-storer) but had seen them being made. Compared to in-private trials, ravens delayed approaching the caches only in the presence of storers. Furthermore, they quickly engaged in searching away from the caches when together with dominant storers but directly approached the caches when together with dominant non-storers. These findings demonstrate that ravens selectively alter their pilfer behaviour with those individuals that are likely to defend the caches (storers) and support the interpretation that they are deceptively manipulating the others' behaviour.
	Address	Department of Biology, University of Vermont, Burlington, VT 05405, USA. thomas.bugnyar@univie.ac.at
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16909235			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2449
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	Author	Nicol, C.J.
	Title	Development, direction, and damage limitation: social learning in domestic fowl			Type	Journal Article
	Year	2004	Publication	Learning & behavior : a Psychonomic Society publication	Abbreviated Journal	Learn Behav
	Volume	32	Issue	1	Pages	72-81
	Keywords	Adaptation, Psychological; Age Factors; Animals; Behavior, Animal; Chickens; Feeding Behavior; Food Preferences; Imitative Behavior; Imprinting (Psychology); Learning; Maternal Behavior; Reinforcement (Psychology); Social Environment; *Social Facilitation
	Abstract	This review highlights two areas of particular interest in the study of social learning in fowl. First, the role of social learning in the development of feeding and foraging behavior in young chicks and older birds is described. The role of the hen as a demonstrator and possible teacher is considered, and the subsequent social influence of brood mates and other companions on food avoidance and food preference learning is discussed. Second, the way in which work on domestic fowl has contributed to an understanding of the importance of directed social learning is examined. The well-characterized hierarchical social organization of small chicken flocks has been used to design studies which demonstrate that the probability of social transmission is strongly influenced by social relationships between birds. The practical implications of understanding the role of social learning in the spread of injurious behaviors in this economically important species are briefly considered.
	Address	Department of Clinical Veterinary Science, University of Bristol, Langford, Bristol, England. c.j.nicol@bristol.ac.uk
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1543-4494	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:15161142			Approved	no
	Call Number	refbase @ user @			Serial	75
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	Author	Sundaresan, S.R.; Fischhoff, I.R.; Dushoff, J.; Rubenstein, D.I.
	Title	Network metrics reveal differences in social organization between two fission-fusion species, Grevy's zebra and onager			Type	Journal Article
	Year	2007	Publication	Oecologia	Abbreviated Journal	Oecologia
	Volume	151	Issue	1	Pages	140-149
	Keywords	Analysis of Variance; Animals; Behavior, Animal/physiology; Environment; Equidae/physiology; Female; India; Kenya; Male; Models, Theoretical; Social Behavior; Social Environment
	Abstract	For species in which group membership frequently changes, it has been a challenge to characterize variation in individual interactions and social structure. Quantifying this variation is necessary to test hypotheses about ecological determinants of social patterns and to make predictions about how group dynamics affect the development of cooperative relationships and transmission processes. Network models have recently become popular for analyzing individual contacts within a population context. We use network metrics to compare populations of Grevy's zebra (Equus grevyi) and onagers (Equus hemionus khur). These closely related equids, previously described as having the same social system, inhabit environments differing in the distribution of food, water, and predators. Grevy's zebra and onagers are one example of many sets of coarsely similar fission-fusion species and populations, observed elsewhere in other ungulates, primates, and cetaceans. Our analysis of the population association networks reveals contrasts consistent with their distinctive environments. Grevy's zebra individuals are more selective in their association choices. Grevy's zebra form stable cliques, while onager associations are more fluid. We find evidence that females associate assortatively by reproductive state in Grevy's zebra but not in onagers. The current approach demonstrates the utility of network metrics for identifying fine-grained variation among individuals and populations in association patterns. From our analysis, we can make testable predictions about behavioral mechanisms underlying social structure and its effects on transmission processes.
	Address	Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544, USA. ssundare@princeton.edu
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0029-8549	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16964497			Approved	no
	Call Number				Serial	1863
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	Author	Sovrano, V.A.; Bisazza, A.; Vallortigara, G.
	Title	How fish do geometry in large and in small spaces			Type	Journal Article
	Year	2007	Publication	Animal Cognition	Abbreviated Journal	Anim. Cogn.
	Volume	10	Issue	1	Pages	47-54
	Keywords	Animals; Association Learning; Color Perception; Cues; Discrimination Learning; Distance Perception; Fishes; Male; Pattern Recognition, Visual; Social Environment; *Space Perception; Visual Perception
	Abstract	It has been shown that children and non-human animals seem to integrate geometric and featural information to different extents in order to reorient themselves in environments of different spatial scales. We trained fish (redtail splitfins, Xenotoca eiseni) to reorient to find a corner in a rectangular tank with a distinctive featural cue (a blue wall). Then we tested fish after displacement of the feature on another adjacent wall. In the large enclosure, fish chose the two corners with the feature, and also tended to choose among them the one that maintained the correct arrangement of the featural cue with respect to geometric sense (i.e. left-right position). In contrast, in the small enclosure, fish chose both the two corners with the features and the corner, without any feature, that maintained the correct metric arrangement of the walls with respect to geometric sense. Possible reasons for species differences in the use of geometric and non-geometric information are discussed.
	Address	Department of General Psychology, University of Padua, Via Venezia, 8, 35131, Padova, Italy. valeriaanna.sovrano@unipd.it
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	1435-9448	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16794851			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	2462
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