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	Author	Mejdell, C.M.; Buvik, T.; Jørgensen, G.H.M.; Bøe, K.E.
	Title	Horses can learn to use symbols to communicate their preferences			Type	Journal Article
	Year	2016	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	184	Issue		Pages	66-73
	Keywords	Operant conditioning; Blanket; Rug; Thermoregulation; Cognition; Clicker training
	Abstract	Abstract This paper describes a method in which horses learn to communicate by touching different neutral visual symbols, in order to tell the handler whether they want to have a blanket on or not. Horses were trained for 10–15 min per day, following a training program comprising ten steps in a strategic order. Reward based operant conditioning was used to teach horses to approach and touch a board, and to understand the meaning of three different symbols. Heat and cold challenges were performed to help learning and to check level of understanding. At certain stages, a learning criterion of correct responses for 8–14 successive trials had to be achieved before proceeding. After introducing the free choice situation, on average at training day 11, the horse could choose between a “no change” symbol and the symbol for either “blanket on” or “blanket off” depending on whether the horse already wore a blanket or not. A cut off point for performance or non-performance was set to day 14, and 23/23 horses successfully learned the task within this limit. Horses of warm-blood type needed fewer training days to reach criterion than cold-bloods (P < 0.05). Horses were then tested under differing weather conditions. Results show that choices made, i.e. the symbol touched, was not random but dependent on weather. Horses chose to stay without a blanket in nice weather, and they chose to have a blanket on when the weather was wet, windy and cold (χ2 = 36.67, P < 0.005). This indicates that horses both had an understanding of the consequence of their choice on own thermal comfort, and that they successfully had learned to communicate their preference by using the symbols. The method represents a novel tool for studying preferences in horses.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0168-1591	ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	6022
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	Author	Rapin, V.; Poncet, P.A.; Burger, D.; Mermod, C.; Richard, M.A.
	Title	[Measurement of the attention time in the horse]			Type	Journal Article
	Year	2007	Publication	Schweizer Archiv fur Tierheilkunde	Abbreviated Journal	Schweiz Arch Tierheilkd
	Volume	149	Issue	2	Pages	77-83
	Keywords	Age Factors; Animals; Attention/physiology; Conditioning, Operant; Cross-Over Studies; Female; Horses/physiology; Learning/physiology; Male; Memory/physiology; Time Factors
	Abstract	A study carried out on 49 horses showed that it is possible to measure the attention time by operant conditioning. After teaching horses an instrumental task using a signal, we were then able to test their attention time by asking them to prolong it increasingly while setting success and failure criteria. Two tests were performed 3 weeks apart. The 2nd test was feasible without relearning, a proof of memory, and was repeatable, a proof of consistency in the attention time. A significant difference was observed between the 3 age groups. Young horses often performed very well during the 1st test but their attention dropped in the 2nd test while older horses were more stable with respect to attention and even increased it slightly. The study shows that there are individual differences but it was not possible to prove a significant influence of breed, gender and paternal influence. Consequently, learning appears to be one of the most interesting approaches for evaluating the attention of horses and for observing their behaviour.
	Address	Laboratoire d'Ecoethologie, Universite de Neuchatel, Suisse. veronique.rapin@unine.ch
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	French	Summary Language		Original Title	Mesure de la duree d'attention chez le cheval
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0036-7281	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:17343134			Approved	no
	Call Number				Serial	1770
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	Author	Chalmeau, R.; Gallo, A.
	Title	Social constraints determine what is learned in the chimpanzee			Type	Journal Article
	Year	1993	Publication	Behavioural Processes	Abbreviated Journal	Behav. Process.
	Volume	28	Issue	3	Pages	173-179
	Keywords	Operant conditioning; Social influence; Individual strategy; Chimpanzee
	Abstract	A group of six chimpanzees was placed in a social learning situation, without training. The learning task was an operant conditioning situation; that is, a subject had to pull two handles simultaneously to cause a piece of fruit to fall into the cage. Only three individuals acquired the operant behaviour. For the operant individuals, social influences on the expression of the learning task were then examined; the dominant chimpanzee during feeding had an inhibiting effect when close to the operant subjects. Depending on the subject, social factors may influence not only the specific expression of what is learnt, but also the nature of what is learnt. Chimpanzees appear to experience situations differently: they develop an individual problem-solving strategy according to their social relationships even if the experimental procedure is the same for all.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	refbase @ user @			Serial	569
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	Author	Shapiro, A.D.; Janik, V.M.; Slater, P.J.B.
	Title	*A gray seal's (Halichoerus grypus) responses to experimenter-given pointing and directional cues*			Type	Journal Article
	Year	2003	Publication	Journal of Comparative Psychology	Abbreviated Journal	J Comp Psychol
	Volume	117	Issue	4	Pages	355-362
	Keywords	Animals; Behavior, Animal/physiology; Cognition/physiology; Conditioning, Operant/physiology; Cues; Eye Movements/physiology; Female; Seals, Earless
	Abstract	A gray seal (Halichoerus grypus) was trained to touch a target on its left or right by responding to pointing signals. The authors then tested whether the seal would be able to generalize spontaneously to altered signals. It responded correctly to center pointing and head turning, center upper body turning, and off-center pointing but not to head turning and eye movements alone. The seal also responded correctly to brief ipsilateral and contralateral points from center and lateral positions. Pointing gestures did not cause the seal to select an object placed centrally behind it. Like many animals in similar studies, this gray seal probably did not understand the referential character of these gestures but rather used signal generalization and experience from initial operant conditioning to solve these tasks.
	Address	School of Biology, University of St Andrews, St Andrews, Fife, United Kingdom
	Corporate Author				Thesis
	Publisher		Place of Publication	Washington, D.C. : 1983	Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0735-7036	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:14717636			Approved	yes
	Call Number	Equine Behaviour @ team @			Serial	4977
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	Author	Sloet van Oldruitenborgh-Oosterbaan, M.M.; Blok, M.B.; Begeman, L.; Kamphuis, M.C.D.; Lameris, M.C.; Spierenburg, A.J.; Lashley, M.J.J.O.
	Title	Workload and stress in horses: comparison in horses ridden deep and round ('rollkur') with a draw rein and horses ridden in a natural frame with only light rein contact			Type	Journal Article
	Year	2006	Publication	Tijdschrift Voor Diergeneeskunde	Abbreviated Journal	Tijdschr Diergeneeskd
	Volume	131	Issue	5	Pages	152-157
	Keywords	Animal Husbandry/methods; Animals; Biomechanics; Blood Glucose/analysis; Female; Heart Rate/physiology; Hematocrit/veterinary; Horses/blood/physiology; Hydrocortisone/blood; Lactic Acid/blood; Physical Conditioning, Animal/adverse effects/physiology; Stress, Physiological/blood/etiology/veterinary
	Abstract	'Rollkur' or 'overbending' is the low and deep riding of a dressage horse during training or warming up. Lately, this technique has been criticized, and not necessarily objectively, on welfare grounds. To be able to evaluate these criticisms, more needs to be known about the workload and stress of horses being ridden 'rollkur'. The aim of the present study was to compare the workload of eight riding-school horses when being ridden deep and round with a draw rein ('rollkur') and when being ridden in a natural frame with only light rein contact ('free'). Workload (as measured by heart rate and blood lactate concentration) was slightly higher when horses were ridden 'rollkur' than when they were ridden 'free'. There were no differences in packed cell volume, or glucose and cortisol concentrations. No signs of uneasiness or stress could be determined when the horses were ridden 'rollkur'. Subjectively, all horses improved their way of moving during 'rollkur' and were more responsive to their rider.
	Address	Department of Equine Sciences, Faculty of Veterinary Medicine, Utrecht University, Yalelaan 12, 3584 CM Utrecht, The Netherlands. m.sloet@vet.uu.nl
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0040-7453	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:16532786			Approved	no
	Call Number	Equine Behaviour @ team @			Serial	5638
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	Author	Williams, J.L.; Friend, T.H.; Nevill, C.H.; Archer, G.
	Title	The efficacy of a secondary reinforcer (clicker) during acquisition and extinction of an operant task in horses			Type	Journal Article
	Year	2004	Publication	Applied Animal Behaviour Science	Abbreviated Journal	Appl. Anim. Behav. Sci.
	Volume	88	Issue	3-4	Pages	331-341
	Keywords	Equine; Horse; Clicker training; Operant conditioning; Secondary reinforcers
	Abstract	“Clicker training” is a popularly promoted training method based on operant conditioning with the use of a secondary reinforcer (the clicker). While this method draws from theories of learning and is used widely, there has been little scientific investigation of its efficacy. We used 60 horses, Equus callabus, and assigned each horse to one of six reinforcement protocols. The reinforcement protocols involved combinations of reinforcers administered (primary versus secondary plus primary), schedule of reinforcement (continuous versus variable ratio), and reinforcers applied during extinction (none or secondary). There were no differences (P>=0.11) between horses which received a secondary reinforcer (click) followed by the primary reinforcer (food) and those which received only the primary reinforcer (food) in the number of trials required to train the horses to touch their noses to a plastic cone (operant response). There also were no differences (P>=0.12) between horses which received the secondary reinforcer plus primary reinforcer and those which received only the primary reinforcer in regards to the number of trials to extinction. We conclude that there is no difference in the amount of training required to learn the operant task or in the task's resistance to extinction between horses that received a secondary reinforcer followed by a primary reinforcer versus horses which received only a primary reinforcer.
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes				Approved	no
	Call Number	Equine Behaviour @ team @			Serial	3581
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	Author	Cook, M.; Mineka, S.; Wolkenstein, B.; Laitsch, K.
	Title	Observational conditioning of snake fear in unrelated rhesus monkeys			Type	Journal Article
	Year	1985	Publication	Journal of abnormal psychology	Abbreviated Journal	J Abnorm Psychol
	Volume	94	Issue	4	Pages	591-610
	Keywords	Animals; Conditioning, Classical; Fear; Female; Macaca mulatta; Male; Snakes
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0021-843X	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:4078162			Approved	no
	Call Number	refbase @ user @			Serial	707
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	Author	McDonnell, S.M.; Henry,M. Bristol, F.
	Title	Spontaneous erection and masturbation in equids Proc 35th			Type	Journal Article
	Year	1991	Publication		Abbreviated Journal	J. Reprod. Fert. Suppl
	Volume	44	Issue		Pages	664-665
	Keywords	Horse; Masturbation; Erection; Aversive conditioning; Sexual behavior; Ejaculation; Semen
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language		Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN		ISBN		Medium
	Area		Expedition		Conference
	Notes	This paper is part of the special issue entitled “Proceedings of the 4th International Symposium on Stallion Reproduction”, Guest Edited by Dr. Edward Squires.			Approved	no
	Call Number				Serial	1378
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	Author	Shettleworth, S.J.
	Title	Foraging, memory, and constraints on learning			Type	Journal Article
	Year	1985	Publication	Annals of the New York Academy of Sciences	Abbreviated Journal	Ann N Y Acad Sci
	Volume	443	Issue		Pages	216-226
	Keywords	Animals; Animals, Wild; Appetitive Behavior; Avoidance Learning; Birds; Conditioning, Classical; Discrimination Learning; Food Preferences; Memory; Mental Recall; Motivation; Predatory Behavior; Rats; *Taste
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0077-8923	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:3860072			Approved	no
	Call Number	refbase @ user @			Serial	384
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	Author	Shettleworth, S.J.
	Title	Stimulus relevance in the control of drinking and conditioned fear responses in domestic chicks (Gallus gallus)			Type	Journal Article
	Year	1972	Publication	Journal of comparative and physiological psychology	Abbreviated Journal	J Comp Physiol Psychol
	Volume	80	Issue	2	Pages	175-198
	Keywords	Acoustic Stimulation; Animals; Auditory Perception; Chickens; Conditioning (Psychology); Conditioning, Classical; Discrimination Learning; Drinking Behavior; Electroshock; Fear; Light; Motor Activity; Photic Stimulation; Punishment; Quinine; *Sound; Taste; Visual Perception
	Abstract
	Address
	Corporate Author				Thesis
	Publisher		Place of Publication		Editor
	Language	English	Summary Language		Original Title
	Series Editor		Series Title		Abbreviated Series Title
	Series Volume		Series Issue		Edition
	ISSN	0021-9940	ISBN		Medium
	Area		Expedition		Conference
	Notes	PMID:5047826			Approved	no
	Call Number	refbase @ user @			Serial	390
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