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Ord, T. J., & Evans, C. S. (2002). Interactive video playback and opponent assessment in lizards. Behav. Process., 59(2), 55–65.
Abstract: Video playback has been used to explore many issues in animal communication, but the scope of this work has been constrained by the lack of stimulus-subject interaction. In many natural contexts, each participant's signalling behaviour is dependent from moment-to-moment on that of the other. Analyses of acoustic communication demonstrate the value of reproducing such social contingencies. We assessed the utility of interactive playback for studies of visual signalling by comparing the responses of male Jacky dragons, Amphibolurus muricatus, to interactive and non-interactive digital video playbacks of a life-sized conspecific. Displays produced by lizards in the interactive condition had the effect of suppressing the aggressive display of their simulated opponent. Each stimulus sequence generated during an interactive playback was subsequently played to a size-matched control animal. Males that could interact with the video stimulus responded principally with aggressive displays, while those that could not produced a mixture of aggressive and appeasement signals. Adding a degree of receiver responsiveness is hence sufficient to alter the type of signal evoked, even when video stimuli are physically identical. Interactive playback permits the experimental study of a broader range of theoretical topics and can enhance the realism of video stimuli.
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Hodson, E. F., Clayton, H. M., & Lanovaz, J. L. (1999). Temporal analysis of walk movements in the Grand Prix dressage test at the 1996 Olympic Games. Appl. Anim. Behav. Sci., 62(2-3), 89–97.
Abstract: Video analysis was used to measure temporal characteristics of the collected walk, extended walk and half pirouette at walk of eleven competitors during the team dressage competition at the 1996 Summer Olympic Games in Atlanta, GA. Forelimb stance durations, hind limb stance durations, lateral step intervals and diagonal step intervals were symmetrical for the right and left sides in the collected and extended walk strides, but there were left-right asymmetries in the forelimb stance duration and in the lateral step interval in the half pirouette strides. For both collected and extended walk strides, hind limb stance duration was significantly longer than forelimb stance duration. The mean values for the group of eleven horses showed that the collected and extended walks had a regular rhythm. The half pirouette strides showed an irregularity in which there was a short interval between footfalls of the outside forelimb and inside hind limb, and along interval between footfalls of the inside hind limb and inside forelimb. This irregularity reflected an early placement of the inside hind limb. The stance times of both hind limbs were prolonged and this finding, in combination with the early placement of the inside hind limb, led to an increase in the period of tripedal support in each stride of the half pirouette. This was interpreted as a means of maintaining the horses' balance in the absence of forward movement.
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Windschnurer, I., Zeitlinger, S., Troxler, J., & Waiblinger, S. (2012). Human and horse behaviour during veterinary treatments. In K. Krueger (Ed.), Proceedings of the 2. International Equine Science Meeting (Vol. in press). Wald: Xenophon Publishing.
Abstract: Veterinary treatments are often aversive or painful, resulting in stress, avoidance, or aggressive behaviour. This preliminary study investigated which behaviours horses and humans display during veterinary examinations or treatments and if specific behaviours are shown more often during particular situations . Moreover, possible links between human and equine behaviour were investigated. Behaviours of horses, veterinarians, and assisting people were recorded at 5 clinics during 49 routine treatments that were divided into 111 situations (assigned to 7 categories: giving injections, waiting, leading horses, inspections with light body contact, invasive inspections, leg inspections, spraying/shaving). Behaviours of humans present during a situation were summarised. Behaviours (frequencies/minute) were analysed by Kruskal-Wallis tests and Spearman rank correlations. Horses showed insecurity (stopping, baulking, defecation) with different frequencies in the different categories (Chi2=23.38, df=6, P=0.001), e.g., often during leading. In humans, activating behaviour (e.g., arm waving, whistling, tongue clicking) was displayed with different frequencies (Chi2=20.42, df=6, P=0.002), e.g., frequently during leading. Restlessness in horses (e.g., stepping, head movements) tended to occur with different frequencies (Chi2=12.37, df=6, P=0.054), e.g., often during invasive inspections. Humans applied restraining techniques considered to be painful (e.g., ear twisting) with different frequencies (Chi2=26.20, df=6, P<0.001), e.g., mostly during invasive inspections. Restlessness in horses correlated with negative human interactions (e.g., hitting, talking impatiently, shouting: rs=0.28, P=0.002, n=111) and neutral human interactions (e.g., slight hits, talking dominantly: rs=0.45, P<0.001, n=111) as well as with the use of painful restraint techniques (rs=0.46, P<0.001, n=111). Moreover, avoidance behaviour of the horses (turning and stepping away) correlated with negative human interactions (rs=0.30, P=0.002, n=111). Thus, humans and horses showed certain behaviours in different situations with differing frequencies. Moreover, links between human and animal behaviour were found. Future studies should analyse causal connections, i.e., if the observed animal behaviour is the result of/or the reason for human interactions by sequence analysis. KW -
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Waiblinger, S., Menke, C., Korff, J., & Bucher, A. (2004). Previous handling and gentle interactions affect behaviour and heart rate of dairy cows during a veterinary procedure. Appl. Anim. Behav. Sci., 85(1-2), 31–42.
Abstract: Veterinary and management procedures often are aversive to the animals, resulting in physiological and behavioural stress reactions, which increase the risk of accidents and might lower performance. We investigated the effects of previous positive handling and of gentle interactions during the procedure on behaviour and heart rate in dairy cows during rectal palpation with sham insemination. Twenty cows were allocated randomly into two groups of 10 animals: handling, received additional positive handling over a period of four weeks by one person (handler); control, only routine handling by different caretakers. The week after the handling period, tests lasting 9 min, including 4 min rectal palpation, were carried out with each animal on four successive days in four situations in a balanced order: cow is alone during the test, with the handler, with an usual caretaker, or with an unknown person. Behaviour and heart rate were recorded. Previously handled animals had lower heart rate during tests (P<=0.05, n=19), kicked less when alone (P<=0.05, n=19) and tended to show less restless behaviour (P<=0.1, n=19). Cows were further calmed by gentle interactions during the test, but people differed remarkably. Cows showed less restless behaviour when gentled by the handler, both in the 4 min of rectal palpation and in the 9 min test period (each: P<=0.001, n=19). No significant stress reducing effect was found for the other two persons. In conclusion, stress reactions of cows during rectal palpation/insemination can be reduced by previous positive handling as well as by a person providing positive, gentle interactions during the procedure. The results underline the importance of positive, gentle interactions with the animals to enhance animal welfare and reduce the risk of accidents. They also show that people differ in the success to calm down the animals in aversive situations and indicate the need to investigate the characteristics responsible for the differences and identify the preconditions for a stress reducing effect in future research.
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Villani, M., Cairoli, F., Kindahl, H., Galeati, G., Faustini, M., Carluccio, A., et al. (2006). Effects of mating on plasma concentrations of testosterone, cortisol, oestrone sulphate and 15-ketodihydro-PGF2alpha in stallions. Reprod Domest Anim, 41(6), 544–548.
Abstract: Very little information is available regarding the physiological mechanisms involved in the normal sexual activity in the stallion and, in particular, the endocrine control of reproduction is still not clearly understood. This experiment was designed to determine the short-term effect of sexual stimulation on plasma concentrations of testosterone, cortisol, oestrone sulphate and 15-ketodihydro-PGF(2alpha) in stallions. Semen samples were collected from 10 lighthorse stallions of proven fertility using a Missouri model artificial vagina. At the same time, blood samples were collected from the jugular vein with heparinized tubes, 20 and 10 min before oestrous mare exposure, at exposure and 10, 20, 30 min after dismounting. Testosterone concentrations showed a sharp rise 10 min after mating (p < 0.001), reached a plateau, and then showed a further increase 30 min after mating (p < 0.001). Cortisol concentrations increased 10 min after mating (p < 0.001) and remained at high levels in the subsequent samples taken. A peak of oestrone sulphate was observed 10 min after mating (p < 0.001). 15-Ketodihydro-PGF(2alpha) concentrations decreased rapidly at the moment of the exposure of the stallions to an oestrous mare (p < 0.05), returned to pre-mating concentrations and then decreased again 30 min after mating (p < 0.05).
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Seyfarth, R. M., Cheney, D. L., & Marler, P. (1980). Monkey responses to three different alarm calls: evidence of predator classification and semantic communication. Science, 210(4471), 801–803.
Abstract: Vervet monkeys give different alarm calls to different predators. Recordings of the alarms played back when predators were absent caused the monkeys to run into trees for leopard alarms, look up for eagle alarms, and look down for snake alarms. Adults call primarily to leopards, martial eagles, and pythons, but infants give leopard alarms to various mammals, eagle alarms to many birds, and snake alarms to various snakelike objects. Predator classification improves with age and experience.
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Töpfer, D., Wolter, R., & Krueger, K. (2014). Fallstudie zum Platzangebot, Verhalten und Wohlbefinden der Pferde (Equus caballus) in Mehrraum-Außenlaufställen mit Aus-lauf und Bewegungsställen [A case study to space, behavior and well-being of horses (Equus caballus) in open stables and open active stables]. KTBL Schriften. Darmstadt: KTBL.
Abstract: Verhaltensbeobachtungen erfolgten an 112 Pferden in Mehrraum-Außenlaufställen mit Auslauf und Bewegungsställen. Bei vermehrtem Platzangebot sinkt das agonistische Verhalten in Bewegungsställen außerhalb des Fressbereiches. Mit zunehmender Stallgröße (überdachte Fläche und Auslauf) steigt in beiden Haltungsverfahren das affiliative Verhalten während das agonistische Verhalten mit Verletzungsrisiko sinkt. Daher wird für beide Haltungsverfahren ein möglichst großzügiger Stall je Pferd empfohlen. Die beobachteten Erkrankungen der letzten zwölf Monate zeigen vermehrt Hautkrankheiten in den Mehrraum-Außenlaufställen mit Auslauf, wohingegen Hufgeschwüre in Bewegungsställen auftraten. Wird der Fütterungsaspekt in den Bewegungsställen betrachtet, so ist die Transponderfütterung in Kombination mit ad libitum Fütterung aufgrund des deutlich geringeren, agonistischen Verhaltens beim Fressen, der reinen Transponderfütterung vorzuziehen. Mehrraum-Außenlaufställe mit Auslauf haben den Vorteil der synchronen Nahrungsaufnahme.
[The behaviour of 112 horses was observed in open stables and open active stables. Horses show less agonistic behaviour outside of the feeding area in open active stables as the space for each horse is augmented. Also the affiliative behaviour increases as a function of more space per horse while agonistic behaviour with a risk for injury decreases. These results are independent from the husbandry system. Due to this fact a stable with a larger area per horse is recommended. Diseases of the last twelve month were recorded and show skin diseases in open stables and abscesses in the hooves in open active stables. To consider the feeding in open active stables it was found that horses which were fed with hay controlled by transponders showed more agonistic behaviour in contrast to horses being fed using transponders but with access to hay ad libitum. Thus transponder feeding in combination with ad libitum feeding is recommended. Open stables have the advantage of synchronized feeding.]
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Klingel, H.. (2012). Soziale Organisation und Sozialverhalten der Equiden. In K. Krueger (Ed.), Proceedings of the 2. International Equine Science Meeting (Vol. in press). Wald: Xenophon Publishing.
Abstract: Verhalten und Ökologie der 6 rezenten Equiden sind in vieler Hinsicht identisch, jedoch in der Sozialen Organisation haben 2 deutliche verschiedene Formen evoluiert, die beide an das Leben in den semi-ariden und ariden Lebensräumen angepasst sind, wo sie zu säsonalen oder opportunistischen Wanderungen gezwungen sind.
Die Verbreitungsgebiete der verschiedenen Arten überlappen, in Südafrika und Namibia von Bergzebra Equus zebra und Steppenzebra E. quagga, in Kenya und Äthiopien von Steppenzebra und Grevy-Zebra E. grevyi, in Äthipien und Somalia von Grevy-Zebra und Afrikanischem Wildesel E. africanus, in China und der Mongolei Asiatischer Wildesel E. hemionus und Przewalski-Pferd E. przewalskii. Obwohl die Vertreter der verschiedenen Arten in den Überschneidungsgebieten die gleichen Ressourcen wie Wasser und Weide nutzen, nehmen sie kaum Kontakt zueinander auf.
Die Vertreter von Typ 1, Steppenzebra Equus quagga, Bergzebra E..zebra, Pferd E przewalskii, leben in nicht-territorialen , dauerhaften 1- Hengst- Familien, in Hengstgruppen und als Einzelgänger.. Die Familienhengste haben die alleinigen Paarungsrechte mit den Stuten in ihrem Harem. Dieser besteht aus bis zu ca. 6 nicht-verwandten Stuten nebst ihren Nachkommen und kann bis 20 Mitglieder haben.
Stuten bleiben bis zu ihrem Tod im Harem..Hengste können mit 5-6 Jahren einen Harem erobern oder gründen, können gleichfalls bis zum Tod die Familie begleiten, werden aber meist vorher von einem anderen Hengst ersetzt. Harems sind auch ohne Hengst stabil, ein Hinweis, dass die Stuten freiwilling im Harem sind und bleiben.. Junge Stuten verlassen ihre elterliche Familie und schliessen sich einem anderen Harem an..Beim Steppenzebra werden die Jungstuten während eines Östrus (Rosse) von Bewerbern entführt, gegen den Widerstand des Familenhengstes = Vaters. Bewerber sind Junggesellen, die so eine Familie gründen, und Familienhengste, die so ihren Harem vergrössern. Junghengste verlassen mit 2-3Jahren ihre elterliche Familie und schliessen sich Jungesellengruppen an, aus denen sich die Familenhengste rekrutieren.
In der Gruppe besteht eine Rangordnung mit dem Henst in der alpha-Position. Sie beruht aud individuellem Kennen und Erkennen der Mitglieder.
Bei Typ 2, Grevy-Zebra, Afrikanischer und Asiatischer Wildesel, monopolisieren Hengste über Jahre Territorien von 10 und mehr km2 , in denen sie die alleinigen Paarungsrechte haben. Territoriale Hengste tolerieren Artgenossen, auch erwachsene Hengste, soweit diese sich unterlegen verhalten. Oder sie stellen sich zum Kampf um den Besitz des Territoriums, eine Vorbedingung für die Fortpflanzung. Stuten im Östrus können von mehreren Hengsten begattet werden, wenn sie sich in deren Territorien aufhalten bzw diese durchwandern.
Stuten und Fohlen und nicht-territoriale Hengste schliessen sich zu anonymen instabilen Gruppen oder Herden zusammen. Feste dauerhafte Bindungen bestehen nur zwischen Stute und Fohlen. Hengste verlassen ihr Territorium für Stunden, Tage, im Extrem auch Monate, um zu Wasserstellen oder Weidegründen zu ziehen, sind aber bei Rückkehr wieder unangefochtene Besitzer.
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Donner, H. D. (1986). Grundausbildung für Reiter und Pferd (H. D. Donner, & D. Specht, Eds.) (25. Aufl. ed.). Warendorf: FN-Verl. d. Dt. Reiterlichen Vereinigung.
Abstract: Verf. gibt einen Ueberblick ueber: 1) Grundausbildung des Reiters: a) Vorbedingungen fuer eine erfolgreiche Ausbildung, b) Sitz und Sitzschulung, c) Einwirkung des Reiters, d) Reiten von Grunduebungen und Dressurlektionen, e) Springausbildung, f) Gelaendereiten sowie 2) Grundausbildung des Pferdes: a) Grundkenntnisse fuer den Ausbilder ueber Charakter und Wesen des Pferdes, Grundsaetze und Erziehung, anatomisch-physiologische Voraussetzungen, b) Allgemeine Grundausbildung, c) Grundausbildung im Springen und im Gelaende, d) Arbeit mit Pferden mit Gebaeude- und Temperamentsfehlern. Schiffer
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Scherer, W. F., Madalengoitia, J., Flores, W., & Acosta, M. (1975). Ecologic studies of Venezuelan encephalitis virus in Peru during 1970-1971. Am J Epidemiol, 101(4), 347–355.
Abstract: Venezuelan encephalitis (VE) virus has intermittently produced epidemics and equine epizootics on the dry Pacific coastal plain of Peru since at least the 1930's. However, evidence that the virus exists in the Amazon region of Peru to the east of the Andes mountains was not obtained until antibodies were found in human sera collected in 1965, and 10 strains of the virus were isolated in a forest near the city of Iquitos, Peru during February and March 1971. Eight strains came from mosquitoes and two from dead sentinel hamsters. Three hamsters exposed in forests near Iquitos developed VE virus antibodies suggesting that hamster-benign strains also exist there. Antibody tests of equine sera revealed no evidence that VE virus was actively cycling during the late 1950's or 1960's in southern coastal Peru, where equine epizootics had occurred in the 1930's and 1940's. In northern coastal Peru bordering Ecuador, antibodies were present in equine sera, presumably residual from the 1969 outbreak caused by subtype I virus, since neutralizing antibody titers were higher to subtype I virus than to subtypes III or IV. No VE virus was detected in this northern region during the dry season of 1970 by use of sentinel hamsters. The possibility is considered that VE epidemics and equine epizootics on the Pacific coast of Peru are caused by movements of virus in infected vertebrates traversing Andean passes or in infected vertebrates or mosquitoes carried in airplanes from the Amazon region.
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