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Davies Morel, M.C.G.; Newcombe, J.R.; Holland, S.J. |
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Factors affecting gestation length in the Thoroughbred mare |
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2002 |
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Animal Reproduction Science |
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74 |
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3-4 |
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175-185 |
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Gestation length; Mare age; Foals; Month of year; Ovulation-mating interval |
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In order to assist in the accurate prediction of the timing of parturition in the mare true gestation length, along with the potential effect of a number of factors, was investigated. Data from 433 Thoroughbred foal pregnancies were used. Sequential ultrasonic scanning allowed the true gestation length (fertilisation-parturition) to be ascertained, as apposed to previous work, which used the mating-parturition interval. An average gestation length of 344.1+/-0.49 days was evident. Colt foal pregnancies were significantly (P<0.001) longer (346.2+/-0.72) than fillies (342.4+/-0.65). Month of birth had a significant effect on gestation length in all foals (P<0.001). With foals born in January having the shortest gestation lengths and those born in April the longest. Mare age, year of birth, stallion age, stud farm and the interval between ovulation and mating had no significant effect. It is concluded that (i) the gestation length range (315-388 days), all resulting in viable foals is noteworthy and of clinical importance when considering the classification of dysmaturity in foals, (ii) mares carrying colt foals due to be born in the middle of the breeding season (April) are likely to have the longer gestation lengths. |
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Equine Behaviour @ team @ |
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3638 |
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Ernst, K.; Puppe, B.; Schon, P.C.; Manteuffel, G. |
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A complex automatic feeding system for pigs aimed to induce successful behavioural coping by cognitive adaptation |
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2005 |
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Applied Animal Behaviour Science |
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Appl. Anim. Behav. Sci. |
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91 |
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3-4 |
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205-218 |
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Learning; Cognition; Reward; Welfare; Pig |
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In modern intensive husbandry systems there is an increasing tendency for animals to interact with technical equipment. If the animal-technology interface is well-designed this may improve animal welfare by offering challenges for cognitive adaptation. Here a system and its application is presented that acoustically calls individual pigs out of a group (n = 8) to a feeding station. In three different learning phases, the computer-controlled “call-feeding-station” (CFS) trained the animals to recognize a specific acoustic signal as a summons for food, using a combination of classical and operant conditioning techniques. The experimental group's stall contained four CFSs, at each of which one animal at a time was able to feed. When an animal had learned to discriminate and recognize its individual acoustic signal it had to localize the particular CFS that was calling and to enter inside it. Then, it received a portion of feed, the amount of which was adapted to the respective age of the animals. Each animal was called at several, unpredictable times each day and the computer programme ensured that the total feed supply was sufficient for each animal. In the last phase of the experiment the animals, in addition, had to press a button with an increasing fixed ratio for the delivery of feed. It was demonstrated that the pigs were able to adapt quickly to the CFSs. Although they were challenged over 12 h daily by requirements of attention, sensory localization and motor efforts to gain comparatively low amounts of feed, they performed well and reached fairly constant success rates between 90 and 95% and short delays between 14 and 16 s between a summons and the food release in the last phase of the experiment. The weight gain during the experiment was the same as in a conventionally fed control group (n = 8). We therefore conclude that CFSs present a positive challenge to the animals with no negative effects on performance but with a potentially beneficial role for welfare and against boredom. The system is also a suitable experimental platform for research on the effects of successful adaptation by rewarded cognitive processes in pigs. |
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Equine Behaviour @ team @ |
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2898 |
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Wolter, R.; Krueger, K. |
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The analysis of social bonds in feral horses |
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Conference Article |
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2015 |
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Proceedings of the 3. International Equine Science Meeting |
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Proc. 3. Int. Equine. Sci. Mtg |
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social bonds, grooming, nearest neighborhood, rank |
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In many social mammals, individuals preferentially affiliate with a small subset of available partners instead of distributing their social behaviors equally among all group members. The resulting social bonds have been investigated in several mammalian taxa, especially in primate societies, but also in other taxa such as birds, dolphins, rodents and ungulates. In feral horses, a great number of studies on social bonds can be found, but with a huge variety between methods for the analysis. There seems to be a lack of a clear and common definition of social bonds in horses and of comparable analyses. For example, there are irregularities between the studies regarding the research designs, the selection of recording methods and the interpretation of the measurements. Mutual grooming is used most often for the analysis of social relationships in many species. As mutual grooming is rare in horses, especially measurements of spatial proximity are commonly used for the analysis of social bonds in addition to other behavioral patterns. However, the combination of mutual grooming and nearest neighborhood analyses for the analysis of social bond is debatable, as in contrast to mutual grooming, which must occur deliberately by both grooming partners, the spatial distribution can be influenced by one partner alone, which may even force the other horse to keep a certain distance or to stay in close proximity.
In this study, we investigated the comparability of mutual grooming and nearest neighborhood data for social bond analyses in feral horses. Therefore, we observed five groups of semi-wild living Przewalski’s horses and six groups of feral horses.
We analysed the horses’ social ranks by applying an Average Dominance Index, we recorded the distances between the animals and observed the number of mutual grooming events as well as friendly approaches.
Our results show that there was only a weak correlation between the frequency of staying in nearest neighborhood and mutual grooming in all observed horse groups. In contrast to this, the correlation between the number of friendly approaches and mutual grooming events was higher in most groups.
Hierarchies did not affect social bonds, as mutual grooming was similarly induced by higher and lower ranking animals and the social rank did not affect the choice of the grooming partner. Similarly, likelihoods of staying in the neighborhood of particular animals were not affected by the animals’ social rank.
The grooming frequencies differed between the different horse groups and between the individual horses living in the particular groups. They seem to be effected by individual predisposition.
Altogether we suggest that the ratio of mutual grooming seems to be a better indicator for social bonds in feral horses than the frequency of staying in the nearest neighborhood. Mutual grooming occurs deliberately and is bidirectional, whereas nearest neighborhoods could be enforced and unidirectional. For the calculation of social bonds in horses, we consider it to be more reliable to combine the frequency of mutual grooming with the frequency of friendly approaches. |
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Wolter, R. |
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978-3-95625-000-2 |
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Equine Behaviour @ team @ |
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5882 |
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Cameron, E.Z.; Setsaas, T.H.; Linklater, W.L. |
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Social bonds between unrelated females increase reproductive success in feral horses |
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2009 |
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Proceedings of the National Academy of Sciences of the United States of America |
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Proc. Natl. Acad. Sci. U.S.A. |
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106 |
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33 |
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13850-13853 |
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In many mammals, females form close social bonds with members of their group, usually between kin. Studies of social bonds and their fitness benefits have not been investigated outside primates, and are confounded by the relatedness between individuals in primate groups. Bonds may arise from kin selection and inclusive fitness rather than through direct benefits of association. However, female equids live in long-term social groups with unrelated members. We present 4 years of behavioral data, which demonstrate that social integration between unrelated females increases both foal birth rates and survival, independent of maternal habitat quality, social group type, dominance status, and age. Also, we show that such social integration reduces harassment by males. Consequently, social integration has strong direct fitness consequences between nonrelatives, suggesting that social bonds can evolve based on these direct benefits alone. Our results support recent studies highlighting the importance of direct benefits in maintaining cooperative behavior, while controlling for the confounding influence of kinship. |
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10.1073/pnas.0900639106 |
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Equine Behaviour @ team @ |
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5152 |
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Leliveld, L.M.C.; Düpjan, S.; Tuchscherer, A.; Puppe, B. |
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Hemispheric Specialization for Processing the Communicative and Emotional Content of Vocal Communication in a Social Mammal, the Domestic Pig |
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2020 |
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Frontiers in Behavioral Neuroscience |
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Front. Behav. Neurosci. |
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14 |
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596758 |
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In humans, speech perception is lateralized, with the left hemisphere of the brain dominant in processing the communicative content and the right hemisphere dominant in processing the emotional content. However, still little is known about such a division of tasks in other species. We therefore investigated lateralized processing of communicative and emotionally relevant calls in a social mammal, the pig (Sus scrofa). Based on the contralateral connection between ears and hemispheres, we compared the behavioural and cardiac responses of 36 young male pigs during binaural and monaural (left or right) playback to the same sounds. The playback stimuli were calls of social isolation and physical restraint, whose communicative and emotional relevance, respectively, were validated prior to the test by acoustic analyses and during binaural playbacks. There were indications of lateralized processing mainly in the initial detection (left head-turn bias, indicating right hemispheric dominance) of the more emotionally relevant restraint calls. Conversely, there were indications of lateralized processing only in the appraisal (increased attention during playback to the right ear) of the more communicative relevant isolation calls. This implies differential involvement of the hemispheres in the auditory processing of vocalizations in pigs and thereby hints at similarities in the auditory processing of vocal communication in non-human animals and speech in humans. Therefore, these findings provide interesting new insight in the evolution of human language and auditory lateralization. |
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1662-5153 |
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Equine Behaviour @ team @ |
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6699 |
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Duncan, I.J.; Petherick, J.C. |
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The implications of cognitive processes for animal welfare |
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1991 |
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Journal of Animal Science |
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J. Anim Sci. |
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69 |
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12 |
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5017-5022 |
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*Animal Welfare; Animals; Animals, Domestic/*psychology; *Cognition |
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In general, codes that have been designed to safeguard the welfare of animals emphasize the importance of providing an environment that will ensure good health and a normal physiological and physical state, that is, they emphasize the animals' physical needs. If mental needs are mentioned, they are always relegated to secondary importance. The argument is put forward here that animal welfare is dependent solely on the cognitive needs of the animals concerned. In general, if these cognitive needs are met, they will protect the animals' physical needs. It is contended that in the few cases in which they do not safeguard the physical needs, it does not matter from a welfare point of view. The human example is given of being ill. It is argued that welfare is only adversely affected when a person feels ill, knows that he or she is ill, or even thinks that he or she is ill, all of which processes are cognitive ones. The implications for welfare of animals possessing certain cognitive abilities are discussed. For example, the extent to which animals are aware of their internal state while performing behavior known to be indicative of so-called states of suffering, such as fear, frustration, and pain, will determine how much they are actually suffering. With careful experimentation it may be possible to determine how negative they feel these states to be. Similarly, the extent to which animals think about items or events absent from their immediate environment will determine how frustrated they are in the absence of the real item or event but in the presence of the cognitive representation. |
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University of Guelph, Canada |
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English |
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0021-8812 |
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PMID:1808195 |
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Equine Behaviour @ team @ |
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2753 |
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König von Borstel, U.; Kienapfel, K.; McLean, A.; Wilkins, C.; Evans, D.; McGreevy, P. |
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Hyperflexing the horse‘s neck: a cost-benefit and meta-analysis |
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Conference Article |
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2015 |
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Proceedings of the 3. International Equine Science Meeting |
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Proc. 3. Int. Equine. Sci. Mtg |
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horse, head-and-neck posture, hyperflexion, welfare, gymnastics |
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In ethical discussions, a cost-benefit analysis requires that welfare costs associated with an activity can be reliably estimated and balanced against the potential benefits of the activity to both humans and animals. The current study applies a meta-analysis to the peer-reviewed evidence for costs and benefits of hyperflexion of the neck in horses; a practice that has attracted enormous public and scientific scrutiny over the past 15 years. A literature review identified 55 studies dealing with horses’ head and neck postures. Fourty-two of these studies examined the impact of various postures on equine welfare, for example, by assessing behavior, physiological stress parameters, health or rider-horse interaction. Thirty-five studies examined the impact of various postures on gymnastics (e.g. kinematics, shifts in weight distribution, muscle activity, airway functioning or overall workload). For the meta-analysis a dataset containing information from each of the individual studies was created. Data included information such as type, degree, duration and circumstances of hyperflexion applied in that particular study as well as information on the horses (e.g., sport discipline, level of training, breed) and on the study design (e.g., size of study and experimental or epidemiological research design). The results of the study regarding the impact of hyperflexion on a) welfare and b) gymnastics were coded as positive (1), insignificant or contradictory (0) or negative (-1). The significant majority of studies (88%) concluded that a hyperflexed head and neck posture negatively impacts welfare. Just one study suggested welfare advantages of training in a hyperflexed head and neck posture. An analysis using a generalized linear mixed model to assess the influence of the above factors collated in the dataset revealed that none of these factors significantly influenced the probability of a study to detect negative welfare implications. Thus hyperflexing the neck appears to impair horses’ welfare regardless of, for example, the duration or the way of achieving hyperflexion. A concurrent assessment of the evidence for gymnastic benefits showed that approximately one quarter of studies conclude that there may be benefits, while another quarter of the studies conclude that hyperflexion has detrimental effects on gymnastics. Thus, on the costs-side there is a clear reduction in equine welfare and some undesirable gymnastic effects, as well as likely a compromised profile of the equestrian sports in public. Benefits, on the other hand, include some desirable gymnastic effects, and potentially increased control of the horse for the rider. On balance, it appears that the costs associated with hyperflexion exceed the potential benefits of the activity to both humans and horses. |
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König von Borstel, U. |
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Equine Behaviour @ team @ |
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5872 |
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Hinrichs, T. |
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Gruppen Management in Aktiv Ställen |
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Conference Article |
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2012 |
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Proceedings of the 2. International Equine Science Meeting |
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Proc. 2. Int. Equine. Sci. Mtg |
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in press |
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In einem Aktivstall leben die Pferde in einer Gruppe und sind im ständigen Sozialkontakt mit ihren Artgenossen. Getrennte Funktionsbereiche kennzeichnen das Konzept. Das Kernstück eines Aktivstalles ist die automatisierte Fütterung für Kraft,- Mineral,- und Raufutter. Sie gewährleistet die bedarfsgerechte Fütterung und regt die Pferde zur vermehrten Bewegungsaktivität an. Die Gestaltung der Funktionsbereiche eines Aktivstalles hängt von mehreren Faktoren wie den örtlichen Gegebenheiten, geplanter Gruppengröße und -zusammensetzung sowie betriebs- und arbeitwirtschaftlichen Aspekten ab.
Die Grundlage für die Planung eines Aktivstall – Konzeptes sind die laufenden Erkenntnisgewinne aus Wissenschaft und Forschung, die praktischen Erfahrungen auf Pferdebetrieben sowie die länderspezifischen, offiziellen Richtlinien und Bestimmungen zur Pferdehaltung.
So sind z. B. in Deutschland die aktuellen Untersuchungen zu den Themen automatisierte Fütterungssysteme und Liegeflächengestaltung im Ruheraum von Frau Dr. Zeitler – Feicht und ihrem Team, den ca. 500 Aktivstallbetrieben und den seit dem Jahr 2009 geltenden Leitlinien zur Pferdehaltung unter Tierschutzgesichtspunkten wesentliche Bestimmungsparameter für die Weiterentwicklung des Aktivstall – Konzeptes.
Auf dem IESM 2012 wird den Teilnehmern das Gruppenmanagement in Aktivställen am Beispiel von Praxisbetrieben vorgestellt. Ziel ist es, eine kontinuierliche Verbesserung der Pferdehaltung im Sinne des Tierschutzes in Verbindung mit der Nutzung der Pferde zu erreichen. |
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Hinrichs, T. |
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Xenophon Publishing |
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Wald |
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Krueger, K. |
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978-3-9808134-26 |
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Public Day |
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Equine Behaviour @ team @ |
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5567 |
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Zeitler-Feicht, M.H.; Streit, S. |
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Checkpunkte für eine tiergerechte Fütterung von Pferden an Abrufautomaten |
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Conference Article |
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2012 |
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Proceedings of the 2. International Equine Science Meeting |
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Proc. 2. Int. Equine. Sci. Mtg |
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in press |
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In der Offenstallhaltung von Pferden wird für die individuelle Fütterung der Fressstand, der eine art-typische synchrone Nahrungsaufnahme erlaubt, empfohlen (BMELV 2009). Seit einigen Jahren kommen jedoch vermehrt Abrufstationen für Kraft- und Raufutter zum Einsatz. Diese ermöglichen nur ein asynchrones Fressen, was das Risiko für Auseinandersetzungen im Fressbereich erhöht.
In einer repräsentativen Studie an insgesamt 567 individuell unterschiedenen Pferden in 32 Offen-laufställen mit Abrufstationen und 10 Betrieben mit Fressständen wurden die Aufenthaltsdauer und Besuchshäufigkeit sowie das Droh- und Meideverhalten der Pferde im Fütterungsbereich und die Häu-figkeit von Blockaden erfasst, des Weiteren Herzfrequenz und Integumentverletzungen. Fazit war, dass es an den Abrufstationen doppelt so oft zu Auseinandersetzungen kam als an den Fressständen. Als Ursache hierfür konnte die starke Frequentierung der Automaten eruiert werden. Umso wichtiger ist somit die tiergerechte Ausführung der Abrufautomaten. In der Studie konnte nachgewiesen werden, dass Durchlaufstationen pferdegerechter sind als Rücklaufstationen. Als nicht tiergerecht wurden Stationen mit Strom führender Austreibehilfe.eingestuft. Die für das fressende Pferd als verhaltens-gerecht empfohlene Variante (Fressstand mit Eingangssperre und ohne Austreibehilfe) führte zu einer Erhöhung der Besuchshäufigkeit und Aufenthaltsdauer und somit auch zu einer erhöhten Anzahl an Drohgesten je Tier und Tag. Die Herzfrequenz lag im Wartebereich bei den meisten Pferden im physio-logischen Bereich (45,1 ± 12,42 Schläge/min), erhöhte sich jedoch in der Abrufstation um ca. 20 Schlä-ge/min. Einige Tiere zeigten möglicherweise stressbedingt auffallend hohe Herzfrequenzwerte ( 100 Schläge/min). Integumentverletzungen im Zusammenhang mit dem Fütterungssystem traten nicht auf. Das Flächengebot und die Konzeption der Offenstallanlage sowie das Management erwiesen sich als maßgebliche Einflussfaktoren auf alle überprüften Kriterien zur Tiergerechtheit.
Um die Anzahl an Auseinandersetzungen im Fütterungsbereich von Abrufstationen zu mindern wird eine Reduzierung der Fütterungshäufigkeit von üblicherweise bis zu 24 Mahlzeiten auf nur 10 Mahlzeiten empfohlen, gemäß dem natürlichen Fressrhythmus von Pferden. Unabhängig vom Fütterungssystem traten die wenigsten Auseinandersetzungen auf, wenn die Pferde täglich etwa 1,5 kg Heu pro 100 kg Körpermasse erhielten bei zusätzlicher ad Libitum Fütterung von Stroh.
Nach bisherigen Erkenntnissen ergeben sich nachfolgende Checkpoints für Abrufautomaten:
Checkpunkte für Abrufstationen
 Schutz des Pferdes durch Trennwände über die gesamte Körperlänge
 Trennwände mit Sichtkontakt zu den anderen Pferden
 Durchlaufstationen mit separatem Ein- und Ausgang
 Eingangssperre für eine stressfreie Futteraufnahme
 Ausgang mit Pendeltüre und Rücklaufsperre
 Ausgang in einen anderen Funktionsbereich
 Ausreichend Ausweichmöglichkeiten für rangniedere Tiere
 Räumliche Trennung von Kraftfutter- und Heuautomaten
 Maximal je 10 Kraft- und Grobfutterrationen je Tier und Tag
 ≥ 1,5 kg Heu bzw. Heulage pro 100 kg Körpermasse je Pferd und Tag
 Stroh zur freien Verfügung (Möglichkeit zum synchronen Fressen)
 Individuelles Anlernen der Pferde am Automaten erforderlich |
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Address |
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Corporate Author |
Zeitler-Feicht, M.H. |
Thesis |
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Publisher |
Xenophon Publishing |
Place of Publication |
Wald |
Editor |
Krueger, K. |
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Language |
Deutsch |
Summary Language |
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Original Title |
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Series Editor |
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Series Title |
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Abbreviated Series Title |
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Series Volume |
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Series Issue |
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Edition |
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ISSN |
978-3-9808134-26 |
ISBN |
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Expedition |
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Conference |
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Notes |
Public Day |
Approved |
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Call Number |
Equine Behaviour @ team @ |
Serial |
5568 |
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| Permanent link to this record |
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Author |
Klingel, H . |
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Title |
Social Organisation and Social Behaviour of the Equids |
Type |
Conference Article |
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Year |
2012 |
Publication |
Proceedings of the 2. International Equine Science Meeting |
Abbreviated Journal |
Proc. 2. Int. Equine. Sci. Mtg |
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Volume |
in press |
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Keywords |
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Abstract |
In contrast to the great similarity in behaviour and ecology of the 6 extant Equid species, 2 distinct types of social organisation have evolved, and both are adapted to life in semi-arid to arid regions where environmental conditions force them to migrate seasonally or opportunistically.
The ranges of the various species overlap: Mountain Zebra Equus zebra and Plains Zebra E. quagga in South Africa and Namibia, Plains Zebra and Grevy's Zebra E. grevyi in Kenya and Ethiopia, Grevy's Zebra and African Wild Ass E. africanus in Ethiopia, Asiatic Wild Ass E. hemionus and Przewalski Horse E. przewalski in Mongolia and China. Although, in the overlap zones, individuals of the different species are using the same resources like water and grazing next to each other, they rarely make closer contacts.
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In the type 1 species, Horse, Plains Zebra and Mountain Zebra, the adults live in non-territorial, stable, one-male families and as single bachelors and in bachelor groups. Family stallions have the exclusive mating rights with the mares in their harems. These consist of up to 6 unrelated mares plus their offspring, totalling up to 20 members.
Mares stay in their harem until death. Stallions' tenure is from age 5-6 years, i.e. when they succeed in controlling a harem, for close to life time, but are replaced when dead or incapacitated. Harems are stable even in the absence of a stallion, indicating voluntary membership. Adolescent mares leave their parental families to become members of another harem.
In Plains Zebra the adolescent mares are abducted, during an oestrus, by suitors who fight the defending family stallion/father. Successful stallions are bachelors who start a family, or family stallions enlarging their harem. Young stallions leave their parental families voluntarily at age 2-3 years and join bachelor stallion groups from where the family stallions are recruited.
An individualised dominance hierarchy excists with the stallion in the alpha position. It is based on individual knowledge and recognition of the members.
In the type 2 species Grevy's Zebra, African Wild Ass and Asiatic Wild Ass adult stallions monopolise territories in which they have the exclusive mating rights. Stallions are tolerant of any conspecifics entering their territory. Bachelor stallions behave subordinately – or fight for the possession of the territory which is a prerequisite for reproduction.
Mares join up to form anonymous and unstable groups or herds. The only stable unit is of a mare and her offspring. In Grevy's Zebra mares with foal join preferentially conspecifics of the same soial status, as do mares without foal.
Matings take place inside the territory. There is no lasting relationship of the mare with a particular stallion, and the mare may be mated by any stallion whose territory she is visiting.
Territories measure up to 10 or more square kilometres, and tenure is for several years.
Grevy Zebra territorial owners leave their territories for a few hours to visit a water hole, or for months when grazing and water conditions are below requirements, and re-occupy it upon return, unchallenged. |
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Address |
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Corporate Author |
Klingel, H . |
Thesis |
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Publisher |
Xenophon Publishing |
Place of Publication |
Wald |
Editor |
Krueger, K. |
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Language |
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Summary Language |
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Original Title |
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Series Editor |
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Series Title |
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Abbreviated Series Title |
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Series Volume |
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Series Issue |
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Edition |
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ISSN |
978-3-9808134-26 |
ISBN |
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Medium |
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Area |
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Expedition |
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Conference |
IESM 2012 |
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Notes |
Invited speaker IESM 2012 |
Approved |
no |
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Call Number |
Equine Behaviour @ team @ |
Serial |
5436 |
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| Permanent link to this record |
